1 urn:lsid:arphahub.com:pub:F9B2E808-C883-5F47-B276-6D62129E4FF4 urn:lsid:zoobank.org:pub:245B00E9-BFE5-4B4F-B76E-15C30BA74C02 Biodiversity Data Journal BDJ 1314-2836 1314-2828 Pensoft Publishers 10.3897/BDJ.6.e11223 11223 5898 Taxonomic Paper Tachinidae Biodiversity & Conservation Systematics Costa Rica Central America and the Caribbean Central America Americas Revision of the Mesoamerican species of Calolydella Townsend (Diptera: Tachinidae) and description of twenty-three new species reared from caterpillars in Area de Conservación Guanacaste, northwestern Costa Rica Fleming AJ ajfleming604@gmail.com https://orcid.org/0000-0002-0943-8047 1 Wood D. Monty 1 Smith M. Alex https://orcid.org/0000-0002-8650-2575 2 Hallwachs Winnie 3 Janzen Daniel H https://orcid.org/0000-0002-7335-5107 3 Agriculture Agri-Food Canada, Rm. 3099I, 960 Carling Avenue, Ottawa, Ontario, Canada Agriculture Agri-Food Canada Rm. 3099I, 960 Carling Avenue, Ottawa, Ontario Canada Department of Integrative Biology, Guelph University, Guelph, Ontario, Canada Department of Integrative Biology, Guelph University Guelph, Ontario Canada Department of Biology, University of Pennsylvania, Philadelphia, Pennsylvania 19104, United States of America Department of Biology, University of Pennsylvania Philadelphia, Pennsylvania 19104 United States of America

Corresponding author: AJ Fleming (ajfleming604@gmail.com).

Academic editor: Daniel Whitmore

 2018 02 02 2018 6 e11223 FFFAC455-6D73-FFE7-CF77-FF81FFBDCA2D 42F17C4E-B48C-4EA8-B1FC-4128BB7BF0B7 1222204 19 11 2016 25 01 2018 AJ Fleming, D. Monty Wood, M. Alex Smith, Winnie Hallwachs, Daniel H Janzen This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited. Background

Twenty-three new species of the genus Calolydella Townsend, 1927 (Diptera: Tachinidae) are described, all reared from multiple species of wild-caught caterpillars across a wide variety of families (Lepidoptera: Crambidae; Erebidae; Geometridae; Hesperiidae; Lycaenidae; Nymphalidae; Pieridae; Riodinidae; and Sphingidae). All caterpillars were collected within Area de Conservación Guanacaste (ACG), in northwestern Costa Rica. This study provides a concise description of each new species using morphology, life history, molecular data, and photographic documentation. In addition to the new species, we also provide a generic redescription and revised key to species of the genus Calolydella from Central and South America.

 New information

The following 23 new species of Calolydella are described by Fleming and Wood: C. adelinamoralesae sp. n., C. alexanderjamesi sp. n., C. argentea sp. n., C. aureofacies sp. n., C. bicolor sp. n., C. bifissus sp. n., C. crocata sp. n., C. destituta sp. n., C. discalis sp. n., C. erasmocoronadoi sp. n., C. felipechavarriai sp. n., C. fredriksjobergi sp. n., C. inflatipalpis sp. n., C. interrupta sp. n., C. nigripalpis sp. n., C. omissa sp. n., C. ordinalis sp. n., C. renemalaisei sp. n., C. susanaroibasae sp. n., C. tanyadapkeyae sp. n., C. tenebrosa sp. n., C. timjamesi sp. n., C. virginiajamesae sp. n. Lydella frugale Curran, 1934 is proposed as a new synonym of Pygophorinia peruviana Townsend, 1927, syn. n., under the combination Calolydella frugale (Curran, 1934), comb. n.

 Blondeliini caterpillars cloud forest dry forest fly Neotropics parasitoid rain forest Introduction

The New World genus Calolydella Townsend, 1927 (Tachinidae: Blondeliini) was erected for one male and several female specimens collected by Townsend himself at Itaquaquecetuba (now a suburb of São Paolo), Brazil and described as Calolydella geminata Townsend, 1927. When Wood (1985) revised the tribe Blondeliini, he synonymized three other genera under Calolydella: Olindopsis Townsend, 1927, Pygophorinia Townsend, 1927, and Prodexodes Townsend, 1927. These synonymies raised the number of species in the genus from one to 13 (Wood 1985).

Here, we describe 23 new species of Calolydella reared from wild-caught caterpillars collected as part of the ongoing inventory of Lepidoptera larvae (and their parasitoids) in the terrestrial portion of Area de Conservación Guanacaste (ACG), an area of 120,000 hectares in northwestern Costa Rica (Janzen et al. 2009, Janzen and Hallwachs 2016). Each new species is distinguished by morphological differences, differences in CO1 (cox1 or cytochrome oxidase 1) gene sequences, and comparison (by DMW and AJF) with previously described species of Calolydella. Although many of the species of Calolydella are common throughout the Neotropics (Wood 1985), none of the previously known species have been found in ACG to date. Individual and comparative details of the parasitization ecology of these flies will be treated in later papers, once the overall knowledge of the caterpillar-attacking tachinids found in ACG is more complete (Smith et al. 2006, Smith et al. 2007, Smith et al. 2008, Smith et al. 2012, Fleming et al. 2014a, Fleming et al. 2014b, Fleming et al. 2015a, Fleming et al. 2015b, Fleming et al. 2015c, Fleming et al. 2015d, Fleming et al. 2016a, Fleming et al. 2016b).

 Materials and methods

All flies and rearing information described herein were collected as part of the ongoing ACG inventory of caterpillars, their food plants, and their parasitoids, throughout the three major ACG terrestrial ecosystems (Janzen et al. 2009, Janzen and Hallwachs 2011, Janzen and Hallwachs 2015, Janzen and Hallwachs 2016). The parasitoid rearing methods are described at http://janzen.bio.upenn.edu/caterpillars/methodology/how/parasitoid_husbandry.htm. Since its inception in 1978, this inventory has reared more than 750,000 wild-caught caterpillars. All frequencies of parasitization reported here need to be considered against this background inventory.

Our present treatment of the genus Calolydella is limited to species from the Mesoamerican region, from the Mexican provinces of Guerrero and Oaxaca to the Colombian border with Panama. While we included all known species in our comparisons with the new species, only the species distributed within this region are included in our key.

Acronyms for depositories

AMNH American Museum of Natural History, New York, USA

NHMUK Natural History Museum [formerly British Museum (Natural History)], London, UK

CNC Canadian National Collection of Insects, Arachnids and Nematodes, Ottawa, Canada

MNCR Museo Nacional de Costa Rica [formerly Instituto Nacional de Biodiversidad – INBio], Santo Domingo de Heredia, Costa Rica

NMW Naturhistorisches Museum Wien, Vienna, Austria

USNM National Museum of Natural History [formerly United States National Museum], Washington, D.C., USA

 Voucher specimen management

The management of voucher specimens has been detailed in previous papers in this series (Janzen and Hallwachs 2016). In brief, caterpillars reared from the ACG inventory each receive a unique voucher code in the format yy–SRNP–xxxxx. Parasitoids emerging from these caterpillars receive the same voucher code, indicating the rearing event; when/if individually processed for DNA barcoding, each parasitoid receives a second voucher code unique to it, in the format DHJPARxxxxxxx. The associated data for each voucher code are available at http://janzen.bio.upenn.edu/caterpillars/database.lasso.

All DHJPARxxxxxxx-coded tachinids had one leg removed for DNA barcoding and couriered to the Center for Biodiversity Genomics of the Biodiversity Institute of Ontario (BIO) at the University of Guelph (Canada). All associated data and successful barcodes are permanently and publicly deposited in the Barcode of Life Data System (BOLD) (Ratnasingham and Hebert 2007), and a select set of these data are subsequently migrated to GenBank. Each barcoded specimen also receives accession numbers from the Barcode of Life Data System (BOLD) and GenBank, respectively. The dynamic nature of the inventory means that it is continually adding new specimens; newly added Calolydella specimens can be found by searching for the genus in BOLD.

All inventoried specimens discussed herein were collected under Costa Rican government research permits issued to DHJ; the Tachinidae samples were exported under permit by DHJ from Costa Rica to their final depository in the CNC. Tachinid identifications for the inventory are done by DHJ in coordination with a) visual inspection of morphology by AJF and DMW, b) DNA barcoding by MAS and BIO, and c) databasing and association with host caterpillars by DHJ and WH through the inventory itself.

The date of capture of fly specimens is the date of eclosion of the fly and not the date of capture of the caterpillar. The fly eclosion date is much more representative of the time when that species is on the wing than is the date of capture of the parasitized caterpillar. The “collector” is the parataxonomist who collected the caterpillar, rather than the person who later retrieved the newly eclosed fly and processed it by freezing, pinning, labeling and oven-drying. The biology and parasitization rates of individual tachinid species will be the subject of later papers. The holotypes of the new species described herein are deposited in the Diptera collection of the CNC, Ottawa, Canada.

 Imaging

Habitus and terminalia photographs were taken using the methods outlined in Fleming et al. (2014a). Raw image files were first processed with Adobe Photoshop CS6, then digitally stacked to produce a final composite image using Zerene Stacker Software v1.04.

Adult flies were dissected following standard practice (O’Hara 1983). The morphological terminology follows Cumming and Wood (2009). If only one male was available, it was designated as the holotype and not subjected to dissection.

 Interim names for undescribed host species

Names of undescribed host species follow a standardized, interim naming system used for taxonomic units considered to be distinct species and identified by a combination of DNA barcode, morphology, and host-plant information. The interim names are given in the format "Eois Janzen52", where the species epithet is composed of the name of the taxonomist who first singled out the species and a number. This prevents confusion with already described species while maintaining traceability of each undescribed species within the ACG project.

 DNA barcoding

We analyzed DNA barcodes (the 5’ region of the cytochrome c oxidase I (CO1) gene, Hebert et al. 2004), where available, for all specimens of ACG Calolydella. The 658 bp DNA barcodes were amplified from total DNA extracts obtained from single legs with a standard glass fibre protocol, using the standard insect primers (LepF1–LepR1) (Ivanova et al. 2006,Smith et al. 2006, Smith et al. 2007, Smith et al. 2008).

 Taxon treatments 994AC7D6-B831-59ED-BA31-D4CB08B91DA0 Calolydella Townsend, 1927 Calolydella Townsend, 1927a: 278. Type-species: Calolydella geminata Townsend, 1927, by original designation. Olindopsis Townsend, 1927a: 274. Type-species: Olindopsis andinensis Townsend, 1927, by original designation. Synonymy proposed by Wood (1985). Pygophorinia Townsend, 1927a: 274. Type-species: Pygophorinia peruviana Townsend, 1927, by original designation. Synonymy proposed by Wood (1985). Prodexodes Townsend, 1927a: 280. Type-species: Prodexodes rufiventris Townsend, 1927, by original designation. Synonymy proposed by Wood (1985). Calolydella Calolydella geminata Townsend, 1927 Description

Calolydella belongs to the tribe Blondeliini. Wood (1985) defined the tribe by thethe following combination of character states: prosternum setose; first postsutural supra-alar seta smaller than first postsutural dorsocentral seta; bend of vein M rounded, obtuse; subapical scutellar setae long, stout, and divergent, their bases situated close together; wing veins R4+5 and M ending at or near wing tip; often elongate and narrow body plan; mid-dorsal depression of syntergite 1+2 (ST1+2) rarely reaching hind margin of syntergite; mid tibia, in many genera, with a single long anterodorsal seta.

