Introduction

Materials and methods

Study area

The study area is situated оn the Akhtuba River (fork of Volga River) about 90 km upstream of Volga estuary, Astrakhan Province, European Russia. We were based in Dosang Railway Station (N 46°54' E 47°55'). All species listed below and the majority of specimens were collected within 1 km from the station i.e. within an area of about 3 km².

Four main biotopes were sampled: fixed sands (Fig. 1), barchan sands (Fig. 2), floodplain of Akhtuba River (Fig. 3), and riverside secondary forest (mainly of ash) (Fig. 4).

Figure 1.  

Fixed sands near Dosang, Astrakhan Province, Russia.

Figure 2.  

Barchan sands North-East of Dosang, Astrakhan Province, Russia.

Figure 3.  

Right side of Akhtuba River, floodplain, Dosang environs, Astrakhan Province, Russia.

Figure 4.  

Left side of Akhtuba River, riverine secondary forest near Dosang, Astrakhan Province, Russia.

Taxon treatment

Analysis

Discussion

Acknowledgements

References

Akhmetova LA (2011)
Beetles of the tribe Aphodiini (Coleoptera, Scarabaeidae) of Russia. Dissertation to fulfill the requirement for the degree of Doctor of Philosophy.
Zoological Institute RAS
,
Sankt-Petesburg
,
299
pp. [In
Russian
].
Akhmetova LA, Frolov AV (2008a)
New records of Aphodius species from Russia (Coleoptera: Scarabaeidae)
.
Zoosystematica Rossica
17
:
72
. [In
English
].
Akhmetova LA, Frolov AV (2008b)
A Review of Scarabs of the Subgenus Nobius Mulsant et Rey, Genus Aphodius Illiger (Coleoptera, Scarabaeidae), of the Fauna of Russia and Neighboring Countries.
Entomological Review
88
(
4
):
421
‑
434
. [In
English
]. https://doi.org/10.1134/S0013873808040052
Akhmetova LA, Frolov AV (2009)
New to Russia and little known species of the genus Aphodius Illiger (Coleoptera, Scarabaeidae).
Zoosystematica Rossica
18
(
2
):
278
‑
284
. [In
English
].
Emeljanov AF (1974)
Proposals on the classification and nomenclature of areals
.
Entomologicheskoe Obozrenie
53
(
3
):
497
‑
522
. [In
Russian
].
Frolov AV (2002)
A Review of Aphodiines of the Subgenus Chilothorax Motschulsky, Genus Aphodius Illiger (Coleoptera, Scarabaeidae), from Russia and Neighboring Countries.
Entomological Review
82
(
1
):
1
‑
18
. [In
English
].
Frolov AV (2009)
Larval morphology of Aphodius sus (Herbst) and A. variicolor Koshantschikov (Coleoptera: Scarabaeidae: Aphodiinae)
.
Zootaxa
2169
:
45
‑
54
.
Kabakov ON, Frolov AV (1996)
A Review of the Beetles of the Genus Aphodius Ill. (Coleoptera, Scarabaeidae) from Russia and Adjacent Countries, Related to the Subgenus Acrossus Muls.
Entomological Review
76
(
9
):
1165
‑
1181
. [In
English
].
Laktionov AP (2010)
Flora Astrakhanskoj Oblasti.
[
Flora of Astrakhan Province
].
Astrakhanskij Universitet
,
Astrakhan
,
296
pp. [In
Russian
].
Nikolajev GV (1987)
Plastinchatousye zhuki Kazakhstana i Srednej Azii
. [
Scarab beetles of Kazakhstan and Middle Asia
].
Nauka
,
Alma-Ata
,
232
pp. [In
Russian
].
Shokhin IV (2007)
Contribution to the fauna of lamellicorn beetles (Coleoptera, Scarabaeoidea) of Southern Russia, with some nomenclatural changes in the family Scarabaeidae.
Caucasian Entomological Bulletin
3
(
2
):
105
‑
185
. [In
Russian
].
Wilson CJ, Angus RB (2006)
A chromosomal analysis of eight species of Aphodius Illiger, subgenera Agiolinus Schmidt, Agrilinus Mulsant & Rey and Planolinus Mulsant & Rey (Coleoptera: Aphodiidae)
.
Proceedings of the Russian Entomological Society
77
:
28
‑
33
.

Supplementary material