Corresponding author: Vladimir Blagoderov (
Academic editor: Torsten Dikow
A new species of
Descriptive terminology follows
Male. Measurements (mm). Head height 0.38, palpi 0.65, proboscis 1.05, antenna 0.9, thorax length 0.63, thorax height 0.71, metepisternum anterior margin 0.15, posterior margin 0.25; coxa 1 0.53; coxa 2 0.47; coxa 3 0.4; wing 1.9.
Colouration. Body, head and antennae entirely dark brownish-grey, almost black, halters yellowish, legs yellowish-brown.
Head (Fig.
Thorax, legs, and abdomen uniformly dark brown (Fig.
Wing (Fig.
Legs (Fig.
Terminalia (Figs
Female. Unknown.
The species differs from all Afrotropical species of
The specific epithet is an adjective in reference to the place of origin of the specimens.
South Africa: Eastern Cape.
1 | Wings with distinct brown spots |
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– | Wings hyaline or smoky, without distinct spots; antennae monochromatic, brown or yellow, at least the first four flagellomeres significantly longer than wide |
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2 | Antennae flagellomeres 1–4 brown, 5 orange, 6 bright yellow, 7–11 brown, 12 bright yellow, 5–13-14 light brown; flagellomeres 1–4 twice as long as wide, 5–12 – approximately as long as wide. Gonostili slender, with apical tooth length twice its width. Côte d'Ivoire | |
– | Antennae yellow, with flagellomeres 1–3, 7–11 and 14 brown; flagellomeres with length equal to width, except the last two, which are significantly longer than wide. Gonostyli wide, with apical tooth as long as wide. Gabon, Central African Republic | |
3 | Wings uniformly brownish |
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– | Wings hyaline or very faintly smoky at the apex |
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4 | Antennae up to twice the length of the thorax and head together; proboscis yellow; subcosta short obliterated at apex, femora and tibiae brownish yellow, abdominal segments brown with yellowish white edges. Grande Comore | |
– | Antennae not longer than head and thorax together; proboscis brown; subcosta longer; ending on costa, femur and tibia dark brown; abdomen uniformly brown. Zaire | |
5 | Antennae entirely dark |
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– | Antennae entirely yellow; wings hyaline; tibial spurs II equal and shorter than apical width of tibia; internal tibial spur III twice the width of tibia apex. Madagascar | |
6 | Wings very weakly smoky at apex; thorax and abdomen brown to dark brown; proboscis 4x the length of coxa I; tergite 9 elongated, its apodeme thin, with narrow stem; internal tibial spur III 3x the width of tibia apex. Uganda, Kenya, Tanzania, Zaire | |
– | Wings hyaline; body very dark brown, almost black, proboscis ~2x the length of coxa I; tergite 9 of male roughly rounded, its apodeme very wide and thick; internal tibial spur III 2.4x the width of tibia apex. South Africa |
(based on
Authors are very grateful to all reviewers whose comments helped to improve the paper greatly.
tip of fore tibia
tip of mid tibia
apical tarsomeres of fore tarsus
apical tarsomeres of mid tarsus
gonocoxites and gonostyli, dorsal view. Scale: 0.1 mm.
gonocoxites and gonostyli, ventral view
Tergite 9, dorsal view
Tergite 9, ventral view
frontal view
lateral view
Tergite 9, dorsal view
apodeme of tergite 9, dorsal view
gonostylus, dorsal view
gonocoxite, posterior view from base, dorsal surface to right
cerci
aedeagal complex