The following redescription applies to both males and females of Calolydella; any differences between the sexes are noted. Head: males lacking proclinate orbital setae, females with 2 pairs of proclinate orbital setae; first reclinate orbital seta either equal to or longer than uppermost frontal seta; ocellar setae long; eye bare, if haired then hairs minute and inconspicuous; parafacial bare; facial margin level with vibrissa and not visible in profile, with a few small supra-vibrissal setae; subvibrissal ridge short, with three or fewer setae; margin of postgena straight or slightly convex anteriorly, genal dilation weak or absent; palps normally slender and yellow, some species with black or inflated palps; postpedicel of similar length in both sexes; arista minutely pubescent, slightly thickened at base. Thorax: prosternum setose; proepisternum bare; postpronotum with 2–4 setae in a narrow triangle or a straight line (rarely inner basal seta absent); acrostichal setae 1–3:3–4; dorsocentral setae 1–3:2–3; intra-alar setae 1–3:2–3; supra-alar setae 1–2:3; katepisternal setae 2–3 [Wood (1985) described the genus as having three katepisternal setae; however, we show that this trait can vary in females]; lateral scutellar setae shorter than subapical setae and curved medially; fore tibia with one posterior seta; mid tibia with one anterodorsal seta; wings ranging from smoky gray to pale translucent brown; wing vein R4+5 with 2–7 setulae dorsally at base, these sometimes extending to crossvein R-M. Abdomen: ST1+2 with one pair of median marginal setae, mid-dorsal depression not extending to hind margin of syntergite; 3rd and 4th abdominal tergites with 1–2 pairs of discal setae. Abdominal tergites marginally banded with gold pollinosity; in some species banding interrupted by a dark median stripe; male abdomen elongated, conical, tapering to a sharp point; in females, abdomen rounded and downwardly curved; in some species the coloration and pattern of pollinosity are sexually dimorphic.

Other species included in Calolydella Townsend

andinensis Townsend, 1927: 339 (Olindopsis). Holotype female (USNM), by original designation [examined by DMW]. Type locality: Peru, Cordillera Oriental, near Tabalosas, 3000ft.

blandita Wulp, 1890: 142 (Hypostena). Lectotype male (NHMUK), by fixation of Wood (1985:28). Type locality: Mexico, Guerrero, Sierra de las Aguas Escondidas, 7000ft.

cingulata Schiner, 1868: 327 (Meigenia). Holotype male (NMW), by original designation [examined by DMW]. Type locality: Brazil.

cylindriventris Wulp, 1890: 145 (Hypostena). Lectotype male (NHMUK), by fixation of Wood (1985:28). Type locality: Mexico, Guerrero, Sierra de las Aguas Escondidas, 7000ft.

geminata Townsend, 1927: 293 (Calolydella). Multiple syntypes (USNM) [1 female and several males examined by DMW]. Type locality: Brazil, São Paulo, Itaquaquecetuba.

gentica Walker, 1860: 302 (Masicera). Lectotype male (NHMUK), by fixation of Wood (1985:28) (examination of “Holotype: ♂” from Mexico in NHMUK is regarded as a lectotype fixation [JE O'Hara, pers. comm.]) [examined by DMW]. Type locality: Mexico.

lathami Curran, 1925: 284 (Lydella). Holotype male (CNC), by original designation [examined by DMW]. Type locality: USA, New York, Long Island, Greenpoint.

leucophaea Wulp, 1890: 141 (Hypostena). Lectotype female (NHMUK), by fixation of Wood (1985:28). Type locality: Mexico, Guerrero, Sierra de las Aguas Escondidas, 9500ft.

peruviana Townsend, 1927: 355 (Pygophorinia). Holotype female (USNM), by original designation [examined by DMW]. Type locality: Peru, [Puno], Casahuiri.

frugale Curran, 1934: 511 (Lydella). Holotype female (AMNH), by original designation [examined by DMW]. Type locality: Guyana, Bartica, Kartabo. Syn n., comb. n.

rufiventris Townsend, 1927: 350 (Prodexodes). Holotype male (USNM), by original designation [examined by DMW]. Type locality: Brazil, Sao Paulo, Itaquaquecetuba.

summatis Reinhard, 1974: 1158 (Calolydella). Holotype male (CNC), by original designation [examined by DMW]. Type locality: Mexico, Durango, 14 miles southwest of El Salto, 8000ft.

triangulifera Bigot, 1889: 268 (Homodexia). Holotype male (NHMUK), by monotypy [examined by DMW]. Type locality: Mexico.

pictigaster Bigot, 1889: 261 (Ceromasia). Holotype female (NHMUK), by monotypy [examined by DMW]. Type locality: Mexico (NHUMK).

trifasciata Walker, 1837: 350 (Tachina). Lectotype female (NHMUK), by fixation of Wood (1985:29) (examination of “Holotype: ♀” from South America in NHMUK is regarded as a lectotype fixation [JE O'Hara, pers. comm.]) [examined by DMW]. Type locality: South America, precise locality uncertain (Austen 1907).

quadrivittata Bigot, 1889: 261 (Ceromasia). Holotype male (NHMUK) [published as female], by monotypy [examined by DMW]. Type locality: Mexico.

quadristriata Wulp, 1890: 146 (Hypostena). Holotype male (NHMUK), by original designation [examined by DMW]. Type locality: Costa Rica, Volcán de Irazu, 6000ft.

 Diagnosis

Calolydella are slender-bodied flies characterized by an abdomen longer than it is wide, with bold abdominal markings of yellow-gold pollinose stripes on a black background. According to Wood (1985), they lack distinctive chaetotactic characters and are difficult to characterize to genus except by their general appearance. Wood (1985) suggested the presence of only two pairs of presutural acrostichal setae as a diagnostic character; however, our current examination of this trait suggests that it varies among species, as does the rest of the thoracic chaetotaxy. Sternite 5 with only short hair-like setae, all more or less equal in size. Sternites 6 and 7 of female larger and more heavily sclerotized.

 Distribution

Essentially a Neotropical genus save for one species, Calolydella lathami, which is present in the eastern USA and adjacent Canada.

 Ecology

All species of Calolydella are parasitoids of Lepidoptera larvae in the families Crambidae, Erebidae, Geometridae, Hesperiidae, Lycaenidae, Notodontidae, Nymphalidae, Pieridae, Riodinidae, and Sphingidae.

 D92E8F57-E496-5146-BA28-6F1DEA8EEC92Calolydella adelinamoralesaeurn:lsid:zoobank.org:act:AE0FC424-12A9-4FDE-A8AF-53FE35C429F3 Fleming & Wood sp. n. Materials

Type status: Holotype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017775; recordedBy: D.H. Janzen, W. Hallwachs & Carolina Cano; individualID: DHJPAR0017775; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 04-SRNP-780, BOLD:AAA7512, ASTAR486-07; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector San Cristobal; locality: Area de Conservacion Guanacaste; verbatimLocality: Puente Palma; verbatimElevation: 460; verbatimLatitude: 10.9163; verbatimLongitude: -85.3787; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 01-Mar-2004; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017079; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: DHJPAR0017079; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 07-SRNP-31260, BOLD:AAA7512, ASTAP517-07; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Pasmompa; verbatimElevation: 440; verbatimLatitude: 11.0193; verbatimLongitude: -85.41; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 14-Mar-2007; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0035793; recordedBy: D.H. Janzen, W. Hallwachs & Keiner Aragon; individualID: DHJPAR0035793; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 09-SRNP-67604, BOLD:AAA7512, ASHYD1174-09; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Palomo; verbatimElevation: 96; verbatimLatitude: 10.9619; verbatimLongitude: -85.2804; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 19-Jul-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017085; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: DHJPAR0017085; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 07-SRNP-31239, BOLD:AAA7512, ASTAP523-07; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Pasmompa; verbatimElevation: 440; verbatimLatitude: 11.0193; verbatimLongitude: -85.41; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 16-Mar-2007; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0037401; recordedBy: D.H. Janzen, W. Hallwachs & Cirilo Umana; individualID: DHJPAR0037401; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 09-SRNP-76683, BOLD:AAA7512, ASHYC4146-10; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Estacion Llanura; verbatimElevation: 135; verbatimLatitude: 10.9333; verbatimLongitude: -85.2533; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 20-Dec-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0035728; recordedBy: D.H. Janzen, W. Hallwachs & Cirilo Umana; individualID: DHJPAR0035728; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 09-SRNP-44796, BOLD:AAA7512, ASHYD1109-09; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Estacion Llanura; verbatimElevation: 135; verbatimLatitude: 10.9333; verbatimLongitude: -85.2533; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 29-Jul-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011693; recordedBy: D.H. Janzen, W. Hallwachs & Roster Moraga; individualID: DHJPAR0011693; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 04-SRNP-25478, BOLD:AAA7512, ASTAS419-06; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste; verbatimLocality: Uncaria; verbatimElevation: 370; verbatimLatitude: 11.0175; verbatimLongitude: -85.4741; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 25-Oct-2004; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0030226; recordedBy: D.H. Janzen, W. Hallwachs & Duvalier Briceno; individualID: DHJPAR0030226; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 09-SRNP-65018, BOLD:AAA7512, ASHYB970-09; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Brasilia; locality: Area de Conservacion Guanacaste; verbatimLocality: Moga; verbatimElevation: 320; verbatimLatitude: 11.0123; verbatimLongitude: -85.3493; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 24-Jan-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017077; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: DHJPAR0017077; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 07-SRNP-31268, BOLD:AAA7512, ASTAP515-07; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Pasmompa; verbatimElevation: 440; verbatimLatitude: 11.0193; verbatimLongitude: -85.41; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 14-Mar-2007; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0009962; recordedBy: D.H. Janzen, W. Hallwachs & Yessenia Mendoza; individualID: DHJPAR0009962; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-3195, BOLD:AAA7512, ASTAV563-06; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector San Cristobal; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Huerta; verbatimElevation: 527; verbatimLatitude: 10.9305; verbatimLongitude: -85.3722; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 12-May-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011699; recordedBy: D.H. Janzen, W. Hallwachs & Roster Moraga; individualID: DHJPAR0011699; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 04-SRNP-25477, BOLD:AAA7512, ASTAS425-06; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste; verbatimLocality: Uncaria; verbatimElevation: 370; verbatimLatitude: 11.0175; verbatimLongitude: -85.4741; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 26-Oct-2004; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0038669; recordedBy: D.H. Janzen, W. Hallwachs & Keiner Aragon; individualID: DHJPAR0038669; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 10-SRNP-67234, BOLD:AAA7512, ASHYD2242-10; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Palomo; verbatimElevation: 96; verbatimLatitude: 10.9619; verbatimLongitude: -85.2804; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 17-Mar-2010; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0035829; recordedBy: D.H. Janzen, W. Hallwachs & Duvalier Briceno; individualID: DHJPAR0035829; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 09-SRNP-65918, BOLD:AAA7512, ASHYD1210-09; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Brasilia; locality: Area de Conservacion Guanacaste; verbatimLocality: Moga; verbatimElevation: 320; verbatimLatitude: 11.0123; verbatimLongitude: -85.3493; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 14-Jul-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017088; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: DHJPAR0017088; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 07-SRNP-31244, BOLD:AAA7512, ASTAP526-07; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Pasmompa; verbatimElevation: 440; verbatimLatitude: 11.0193; verbatimLongitude: -85.41; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 16-Mar-2007; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0035784; recordedBy: D.H. Janzen, W. Hallwachs & Cirilo Umana; individualID: DHJPAR0035784; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 09-SRNP-67687, BOLD:AAA7512, ASHYD1165-09; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Palomo; verbatimElevation: 96; verbatimLatitude: 10.9619; verbatimLongitude: -85.2804; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 19-Jul-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0030217; recordedBy: D.H. Janzen, W. Hallwachs & Duvalier Briceno; individualID: DHJPAR0030217; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 09-SRNP-65019, BOLD:AAA7512, ASHYB961-09; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Brasilia; locality: Area de Conservacion Guanacaste; verbatimLocality: Moga; verbatimElevation: 320; verbatimLatitude: 11.0123; verbatimLongitude: -85.3493; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 28-Jan-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0037372; recordedBy: D.H. Janzen, W. Hallwachs & Cirilo Umana; individualID: DHJPAR0037372; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 09-SRNP-76670, BOLD:AAA7512, ASHYC4117-10; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Estacion Llanura; verbatimElevation: 135; verbatimLatitude: 10.9333; verbatimLongitude: -85.2533; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 19-Dec-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0035795; recordedBy: D.H. Janzen, W. Hallwachs & Cirilo Umana; individualID: DHJPAR0035795; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 09-SRNP-67644, BOLD:AAA7512, ASHYD1176-09; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Palomo; verbatimElevation: 96; verbatimLatitude: 10.9619; verbatimLongitude: -85.2804; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 24-Jul-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017083; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: DHJPAR0017083; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 07-SRNP-31229, BOLD:AAA7512, ASTAP521-07; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Pasmompa; verbatimElevation: 440; verbatimLatitude: 11.0193; verbatimLongitude: -85.41; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 14-Mar-2007; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017081; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: DHJPAR0017081; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 07-SRNP-31234, BOLD:AAA7512, ASTAP519-07; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Pasmompa; verbatimElevation: 440; verbatimLatitude: 11.0193; verbatimLongitude: -85.41; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 15-Mar-2007; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017086; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: DHJPAR0017086; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 07-SRNP-31272, BOLD:AAA7512, ASTAP524-07; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Pasmompa; verbatimElevation: 440; verbatimLatitude: 11.0193; verbatimLongitude: -85.41; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 16-Mar-2007; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0034521; recordedBy: D.H. Janzen, W. Hallwachs & Duvalier Briceno; individualID: DHJPAR0034521; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 09-SRNP-67040, BOLD:AAA7512, ASHYC1173-09; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Palomo; verbatimElevation: 96; verbatimLatitude: 10.9619; verbatimLongitude: -85.2804; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 01-Jun-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017076; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: DHJPAR0017076; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 07-SRNP-31277, BOLD:AAA7512, ASTAP514-07; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Pasmompa; verbatimElevation: 440; verbatimLatitude: 11.0193; verbatimLongitude: -85.41; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 14-Mar-2007; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017075; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: DHJPAR0017075; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 07-SRNP-31263, BOLD:AAA7512, ASTAP513-07; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Pasmompa; verbatimElevation: 440; verbatimLatitude: 11.0193; verbatimLongitude: -85.41; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 15-Mar-2007; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017087; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: DHJPAR0017087; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 07-SRNP-31231, BOLD:AAA7512, ASTAP525-07; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Pasmompa; verbatimElevation: 440; verbatimLatitude: 11.0193; verbatimLongitude: -85.41; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 16-Mar-2007; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0035781; recordedBy: D.H. Janzen, W. Hallwachs & Cirilo Umana; individualID: DHJPAR0035781; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 09-SRNP-67672, BOLD:AAA7512, ASHYD1162-09; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Palomo; verbatimElevation: 96; verbatimLatitude: 10.9619; verbatimLongitude: -85.2804; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 25-Jul-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0035798; recordedBy: D.H. Janzen, W. Hallwachs & Keiner Aragon; individualID: DHJPAR0035798; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 09-SRNP-67599, BOLD:AAA7512, ASHYD1179-09; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Palomo; verbatimElevation: 96; verbatimLatitude: 10.9619; verbatimLongitude: -85.2804; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 25-Jul-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0010349; recordedBy: D.H. Janzen, W. Hallwachs & Minor Carmona; individualID: DHJPAR0010349; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-42838, BOLD:AAA7512, ASTAS180-06; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Cabanya; verbatimElevation: 340; verbatimLatitude: 10.877; verbatimLongitude: -85.2308; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 28-Aug-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0035732; recordedBy: D.H. Janzen, W. Hallwachs & Cirilo Umana; individualID: DHJPAR0035732; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 09-SRNP-44788, BOLD:AAA7512, ASHYD1113-09; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Estacion Llanura; verbatimElevation: 135; verbatimLatitude: 10.9333; verbatimLongitude: -85.2533; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 27-Jul-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0035707; recordedBy: D.H. Janzen, W. Hallwachs & Cirilo Umana; individualID: DHJPAR0035707; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 09-SRNP-44825, BOLD:AAA7512, ASHYD1088-09; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Estacion Llanura; verbatimElevation: 135; verbatimLatitude: 10.9333; verbatimLongitude: -85.2533; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 21-Jul-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0030227; recordedBy: D.H. Janzen, W. Hallwachs & Duvalier Briceno; individualID: DHJPAR0030227; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 09-SRNP-65020, BOLD:AAA7512, ASHYB971-09; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Brasilia; locality: Area de Conservacion Guanacaste; verbatimLocality: Moga; verbatimElevation: 320; verbatimLatitude: 11.0123; verbatimLongitude: -85.3493; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 28-Jan-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011700; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: DHJPAR0011700; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 05-SRNP-30308, BOLD:AAA7512, ASTAS426-06; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Pasmompa; verbatimElevation: 440; verbatimLatitude: 11.0193; verbatimLongitude: -85.41; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0035705; recordedBy: D.H. Janzen, W. Hallwachs & Cirilo Umana; individualID: DHJPAR0035705; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 09-SRNP-44799, BOLD:AAA7512, ASHYD1086-09; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Estacion Llanura; verbatimElevation: 135; verbatimLatitude: 10.9333; verbatimLongitude: -85.2533; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 22-Jul-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011694; recordedBy: D.H. Janzen, W. Hallwachs & Roster Moraga; individualID: DHJPAR0011694; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 04-SRNP-25355, BOLD:AAA7512, ASTAS420-06; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste; verbatimLocality: Uncaria; verbatimElevation: 370; verbatimLatitude: 11.0175; verbatimLongitude: -85.4741; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 26-Oct-2004; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0035799; recordedBy: D.H. Janzen, W. Hallwachs & Cirilo Umana; individualID: DHJPAR0035799; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 09-SRNP-67660, BOLD:AAA7512, ASHYD1180-09; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Palomo; verbatimElevation: 96; verbatimLatitude: 10.9619; verbatimLongitude: -85.2804; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 25-Jul-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0038666; recordedBy: D.H. Janzen, W. Hallwachs & Keiner Aragon; individualID: DHJPAR0038666; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 10-SRNP-67215, BOLD:AAA7512, ASHYD2239-10; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Palomo; verbatimElevation: 96; verbatimLatitude: 10.9619; verbatimLongitude: -85.2804; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 14-Mar-2010; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0035796; recordedBy: D.H. Janzen, W. Hallwachs & Keiner Aragon; individualID: DHJPAR0035796; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 09-SRNP-67618, BOLD:AAA7512, ASHYD1177-09; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Palomo; verbatimElevation: 96; verbatimLatitude: 10.9619; verbatimLongitude: -85.2804; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 24-Jul-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0019750; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: DHJPAR0019750; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 07-SRNP-31242, , ASTAB298-07; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Pasmompa; verbatimElevation: 440; verbatimLatitude: 11.0193; verbatimLongitude: -85.41; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 30-Mar-2007; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0035792; recordedBy: D.H. Janzen, W. Hallwachs & Cirilo Umana; individualID: DHJPAR0035792; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 09-SRNP-67630, BOLD:AAA7512, ASHYD1173-09; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Palomo; verbatimElevation: 96; verbatimLatitude: 10.9619; verbatimLongitude: -85.2804; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 19-Jul-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0019763; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: DHJPAR0019763; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 07-SRNP-31258, BOLD:AAA7512, ASTAB311-07; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Pasmompa; verbatimElevation: 440; verbatimLatitude: 11.0193; verbatimLongitude: -85.41; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 24-Mar-2007; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017082; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: DHJPAR0017082; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 07-SRNP-31269, BOLD:AAA7512, ASTAP520-07; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Pasmompa; verbatimElevation: 440; verbatimLatitude: 11.0193; verbatimLongitude: -85.41; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 15-Mar-2007; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011692; recordedBy: D.H. Janzen, W. Hallwachs & Roster Moraga; individualID: DHJPAR0011692; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 04-SRNP-25476, BOLD:AAA7512, ASTAS418-06; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste; verbatimLocality: Uncaria; verbatimElevation: 370; verbatimLatitude: 11.0175; verbatimLongitude: -85.4741; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 25-Oct-2004; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0010355; recordedBy: D.H. Janzen, W. Hallwachs & Minor Carmona; individualID: DHJPAR0010355; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-42840, BOLD:AAA7512, ASTAS186-06; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Cabanya; verbatimElevation: 340; verbatimLatitude: 10.877; verbatimLongitude: -85.2308; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 27-Aug-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011698; recordedBy: D.H. Janzen, W. Hallwachs & Roster Moraga; individualID: DHJPAR0011698; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 04-SRNP-25331, BOLD:AAA7512, ASTAS424-06; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste; verbatimLocality: Uncaria; verbatimElevation: 370; verbatimLatitude: 11.0175; verbatimLongitude: -85.4741; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 26-Oct-2004; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011697; recordedBy: D.H. Janzen, W. Hallwachs & Roster Moraga; individualID: DHJPAR0011697; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 04-SRNP-25475, BOLD:AAA7512, ASTAS423-06; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste; verbatimLocality: Uncaria; verbatimElevation: 370; verbatimLatitude: 11.0175; verbatimLongitude: -85.4741; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 26-Oct-2004; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0035797; recordedBy: D.H. Janzen, W. Hallwachs & Cirilo Umana; individualID: DHJPAR0035797; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 09-SRNP-67673, BOLD:AAA7512, ASHYD1178-09; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Palomo; verbatimElevation: 96; verbatimLatitude: 10.9619; verbatimLongitude: -85.2804; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 24-Jul-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017078; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: DHJPAR0017078; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 07-SRNP-31251, BOLD:AAA7512, ASTAP516-07; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Pasmompa; verbatimElevation: 440; verbatimLatitude: 11.0193; verbatimLongitude: -85.41; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 14-Mar-2007; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017040; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: DHJPAR0017040; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 07-SRNP-31250, BOLD:AAA7512, ASTAP478-07; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Pasmompa; verbatimElevation: 440; verbatimLatitude: 11.0193; verbatimLongitude: -85.41; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 16-Mar-2007; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0035786; recordedBy: D.H. Janzen, W. Hallwachs & Cirilo Umana; individualID: DHJPAR0035786; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 09-SRNP-67652, BOLD:AAA7512, ASHYD1167-09; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Palomo; verbatimElevation: 96; verbatimLatitude: 10.9619; verbatimLongitude: -85.2804; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 22-Jul-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011703; recordedBy: D.H. Janzen, W. Hallwachs & Roster Moraga; individualID: DHJPAR0011703; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 04-SRNP-25381, BOLD:AAA7512, ASTAS429-06; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste; verbatimLocality: Uncaria; verbatimElevation: 370; verbatimLatitude: 11.0175; verbatimLongitude: -85.4741; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 31-Oct-2004; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017080; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: DHJPAR0017080; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 07-SRNP-31254, BOLD:AAA7512, ASTAP518-07; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Pasmompa; verbatimElevation: 440; verbatimLatitude: 11.0193; verbatimLongitude: -85.41; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 18-Mar-2007; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011702; recordedBy: D.H. Janzen, W. Hallwachs & Roster Moraga; individualID: DHJPAR0011702; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 04-SRNP-25485, BOLD:AAA7512, ASTAS428-06; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste; verbatimLocality: Uncaria; verbatimElevation: 370; verbatimLatitude: 11.0175; verbatimLongitude: -85.4741; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 31-Oct-2004; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0035780; recordedBy: D.H. Janzen, W. Hallwachs & Cirilo Umana; individualID: DHJPAR0035780; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 09-SRNP-67684, BOLD:AAA7512, ASHYD1161-09; Taxon: scientificName: Calolydella adelinamoralesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: adelinamoralesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Palomo; verbatimElevation: 96; verbatimLatitude: 10.9619; verbatimLongitude: -85.2804; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the arctiine erebid moth, Agaraea minuta; verbatimEventDate: 25-Jul-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

 Description

Male (Fig. 1a, b, c). Length: 5–8mm. Head (Fig. 1b): frontal setae extending to base of postpedicel; fronto-orbital plate gold and sparsely setulose; parafacial silver throughout. Thorax (Fig. 1a, c): gold on dorsal surface, silver laterally (>50% coverage); four thoracic vittae, these becoming almost fused postsuturally; postpronotum with three setae in a narrow triangle; 3:3 acrostichal setae; 2:4 dorsocentral setae; 1:3 intra-alar setae; 1:3 supra-alar setae; three katepisternal setae; anatergite bare (rarely setose, one specimen with two hair-like seta); scutellar discal setae situated as wide apart as subapical scutellar setae. Wing vein R4+5 with at most four small setulae dorsally at base. Abdomen (Fig. 1a, c): ground color dark brown-orange, with transverse marginal pollinose bands; abdominal pollinosity gold dorsally, silver ventrally, with an orange spot lateroventrally at base of ST1+2; T3 with one pair of median marginal setae and 1–2 pairs of discal setae; T4 with 1–2 pairs of discal setae. Terminalia (Fig. 2): sternite 5 with two small lobes and a wide U-shaped median cleft, 0.39X the length of the sternite from lobe to apex (Fig. 2c); inner margin covered by dense pollinosity, appearing darker than surrounding cuticle; lobes of sternite with many short stout setae marginally. Cerci, in dorsal view (Fig. 2b), separated by a narrow gap; cercus, in lateral view, long, upturned, slender and pointed, slightly tapered apically from its already narrow base; cercus setose along its anterior 1/2. Surstylus (Fig. 2a), short and spatulate, 4/5 length of cercus, appearing rounded at apex when viewed laterally; apical half with multiple short setae; tip of surstylus slightly angled inwards when viewed dorsally.

Female (Fig. 1d, e, f). Length: 5–7mm. As male, except for the following characters: fronto-orbital plate 1.6X as wide as in male; 2:3 acrostichal setae; wing vein R4+5 with 6–7 setulae extending to crossvein R-M, and ground color of abdomen appearing slightly darker than in male.

 Diagnosis

Calolydella adelinamoralesae can be distinguished from all other species of Calolydella by the following combination of traits: parafacial all silver, frontal setae extending to base of postpedicel, anatergite bare, and T4 with 1–2 pairs of discal setae.

 Etymology

The specific epithet is in honor of Adelina Morales Chaves of La Cruz, Guanacaste, Costa Rica, in recognition of the many years she dedicated to sorting ACG Malaise trap samples, many of which contained Calolydella flies.

 Distribution

Costa Rica, ACG, Alajuela and Guanacaste provinces, 96–527m.

 Ecology

Calolydella adelinamoralesae has been reared 53 times from Agaraea minuta (Schaus, 1892) (Lepidoptera: Erebidae), in rain forest and dry-rain lowland intergrade.

 9D2DD575-D9E9-54BF-A092-F6B08747E72DCalolydella alexanderjamesiurn:lsid:zoobank.org:act:D19E48EA-474C-4B9D-B784-3D2E275CCEF1 Fleming & Wood sp. n. Materials

Type status: Holotype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017782; recordedBy: D.H. Janzen, W. Hallwachs & gusaneros; individualID: DHJPAR0017782; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 99-SRNP-16175, BOLD:AAW8659, ASTAR493-07; Taxon: scientificName: Calolydella alexanderjamesi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: alexanderjamesi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Santa Rosa; locality: Area de Conservacion Guanacaste; verbatimLocality: Area Administrativa; verbatimElevation: 295; verbatimLatitude: 10.8376; verbatimLongitude: -85.6187; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the erebid moth, Correbia undulata; verbatimEventDate: 16-Nov-1999; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

 Description

Male (Fig. 3a, b, c). Length: 7mm. Head (Fig. 3b): frontal setae extending to base of postpedicel; fronto-orbital plate gold, sparsely setulose throughout; parafacial at least 50% silver pollinose. Thorax (Fig. 3a, c): gold pollinose dorsally with a gold tinge on lateral surfaces; outermost two thoracic vittae twice as wide as innermost two, these becoming fused postsuturally; postpronotum with two setae (inner basal seta absent); 3:3 acrostichal setae; 3:4 dorsocentral setae; 2:2 intra-alar setae; 2:3 supra-alar setae; three katepisternal setae; anatergite bare; scutellar discal setae situated as wide apart as subapical scutellar setae. Wing vein R4+5 with at most three small setulae dorsally, near base. Abdomen (Fig. 3a): ground color dark brown-orange with uninterrupted transverse marginal pollinose bands; abdominal pollinosity gold dorsally, silver ventrally, with an orange spot lateroventrally at base of ST1+2; T3 with one pair of median marginal setae and one pair of discal setae; T4 with one pair of discal setae. Terminalia: not examined.

Female: not known at this time.

 Diagnosis

Calolydella alexanderjamesi can be distinguished from all other species of Calolydella by the following combination of traits: parafacial mostly gold, fronto-orbital plate with small black setulae interspersed among frontal setae, anatergite bare, and thoracic pollinosity gold on both dorsal and lateral surfaces.

 Etymology

The specific epithet is in honor of Alexander James of Levittown, Pennsylvania, in recognition of the moral and family support of his mother, Tanya Dapkey, in her efforts curating and preparing ACG parasitoid flies for DNA barcoding.

 Distribution

Costa Rica, ACG, Guanacaste Province, Area Administrativa, 295m.

 Ecology

Calolydella alexanderjamesi has been reared once from Correbia undulata (Druce, 1884) (Lepidoptera: Erebidae), in dry forest.

 B2282B4F-A1AE-5849-9CF8-753C88B95AF4Calolydella argenteaurn:lsid:zoobank.org:act:DB3A83D1-98DF-498E-B4BA-6A3736CDE596 Fleming & Wood sp. n. Materials

Type status: Holotype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0042289; recordedBy: D.H. Janzen, W. Hallwachs & Cirilo Umana; individualID: DHJPAR0042289; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 11-SRNP-75105, BOLD:ABA7233, ASHYH053-11; Taxon: scientificName: Calolydella argentea; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: argentea; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Finca Esmeralda; verbatimElevation: 123; verbatimLatitude: 10.9355; verbatimLongitude: -85.2531; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the noctuid moth, Antiblemma Poole33; verbatimEventDate: 17-Mar-2011; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

 Description

Female (Fig. 4a, b, c). Length: 6mm. Head (Fig. 4b): frontal setae extending to base of postpedicel; fronto-orbital plate all silver, sparsely setulose throughout; parafacial silver throughout. Thorax (Fig. 4a, c): silver pollinose on both dorsal and lateral surfaces; with four regular thoracic vittae; postpronotum with three setae; 3:3 acrostichal setae; 2:3 dorsocentral setae; 2:3 intra-alar setae; 2:3 supra-alar setae; three katepisternal setae; anatergite with three or more setae, often in a small tuft; scutellar discal setae situated slightly wider apart than subapical scutellar setae. Wing vein R4+5 with 6–7 setulae dorsally, reaching from base almost to crossvein R-M. Abdomen (Fig. 4a): ground color black with uninterrupted transverse marginal pollinose bands covering anterior marginal 2/3 of tergites; both abdominal surfaces (dorsal and ventral) concolorous; base of ST1+2 black lateroventrally; T3 with one pair of median marginal setae and one pair of discal setae; T4 with one pair of discal setae.

Male: not known at this time.

 Diagnosis

Calolydella argentea can be distinguished from all other species of Calolydella by the following combination of traits: parafacial and fronto-orbital plate entirely silver, frontal setae extending to base of postpedicel with intermingled small setulae, 3:3 acrostichal setae, anterior 2/3 of abdominal tergites with silver pollinose bands, anatergite with three or more hair-like setae, and wing vein R4+5 with 6–7 setulae, almost reaching to crossvein R-M.

 Etymology

The specific epithet is derived from the Latin adjective "argentum", meaning silver, in reference to its entirely silver parafacial and fronto-orbital plate, a feature unique to this species.

 Distribution

Costa Rica, ACG, Alajuela Province, Finca Esmeralda, 123m.

 Ecology

Calolydella argentea has been reared once from Antiblemma Poole33 (Lepidoptera: Noctuidae), in rain forest.

 9CF76ED3-2EF0-52A1-8DB4-EF4C797BF323Calolydella aureofaciesurn:lsid:zoobank.org:act:EFC688A2-D2B7-4530-B9CD-47A41F29087C Fleming & Wood sp. n. Materials

Type status: Holotype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0029582; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Pereira; individualID: DHJPAR0029582; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 08-SRNP-36592, ASHYM1003-09; Taxon: scientificName: Calolydella aureofacies; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: aureofacies; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Cacao; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Ponderosa; verbatimElevation: 1060; verbatimLatitude: 10.9146; verbatimLongitude: -85.4626; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larvae of the geometrid moth, Cyclophora Janzen23; verbatimEventDate: 26-Sep-2008; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0029601; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Pereira; individualID: DHJPAR0029601; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 08-SRNP-36589, BOLD:AAH9540, ASHYM1022-09; Taxon: scientificName: Calolydella aureofacies; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: aureofacies; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Cacao; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Ponderosa; verbatimElevation: 1060; verbatimLatitude: 10.9146; verbatimLongitude: -85.4626; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larvae of the geometrid moth, Cyclophora Janzen23; verbatimEventDate: 26-Sep-2008; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

 Description

Female: 6mm (Fig. 5). Head (Fig. 5b): frontal setae extending to base of postpedicel; fronto-orbital plate gold, sparsely setulose along upper half; parafacial almost entirely gold (>90% coverage); palps black basally. Thorax (Fig. 5a, c): gold on both dorsal and lateral surfaces; four regular thoracic vittae; postpronotum with three setae; 2:3 acrostichal setae; 2:3 dorsocentral setae; 2:3 intra-alar setae; 2:3 supra-alar setae; three katepisternal setae; anatergite with three or more hair-like setae, often in a small tuft; scutellar discal setae absent. Wing vein R4+5 with 5–7 setulae dorsally, extending almost to crossvein R-M. Abdomen (Fig. 5a): ground color black, typically with uninterrupted transverse marginal pollinose bands (these can sometimes appear interrupted by a thick median stripe under certain angles of light); both dorsal and ventral surfaces concolorous, silver; base of ST1+2 black lateroventrally; T3 with one pair of median marginal setae and one pair of discal setae; T4 with a pair of discal setae.

Male: not known at this time.

 Diagnosis

Calolydella aureofacies can be distinguished from all other species of Calolydella by the following combination of traits: parafacial mostly gold, female palps black basally, anatergite with three or more hair-like setae, and scutellar discal setae absent.

 Etymology

The specific epithet is derived from the Latin adjective "aurum", meaning gold, and "facies", meaning face, in reference to its gold parafacial and fronto-orbital plate.

 Distribution

Costa Rica, ACG, Guanacaste Province, Sendero Ponderosa, 1060m.

 Ecology

Calolydella aureofacies has been reared once from Cyclophora Janzen23 (Lepidoptera: Geometridae), in cloud forest.

 B0232C9F-35F3-5405-BBF6-6B8B5D15AB00Calolydella bicolorurn:lsid:zoobank.org:act:266003AB-9934-4077-92EC-A89D557EEE43 Fleming & Wood sp. n. Materials

Type status: Holotype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0054050; recordedBy: D.H. Janzen, W. Hallwachs & Sebastian Kleppe; individualID: DHJPAR0054050; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 13-SRNP-31573, BOLD:ACM2409, ASHYD3218-14; Taxon: scientificName: Calolydella bicolor; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: bicolor; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Bernales; verbatimElevation: 660; verbatimLatitude: 10.9835; verbatimLongitude: -85.4212; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Callicore lyca; verbatimEventDate: 26-Nov-2013; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

 Description

Female (Fig. 6a, b, c). Length: 7mm. Head (Fig. 6b): frontal setae extending to base of postpedicel; fronto-orbital plate gold throughout, ranging from bare to almost bare; parafacial silver throughout. Thorax (Fig. 6a, c): gold on dorsal surface, silver laterally (>50% coverage); vittae fused into two prominent dark stripes; postpronotum with three setae; 3:3 acrostichal setae; 2:3 dorsocentral setae; 1:3 intra-alar setae; 2:3 supra-alar setae; three katepisternal setae; anatergite with three or more hair-like setae, often in a small tuft; scutellar discal setae situated slightly closer together than subapical scutellar setae. Wing vein R4+5 with at most 2–3 setulae dorsally at base, not extending to crossvein R-M. Abdomen (Fig. 6a): ground color orange-yellow, with median dark stripe breaking up pollinose marginal banding; abdominal pollinosity gold dorsally, silver ventrally; base of ST1+2 black lateroventrally; T3 with a row of marginal setae and one pair of discal setae; T4 with one pair of discal setae.

Male: not known at this time.

 Diagnosis

Calolydella bicolor can be distinguished from all other species of Calolydella by the following combination of traits: fronto-orbital plate entirely gold, parafacial entirely silver, anatergite with three or more hair-like setae, and gold abdominal pollinose bands interrupted by a median dark stripe.

 Etymology

The specific epithet is derived from the Latin adjective “bicolor”, meaning two-toned, in reference to its gold fronto-orbital plate and silver parafacial.

 Distribution

Costa Rica, ACG, Guanacaste Province, Sendero Bernales, 660m.

 Ecology

Calolydella bicolor has been reared once from Callicore lyca (Doubleday, 1847) (Lepidoptera: Nymphalidae), in rain forest.

 9C934609-E254-5119-95CF-12AF43183EECCalolydella bifissusurn:lsid:zoobank.org:act:11FFAD38-EC3E-48AD-81FB-6F45B23276DE Fleming & Wood sp. n. Materials

Type status: Holotype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017132; recordedBy: D.H. Janzen, W. Hallwachs & Jose Perez; individualID: DHJPAR0017132; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 07-SRNP-40315, BOLD:AAA1939, ASTAP570-07; Taxon: scientificName: Calolydella bifissus; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: bifissus; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Rio Francia Arriba; verbatimElevation: 400; verbatimLatitude: 10.8967; verbatimLongitude: -85.29; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalidae butterfly, Dircenna klugii; verbatimEventDate: 20-Feb-2007; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0036636; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Pereira; individualID: DHJPAR0036636; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 09-SRNP-36303, BOLD:AAA1939, ASHYE1547-09; Taxon: scientificName: Calolydella bifissus; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: bifissus; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Cacao; locality: Area de Conservacion Guanacaste; verbatimLocality: Estacion Cacao; verbatimElevation: 1150; verbatimLatitude: 10.9269; verbatimLongitude: -85.4682; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the geometridae moth, Phyllodonta latrata; verbatimEventDate: 19-Jul-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: 07-SRNP-40315; recordedBy: D.H. Janzen, W. Hallwachs & Jose Perez; individualCount: 2, siblings of holotype; sex: 1M, 1F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 07-SRNP-40315; Taxon: scientificName: Calolydella bifissus; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: bifissus; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Rio Francia Arriba; verbatimElevation: 400; verbatimLatitude: 10.8967; verbatimLongitude: -85.29; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalidae butterfly, Dircenna klugii; verbatimEventDate: 20-Feb-2007; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimens

 Description

Male (Fig. 7a, b, c). Length: 5–6mm. Head (Fig. 7b): frontal setae extending to base of postpedicel; frontal vitta tapering apically; fronto-orbital plate gold throughout, with a single row of fine setulae outside of frontal setae; parafacial at least 50% silver pollinose. Thorax (Fig. 7a, c): gold on dorsal surface, silver laterally (>50% coverage); vittae fused into two prominent dark stripes; postpronotum with two setae (inner basal seta absent); 1:3 acrostichal setae; 2:2 dorsocentral setae; 1:3 intra-alar setae; 1:3 supra-alar setae; three katepisternal setae; anatergite with three or more hair-like setae, often in a small tuft; scutellar discal setae situated slightly closer together than subapical scutellar setae. Wing vein R4+5 with at most 2–3 setulae dorsally at base, not extending to crossvein R-M. Abdomen (Fig. 7a): ground color dark brown-orange with uninterrupted transverse marginal pollinose bands; pollinosity gold dorsally, silver ventrally, with an orange spot lateroventrally at base of ST1+2; T3 with one pair of median marginal setae and one pair of discal setae; T4 with one pair of discal setae. Terminalia (Fig. 8): sternite 5 (Fig. 8c) with a wide U-shaped median cleft, 0.46X the length of the sternite from lobe to apex; each lobe with a smaller protrusion along upper margin of median cleft; inner margin covered by light pollinosity, appearing slightly darker than surrounding cuticle; lobe of sternite entirely covered with short setae, of varying lengths. Cerci with a fused base and cleft apex, slightly pinched at center (Fig. 8b); cercus, in lateral view, broad and with a slight downward bend. Surstylus digitiform and straight (Fig. 8a), appearing subequal to length of cercus when viewed laterally; surstylus with short setae along entire length; tip of surstylus appearing curved inwards when viewed dorsally.

Female (Fig. 7d, e, f). Length: 5mm. As male, except for the following characters: fronto-orbital plate 3X as wide as in male; frontal vitta not tapered, apically 1.2X as wide as in male; two katepisternal setae.

 Diagnosis

Calolydella bifissus can be distinguished from all other species of Calolydella by the following combination of characters: two bold thoracic vittae, abdominal pollinosity concolorous on both dorsal and ventral surfaces, anatergite with a small tuft of hair-like setae, and abdominal ground color dark brown-orange.

 Etymology

The specific epithet is derived from the Latin adjective "bifissus", meaning cleft or cloven, in reference to cleft nature of the cerci, which are fused basally, a character state unique to this species.

 Distribution

Costa Rica, ACG, Alajuela and Guanacaste provinces, 400–1150m.

 Ecology

Calolydella bifissus has been reared twice, from two separate hosts, Dircenna kluggii (Geyer, 1837) (Lepidoptera: Nymphalidae) and Phyllodonta latrata (Guenée, 1857) (Lepidoptera: Geometridae), in rain forest and cloud forest ecosystems.

 225D2FB3-E1DE-5ADF-8113-9F07FA65E18ACalolydella crocataurn:lsid:zoobank.org:act:252596B5-9766-4D84-AD8D-60846458BA5E Fleming & Wood sp. n. Materials

Type status: Holotype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0053232; recordedBy: D.H. Janzen, W. Hallwachs & Ricardo Calero; individualID: DHJPAR0053232; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 13-SRNP-70888, BOLD:ACK3862, ASHYM2586-13; Taxon: scientificName: Calolydella crocata; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: crocata; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Estacion Quica; verbatimElevation: 470; verbatimLatitude: 10.997; verbatimLongitude: -85.3967; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the geometrid moth, Semaeopus Janzen17; verbatimEventDate: 25-Jun-2013; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0055883; recordedBy: D.H. Janzen, W. Hallwachs & Dinia Martinez; individualID: DHJPAR0055883; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 14-SRNP-71256, BOLD:ACK3862, ASHYH2615-14; Taxon: scientificName: Calolydella crocata; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: crocata; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Medrano; verbatimElevation: 380; verbatimLatitude: 11.016; verbatimLongitude: -85.3805; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the geometrid moth, Semaeopus Janzen05; verbatimEventDate: 02-Aug-2014; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

 Description

Male (Fig. 9a, b, c). Length: 6mm. Head (Fig. 9b): frontal setae extending to base of postpedicel; fronto-orbital plate entirely gold, and sparsely setulose only along upper half; parafacial silver. Thorax: (Fig. 9a, c) gold on both dorsal and lateral surfaces; thorax with four vittae visible only under certain angles of light; postpronotum with three setae; 3:3 acrostichal setae; 3:3 dorsocentral setae; 2:3 intra-alar setae; 2:3 supra-alar setae; three katepisternal setae; anatergite bare; scutellar discal setae absent. Wing vein R4+5 with 3–4 setulae dorsally at base. Abdomen (Fig. 9a): ground color black, with thin median dark stripe breaking up pollinose marginal banding; abdominal pollinosity gold dorsally, silver ventrally; base of ST1+2 black lateroventrally; T3 with one pair of median marginal setae and one pair of discal setae; T4 with one pair of discal setae. Terminalia (Fig. 10): sternite 5 (Fig. 10c) with two small lobes and a wide U-shaped median cleft, 0.39X the length of the sternite from lobe to apex; inner margin covered by dense pollinosity, appearing darker than surrounding cuticle; upper margin of sternite with many stout, outwardly-pointing setae. Cerci (Fig. 10b) separated by a narrow gap basally, widening to an oval about midway; cercus, in dorsal view, long and rounded and not significantly tapering from base to tip; cercus, in lateral view, straight and setose along basal half, with a slight bulb at its tip. Surstylus (Fig. 10a) short, 2/3 length of cercus, and knife-like, appearing sharply pointed when viewed laterally; apical half of surstylus with short setae; tip of surstylus slightly angled outwards when viewed dorsally.

Female: not known at this time.

 Diagnosis

Calolydella crocata can be distinguished from all other species of Calolydella by the following combination of traits: fronto-orbital plate mostly gold-pollinose but with some silver pollinosity and sparsely setulose, thorax with four vittae visible only under certain angles of light, anatergite bare, and abdomen with transverse marginal pollinose bands interrupted by a thin dark median stripe.

 Etymology

The specific epithet is derived from the Latin adjective "crocatum", meaning yellow, in reference to the gold pollinosity present over most of the body of the new species.

 Distribution

Costa Rica, ACG, Guanacaste Province, Estación Quica, 470m.

 Ecology

Calolydella crocata has been reared once from Semaeopus Janzen05 (Lepidoptera: Geometridae), in rain forest.

 AA76DCC1-1719-5158-AB02-79B904DA3629Calolydella destitutaurn:lsid:zoobank.org:act:93C849D2-BF4A-4086-95EE-D69A4898598B Fleming & Wood sp. n. Materials

Type status: Holotype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017799; recordedBy: D.H. Janzen, W. Hallwachs & Roster Moraga; individualID: DHJPAR0017799; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 01-SRNP-11844, BOLD:AAE7382, ASTAR510-07; Taxon: scientificName: Calolydella destituta; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: destituta; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste; verbatimLocality: Mena Central; verbatimElevation: 345; verbatimLatitude: 11.0299; verbatimLongitude: -85.4536; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the erebid moth, Ramphia albizona; verbatimEventDate: 20-Nov-2001; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017768; recordedBy: D.H. Janzen, W. Hallwachs & Roster Moraga; individualID: DHJPAR0017768; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 01-SRNP-11820, BOLD:AAE7382, ASTAR479-07; Taxon: scientificName: Calolydella destituta; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: destituta; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste; verbatimLocality: Mena Central; verbatimElevation: 345; verbatimLatitude: 11.0299; verbatimLongitude: -85.4536; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the erebid moth, Ramphia albizona; verbatimEventDate: 20-Nov-2001; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017771; recordedBy: D.H. Janzen, W. Hallwachs & Roster Moraga; individualID: DHJPAR0017771; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 01-SRNP-11844, BOLD:AAE7382, ASTAR482-07; Taxon: scientificName: Calolydella destituta; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: destituta; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste; verbatimLocality: Mena Central; verbatimElevation: 345; verbatimLatitude: 11.0299; verbatimLongitude: -85.4536; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the erebid moth, Ramphia albizona; verbatimEventDate: 20-Nov-2001; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017776; recordedBy: D.H. Janzen, W. Hallwachs & Roster Moraga; individualID: DHJPAR0017776; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 01-SRNP-11820, BOLD:AAE7382, ASTAR487-07; Taxon: scientificName: Calolydella destituta; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: destituta; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste; verbatimLocality: Mena Central; verbatimElevation: 345; verbatimLatitude: 11.0299; verbatimLongitude: -85.4536; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the erebid moth, Ramphia albizona; verbatimEventDate: 20-Nov-2001; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

 Description

Male (Fig. 11a, b, c). Length: 8–9mm. Head (Fig. 11b): frontal setae extending beyond base of postpedicel; fronto-orbital plate gold, sparsely setulose throughout; parafacial silver along lower half. Thorax (Fig. 11a, c): gold on dorsal surface, silver laterally (>50% coverage); four regular thoracic vittae; postpronotum with four setae; 3:3 acrostichal setae; 2:3 dorsocentral setae; 2:3 intra-alar setae; 2:3 supra-alar setae; three katepisternal setae; anatergite with three or more hair-like setae, often in a small tuft; scutellar discal setae situated as wide apart as subapical setae. Wing vein R4+5 with at most three small setulae dorsally at base. Abdomen (Fig. 11a): ground color dark orange-brown, with uninterrupted transverse marginal pollinose bands, and with an orange spot lateroventrally at base of ST1+2; T3 lacking median marginal and discal setae; T4 with one pair of discal setae. Terminalia (Fig. 12): sternite 5 (Fig. 12c) with two small lobes and a wide U-shaped median cleft, 0.37X the length of the sternite from lobe to apex; inner margin covered by dense pollinosity, appearing darker than surrounding cuticle; entire lobe of sternite with medium-length setae, all of equal length. Cerci (Fig. 12b), in dorsal view, separated by a gap, widening at apex; each cercus long and very slightly tapering from its already narrow base; cercus, when viewed laterally, setose along basal half, slender and slightly arched, slightly upturned apically. Surstylus (Fig. 12a) 5/6 the length of cercus, slender and digitiform when viewed laterally, with a sharp pinch where basal section meets apical section, 2X as wide as cercus; surstylus with short setae along its entire length; tip of surstylus curved inwards when viewed dorsally.

Female (Fig. 11d, e, f). Length: 5–6mm. As male, except for the following characters: fronto-orbital plate 2X as wide as in male; and T3 with one pair of median marginal setae and one pair of discal setae.

 Diagnosis

Calolydella destituta can be distinguished from all other species of Calolydella by the following combination of traits: anatergite with a small tuft of setae, pollinose bands on abdomen not interrupted by a dark median stripe, and T3 lacking median marginal setae.

 Etymology

The specific epithet is derived from the Latin adjective "destituta", meaning absent or lacking, in reference to the absence of median marginal setae on T3, a character state unique to this species.

 Distribution

Costa Rica, ACG, Guanacaste Province, 345.

 Ecology

Calolydella destituta has been reared four times from Ramphia albizona (Latreille, 1817) (Lepidoptera: Erebidae), in rain forest and dry-rain lowland intergrade.

 019C4F8D-9A94-53F0-A3E4-DC198518038ECalolydella discalisurn:lsid:zoobank.org:act:584603D5-5F1C-457A-B7FC-6746A9EDA163 Fleming & Wood sp. n. Materials

Type status: Holotype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017774; recordedBy: D.H. Janzen, W. Hallwachs & Androny Valdelomar; individualID: DHJPAR0017774; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 04-SRNP-196, BOLD:AAI6844, ASTAR485-07; Taxon: scientificName: Calolydella discalis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: discalis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector San Cristobal; locality: Area de Conservacion Guanacaste; verbatimLocality: Rio Blanco Abajo; verbatimElevation: 500; verbatimLatitude: 10.9004; verbatimLongitude: -85.3725; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the erebid moth, Turuptiana obliqua; verbatimEventDate: 04-Feb-2004; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017773; recordedBy: D.H. Janzen, W. Hallwachs & Androny Valdelomar; individualID: DHJPAR0017773; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 04-SRNP-211, BOLD:AAI6844, ASTAR484-07; Taxon: scientificName: Calolydella discalis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: discalis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector San Cristobal; locality: Area de Conservacion Guanacaste; verbatimLocality: Rio Blanco Abajo; verbatimElevation: 500; verbatimLatitude: 10.9004; verbatimLongitude: -85.3725; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the erebid moth, Turuptiana obliqua; verbatimEventDate: 03-Feb-2004; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

 Description

Male (Fig. 13a, b, c). Length: 8–9mm. Head (Fig. 13b): frontal setae extending to base of postpedicel; fronto-orbital plate gold, with a single row of short setulae outside of frontal setae; parafacial at least 50% silver pollinose. Thorax (Fig. 13a, c): gold on both dorsal and lateral surfaces; outermost two thoracic vittae twice as wide as innermost two; scutellar discal setae situated as wide apart as subapical scutellar setae; postpronotum with three setae; 3:3 acrostichal setae; 2:3 dorsocentral setae; 3:3 intra-alar setae; 2:3 supra-alar setae; three katepisternal setae; anatergite with three or more hair-like setae, often in a small tuft. Wing vein R4+5 with 4–5 small setulae dorsally at base. Abdomen (Fig. 13a): ground color dark brown-orange with uninterrupted transverse marginal pollinose bands; abdominal pollinosity gold dorsally, silver ventrally, and with an orange spot lateroventrally at base of ST1+2; T3 with one pair of median marginal setae pair and two pairs of discal setae; T4 with two pairs of discal setae. Terminalia (Fig. 14): sternite 5 (Fig. 14c) with two small lobes and a wide U-shaped median cleft, 0.37X the length of the sternite from lobe to apex; inner margin covered by dense pollinosity, appearing darker than surrounding cuticle; entire lobe of sternite covered with short setae, all of equal length. Cerci (Fig. 14b), in dorsal view, narrow and tapered, separated by a narrow gap widening at apex; cercus long, slender and straight when viewed laterally, very slightly tapered from its already narrow base; setose along its basal half. Surstylus (Fig. 14a) subequal to length to cercus, slender and digitiform when viewed laterally; with short setae along entire length; tip of surstylus not curved inwards when viewed dorsally.

Female: not known at this time.

 Diagnosis

Calolydella discalis can be distinguished from all other species of Calolydella by the following combination of traits: parafacial mostly gold, fronto-orbital plate with a single row of small black setulae outside of the frontal setae, anatergite with a small tuft of three or more hair-like setae, and T3 and T4 each with two pairs of discal setae.

 Etymology

The specific epithet refers to the two pairs of discal setae on T3 and T4.

 Distribution

Costa Rica, ACG, Alajuela Province, Rio Blanco Abajo, 500m.

 Ecology

Calolydella discalis has been reared twice from Turuptiana obliqua Walker, 1869 (Lepidoptera: Erebidae: Arctiinae), in rain forest.

 A4265A47-67D8-58B4-A3A4-30C728FB18C8Calolydella erasmocoronadoiurn:lsid:zoobank.org:act:FAC2D446-637C-42C4-B744-69491257D313 Fleming & Wood sp. n. Materials

Type status: Holotype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0039246; recordedBy: D.H. Janzen, W. Hallwachs & Lucia Rios; individualID: DHJPAR0039246; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 10-SRNP-20642, BOLD:AAC8991, ASTAV809-10; Taxon: scientificName: Calolydella erasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste; verbatimLocality: Monte Cristo; verbatimElevation: 525; verbatimLatitude: 11.0137; verbatimLongitude: -85.4253; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe; verbatimEventDate: 14-May-2010; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011706; recordedBy: D.H. Janzen, W. Hallwachs & Osvaldo Espinoza; individualID: DHJPAR0011706; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 04-SRNP-2399, BOLD:AAC8991, ASTAS432-06; Taxon: scientificName: Calolydella erasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector San Cristobal; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Huerta; verbatimElevation: 527; verbatimLatitude: 10.9305; verbatimLongitude: -85.3722; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe; verbatimEventDate: 17-Jun-2004; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0037554; recordedBy: D.H. Janzen, W. Hallwachs & Noe Castillo; individualID: DHJPAR0037554; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 09-SRNP-69806, BOLD:AAC8991, ASHYC4299-10; Taxon: scientificName: Calolydella erasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Jacobo; verbatimElevation: 461; verbatimLatitude: 10.9408; verbatimLongitude: -85.3177; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe; verbatimEventDate: 10-Sep-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011688; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: DHJPAR0011688; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 04-SRNP-55482, BOLD:AAC8991, ASTAS414-06; Taxon: scientificName: Calolydella erasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Mismo; verbatimElevation: 680; verbatimLatitude: 10.9876; verbatimLongitude: -85.4197; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe; verbatimEventDate: 19-Oct-2004; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0039244; recordedBy: D.H. Janzen, W. Hallwachs & Lucia Rios; individualID: DHJPAR0039244; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 10-SRNP-20643, BOLD:AAC8991, ASTAV807-10; Taxon: scientificName: Calolydella erasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste; verbatimLocality: Monte Cristo; verbatimElevation: 525; verbatimLatitude: 11.0137; verbatimLongitude: -85.4253; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe; verbatimEventDate: 11-May-2010; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0039242; recordedBy: D.H. Janzen, W. Hallwachs & Lucia Rios; individualID: DHJPAR0039242; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 10-SRNP-20622, , ASTAV805-10; Taxon: scientificName: Calolydella erasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste; verbatimLocality: Monte Cristo; verbatimElevation: 525; verbatimLatitude: 11.0137; verbatimLongitude: -85.4253; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe; verbatimEventDate: 11-May-2010; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0039245; recordedBy: D.H. Janzen, W. Hallwachs & Lucia Rios; individualID: DHJPAR0039245; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 10-SRNP-20618, BOLD:AAC8991, ASTAV808-10; Taxon: scientificName: Calolydella erasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste; verbatimLocality: Monte Cristo; verbatimElevation: 525; verbatimLatitude: 11.0137; verbatimLongitude: -85.4253; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe; verbatimEventDate: 11-May-2010; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0037556; recordedBy: D.H. Janzen, W. Hallwachs & Noe Castillo; individualID: DHJPAR0037556; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 09-SRNP-69793, BOLD:AAC8991, ASHYC4301-10; Taxon: scientificName: Calolydella erasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Jacobo; verbatimElevation: 461; verbatimLatitude: 10.9408; verbatimLongitude: -85.3177; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe; verbatimEventDate: 10-Sep-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0037555; recordedBy: D.H. Janzen, W. Hallwachs & Noe Castillo; individualID: DHJPAR0037555; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 09-SRNP-69805, BOLD:AAC8991, ASHYC4300-10; Taxon: scientificName: Calolydella erasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Jacobo; verbatimElevation: 461; verbatimLatitude: 10.9408; verbatimLongitude: -85.3177; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe; verbatimEventDate: 10-Sep-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0039243; recordedBy: D.H. Janzen, W. Hallwachs & Lucia Rios; individualID: DHJPAR0039243; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 10-SRNP-20621, BOLD:AAC8991, ASTAV806-10; Taxon: scientificName: Calolydella erasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste; verbatimLocality: Monte Cristo; verbatimElevation: 525; verbatimLatitude: 11.0137; verbatimLongitude: -85.4253; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe; verbatimEventDate: 11-May-2010; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011686; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: DHJPAR0011686; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 04-SRNP-55195, BOLD:AAC8991, ASTAS412-06; Taxon: scientificName: Calolydella erasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Mismo; verbatimElevation: 680; verbatimLatitude: 10.9876; verbatimLongitude: -85.4197; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe; verbatimEventDate: 02-Oct-2004; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011687; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: DHJPAR0011687; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 04-SRNP-55483, BOLD:AAC8991, ASTAS413-06; Taxon: scientificName: Calolydella erasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Mismo; verbatimElevation: 680; verbatimLatitude: 10.9876; verbatimLongitude: -85.4197; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe; verbatimEventDate: 20-Oct-2004; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011685; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: DHJPAR0011685; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 04-SRNP-55194, BOLD:AAC8991, ASTAS411-06; Taxon: scientificName: Calolydella erasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Mismo; verbatimElevation: 680; verbatimLatitude: 10.9876; verbatimLongitude: -85.4197; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe; verbatimEventDate: 03-Oct-2004; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0039251; recordedBy: D.H. Janzen, W. Hallwachs & Lucia Rios; individualID: DHJPAR0039251; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 10-SRNP-20639, BOLD:AAC8991, ASTAV814-10; Taxon: scientificName: Calolydella erasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste; verbatimLocality: Monte Cristo; verbatimElevation: 525; verbatimLatitude: 11.0137; verbatimLongitude: -85.4253; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe; verbatimEventDate: 11-May-2010; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0039250; recordedBy: D.H. Janzen, W. Hallwachs & Lucia Rios; individualID: DHJPAR0039250; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 10-SRNP-20619, BOLD:AAC8991, ASTAV813-10; Taxon: scientificName: Calolydella erasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste; verbatimLocality: Monte Cristo; verbatimElevation: 525; verbatimLatitude: 11.0137; verbatimLongitude: -85.4253; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe; verbatimEventDate: 11-May-2010; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011707; recordedBy: D.H. Janzen, W. Hallwachs & Osvaldo Espinoza; individualID: DHJPAR0011707; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 04-SRNP-2393, BOLD:AAC8991, ASTAS433-06; Taxon: scientificName: Calolydella erasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector San Cristobal; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Huerta; verbatimElevation: 527; verbatimLatitude: 10.9305; verbatimLongitude: -85.3722; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe; verbatimEventDate: 03-Jun-2004; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011689; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: DHJPAR0011689; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 04-SRNP-55450, BOLD:AAC8991, ASTAS415-06; Taxon: scientificName: Calolydella erasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Mismo; verbatimElevation: 680; verbatimLatitude: 10.9876; verbatimLongitude: -85.4197; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe; verbatimEventDate: 18-Oct-2004; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0039248; recordedBy: D.H. Janzen, W. Hallwachs & Lucia Rios; individualID: DHJPAR0039248; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 10-SRNP-20624, BOLD:AAC8991, ASTAV811-10; Taxon: scientificName: Calolydella erasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste; verbatimLocality: Monte Cristo; verbatimElevation: 525; verbatimLatitude: 11.0137; verbatimLongitude: -85.4253; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe; verbatimEventDate: 11-May-2010; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0037553; recordedBy: D.H. Janzen, W. Hallwachs & Noe Castillo; individualID: DHJPAR0037553; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 09-SRNP-69808, BOLD:AAC8991, ASHYC4298-10; Taxon: scientificName: Calolydella erasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Jacobo; verbatimElevation: 461; verbatimLatitude: 10.9408; verbatimLongitude: -85.3177; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe; verbatimEventDate: 08-Sep-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011708; recordedBy: D.H. Janzen, W. Hallwachs & Osvaldo Espinoza; individualID: DHJPAR0011708; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 04-SRNP-2401, BOLD:AAC8991, ASTAS434-06; Taxon: scientificName: Calolydella erasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector San Cristobal; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Huerta; verbatimElevation: 527; verbatimLatitude: 10.9305; verbatimLongitude: -85.3722; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe; verbatimEventDate: 17-Jun-2004; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0039247; recordedBy: D.H. Janzen, W. Hallwachs & Lucia Rios; individualID: DHJPAR0039247; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 10-SRNP-20640, BOLD:AAC8991, ASTAV810-10; Taxon: scientificName: Calolydella erasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste; verbatimLocality: Monte Cristo; verbatimElevation: 525; verbatimLatitude: 11.0137; verbatimLongitude: -85.4253; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe; verbatimEventDate: 14-May-2010; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0039249; recordedBy: D.H. Janzen, W. Hallwachs & Lucia Rios; individualID: DHJPAR0039249; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 10-SRNP-20623, BOLD:AAC8991, ASTAV812-10; Taxon: scientificName: Calolydella erasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste; verbatimLocality: Quebrada Serrano; verbatimElevation: 585; verbatimLatitude: 11.0002; verbatimLongitude: -85.4561; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe; verbatimEventDate: 11-May-2010; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0037558; recordedBy: D.H. Janzen, W. Hallwachs & Noe Castillo; individualID: DHJPAR0037558; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 09-SRNP-69807, BOLD:AAC8991, ASHYC4303-10; Taxon: scientificName: Calolydella erasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Jacobo; verbatimElevation: 461; verbatimLatitude: 10.9408; verbatimLongitude: -85.3177; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe; verbatimEventDate: 14-Sep-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011684; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: DHJPAR0011684; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 04-SRNP-55193, BOLD:AAC8991, ASTAS410-06; Taxon: scientificName: Calolydella erasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Mismo; verbatimElevation: 680; verbatimLatitude: 10.9876; verbatimLongitude: -85.4197; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe; verbatimEventDate: 04-Oct-2004; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: 04-SRNP-55202; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: 04-SRNP-55202; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 04-SRNP-55202; Taxon: scientificName: Calolydella erasmocoronadoi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: erasmocoronadoi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Mismo; verbatimElevation: 680; verbatimLatitude: 10.9876; verbatimLongitude: -85.4197; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid moth, Euselasia chrysippe; verbatimEventDate: 04-Oct-2004; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

 Description

Male (Fig. 15a, b, c). Length: 6–8mm. Head (Fig. 15b): frontal setae extending to base of postpedicel; fronto-orbital plate up to 50% silver pollinose, almost bare, with a few small setulae interspersed among frontal setae; parafacial silver throughout. Thorax (Fig. 15a, c): gold on dorsal surface, silver laterally (>50% coverage); thoracic vittae visible only under certain angles of light; postpronotum with three setae; 3:3 acrostichal setae; 3:3 dorsocentral setae; 1:3 intra-alar setae; 2:3 supra-alar setae; three katepisternal setae; anatergite with three or more hair-like setae, often in a small tuft; scutellar discal setae situated as wide apart as subapical scutellar setae. Wing vein R4+5 with 6–7 setulae dorsally, extending halfway to crossvein R-M. Abdomen (Fig. 15a): ground color black with uninterrupted transverse gold pollinose bands; abdominal pollinosity gold dorsally, silver ventrally; base of ST1+2 black lateroventrally; T3 with one pair of median marginal setae and one pair of discal setae; T4 with one pair of discal setae. Terminalia (Fig. 16): sternite 5 (Fig. 16c) with two small lobes and a wide U-shaped median cleft, 0.34X the length of the sternite from lobe to apex; inner margin covered by dense pollinosity, appearing darker than surrounding cuticle; upper half of sternite with many stout, outwardly-pointing setae. Cerci (Fig. 16b), in dorsal view, separated by a narrow gap; each cercus long and rounded, and not significantly tapering from base to tip; cercus straight when viewed laterally, with a slight downward hook at its tip, setose almost along its entire length. Surstylus (Fig. 16c) stout and digitiform; apical half with short setae; tip of surstylus not lobed or angled inwards when viewed dorsally; subequal in length to cercus.

Female (Fig. 15d, e, f). Length: 4–7mm. Fronto-orbital plate 2.5X as wide as in male.

 Diagnosis

Calolydella erasmocoronadoi can be distinguished from all other species of Calolydella by the following combination of traits: fronto-orbital plate bare, up to 50% silver pollinose, thoracic vittae visible only under certain angles of light, anatergite with three or more setae arranged in a small tuft, wing vein R4+5 with 6–7 setulae, and abdomen with uninterrupted transverse marginal pollinose bands.

 Etymology

The specific epithet is in honor of Erasmo Coronado Caballo of Liberia, Costa Rica, in recognition of his high-quality assistantship to Sigifredo Marin of Liberia and to the ACG parataxonomists who collected and reared the caterpillars from which the new species of ACG Calolydella described here emerged.

 Distribution

Costa Rica, ACG, Guanacaste Province, Sendero Mismo, 680m.

 Ecology

Calolydella erasmocoronadoi has been reared 42 times from Euselasia chryssipe (Bates, 1866) (Lepidoptera: Riodinidae), in rain forest.

 25EA6662-AC46-5661-BDB2-7C83BD465AF9Calolydella felipechavarriaiurn:lsid:zoobank.org:act:538AEEF3-B6C0-458D-B5D3-518CBD726E0E Fleming & Wood sp. n. Materials

Type status: Holotype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0050491; recordedBy: D.H. Janzen, W. Hallwachs & Keiner Aragon; individualID: DHJPAR0050491; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 12-SRNP-79033, BOLD:ACJ3769, ACGBA3083-13; Taxon: scientificName: Calolydella felipechavarriai; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: felipechavarriai; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Palomo; verbatimElevation: 96; verbatimLatitude: 10.9619; verbatimLongitude: -85.2804; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the notodontid moth, Disphragis perses; verbatimEventDate: 08-Jan-2013; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0050489; recordedBy: D.H. Janzen, W. Hallwachs & Keiner Aragon; individualID: DHJPAR0050489; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 12-SRNP-79034, BOLD:ACJ3769, ACGBA3081-13; Taxon: scientificName: Calolydella felipechavarriai; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: felipechavarriai; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Palomo; verbatimElevation: 96; verbatimLatitude: 10.9619; verbatimLongitude: -85.2804; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the notodontid moth, Disphragis perses; verbatimEventDate: 10-Jan-2013; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: 12-SRNP-79034; recordedBy: D.H. Janzen, W. Hallwachs & Keiner Aragon; individualID: 12-SRNP-79034; individualCount: 3; sex: 1M, F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 12-SRNP-79034, BOLD:ACJ3769, ACGBA3081-13; Taxon: scientificName: Calolydella felipechavarriai; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: felipechavarriai; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Palomo; verbatimElevation: 96; verbatimLatitude: 10.9619; verbatimLongitude: -85.2804; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the notodontid moth, Disphragis perses; verbatimEventDate: 10-Jan-2013; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

 Description

Male (Fig. 17a, b, c). Length: 4–6mm. Head (Fig. 17b): frontal setae extending beyond base of postpedicel; fronto-orbital plate gold, almost bare, with a few small setulae interspersed among frontal setae; parafacial silver throughout. Thorax (Fig. 17a, c): gold on dorsal surface, silver laterally (>50% coverage); with four regular thoracic vittae; postpronotum with three setae; 3:3 acrostichal setae; 2:3 dorsocentral setae; 2:3 intra-alar setae; 2:3 supra-alar setae 2:3; three katepisternal setae; anatergite with two hair-like setae; scutellar discal setae slightly closer together than subapical scutellar setae. Wing vein R4+5 with 6–7 small setulae dorsally, extending to crossvein R-M. Abdomen (Fig. 17a): black in ground color, with uninterrupted transverse marginal pollinose bands; abdominal pollinosity gold dorsally, silver ventrally; base of ST1+2 black lateroventrally; T3 with one pair of median marginal setae and one pair of discal setae; T4 with one pair of discal setae. Terminalia (Fig. 18): sternite 5 (Fig. 18c) with two small lobes and a wide U-shaped median cleft, 0.31X the length of the sternite from lobe to apex; inner margin covered by dense pollinosity, appearing darker than surrounding cuticle; upper half of sternite with many stout, outwardly-pointing setae. Cerci (Fig. 18b), in dorsal view, separated by a narrow gap; each cercus long and rounded, and not significantly tapering from base to tip; cercus straight when viewed laterally, with a slight downward hook at its tip, setose almost along its entire length. Surstylus (Fig. 18a) stout and digitiform, bent midway, with a strong downward bias; apical half with short setae; tip of surstylus not lobed or angled inwards when viewed dorsally; subequal in length to cercus.

Female (Fig. 17d, e, f). Length: 4–5mm. Fronto-orbital plate 1.36X as wide as in male.

 Diagnosis

Calolydella felipechavarriai can be distinguished from all other species of Calolydella by the following combination of traits: fronto-orbital plate almost bare, anatergite with at most two hair-like setae, and T4 with one pair of discal setae.

 Etymology

The specific epithet is in honor of Luis Felipe Chavarria Diaz of ACG, Guanacaste, Costa Rica, in recognition of his high quality accounting, photography and administrative support of the Guanacaste Dry Forest Conservation Fund (www.gdfcf.org) and of the ACG parataxonomists who collected and reared the caterpillars from which the new species of ACG Calolydella described here emerged.

 Distribution

Costa Rica, ACG, Alajuela Province, Palomo, 96m.

 Ecology

Calolydella felipechavarriai has been reared twice from Bardaxima perses Druce, 1900 (Lepidoptera: Notodontidae), in rain forest.

 403BB166-90A2-556F-B4EF-A4E8598012B3Calolydella fredriksjobergiurn:lsid:zoobank.org:act:9E389552-FFEF-43B1-B371-081170A9EB6A Fleming & Wood sp. n. Materials

Type status: Holotype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011711; recordedBy: D.H. Janzen, W. Hallwachs & Asitente de Diana; individualID: DHJPAR0011711; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 03-SRNP-21135, BOLD:ABY7242, ASTAS437-06; Taxon: scientificName: Calolydella fredriksjobergi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: fredriksjobergi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Estacion Pitilla; verbatimElevation: 675; verbatimLatitude: 10.9893; verbatimLongitude: -85.4258; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larvae of the erebid moth, Dysschema jansonis; verbatimEventDate: 28-Sep-2003; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0054013; recordedBy: D.H. Janzen, W. Hallwachs & Cirilo Umana; individualID: DHJPAR0054013; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 13-SRNP-77901, BOLD:ABY7242, ASHYD3181-14; Taxon: scientificName: Calolydella fredriksjobergi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: fredriksjobergi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Finca Esmeralda; verbatimElevation: 123; verbatimLatitude: 10.9355; verbatimLongitude: -85.2531; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larvae of the nymphalid butterfly, Danaus plexippus; verbatimEventDate: 21-Nov-2013; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017784; recordedBy: D.H. Janzen, W. Hallwachs & gusaneros; individualID: DHJPAR0017784; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 99-SRNP-8296, BOLD:ABY7242, ASTAR495-07; Taxon: scientificName: Calolydella fredriksjobergi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: fredriksjobergi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Horizontes; locality: Area de Conservacion Guanacaste; verbatimLocality: Sitio La Dama; verbatimElevation: 105; verbatimLatitude: 10.7863; verbatimLongitude: -85.5583; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larvae of the nymphalid butterfly, Anaea aidea; verbatimEventDate: 17-Jun-1999; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0050643; recordedBy: D.H. Janzen, W. Hallwachs & Cirilo Umana; individualID: DHJPAR0050643; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 12-SRNP-77815, BOLD:ABY7242, ACGBA3235-13; Taxon: scientificName: Calolydella fredriksjobergi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: fredriksjobergi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Finca Esmeralda; verbatimElevation: 123; verbatimLatitude: 10.9355; verbatimLongitude: -85.2531; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larvae of the erebid moth, Melese Espinoza01; verbatimEventDate: 16-Dec-2012; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0035673; recordedBy: D.H. Janzen, W. Hallwachs & Mariano Pereira; individualID: DHJPAR0035673; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 09-SRNP-56992, BOLD:ABY7242, ASHYD1054-09; Taxon: scientificName: Calolydella fredriksjobergi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: fredriksjobergi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Mundo Nuevo; locality: Area de Conservacion Guanacaste; verbatimLocality: Camino Pozo Tres; verbatimElevation: 733; verbatimLatitude: 10.7708; verbatimLongitude: -85.3742; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larvae of the nymphalid butterfly, Chlosyne rosita; verbatimEventDate: 09-Aug-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0029600; recordedBy: D.H. Janzen, W. Hallwachs & Dunia Garcia; individualID: DHJPAR0029600; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 08-SRNP-36696, BOLD:ABY7242, ASHYM1021-09; Taxon: scientificName: Calolydella fredriksjobergi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: fredriksjobergi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Cacao; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Ponderosa; verbatimElevation: 1060; verbatimLatitude: 10.9146; verbatimLongitude: -85.4626; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larvae of the crambid moth, Cliniodes opalalis; verbatimEventDate: 02-Oct-2008; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

 Description

Male (Fig. 19a, b, c). Length: 7–9mm. Head (Fig. 19b): frontal setae extending to base of postpedicel; fronto-orbital plate gold, with a single row of fine setulae outside of frontal setae; parafacial silver throughout. Thorax (Fig. 19a, c): gold on both dorsal and lateral surfaces; outermost two thoracic vittae, twice as wide as innermost two; postpronotum with three setae; 3:3 acrostichal setae; 3:3 dorsocentral setae; 2:3 intra-alar setae; 2:3 supra-alar setae; three katepisternal setae; anatergite with three or more hair-like setae, often in a small tuft; scutellar discal setae situated as wide apart as subapical scutellar setae. Wing vein R4+5 with 6–7 setulae dorsally, reaching from base almost to crossvein R-M. Abdomen (Fig. 19a): ground color dark brown-orange, with uninterrupted transverse marginal pollinose bands; abdominal pollinosity gold dorsally, silver ventrally, with an orange spot lateroventrally at base of ST1+2; T3 with one pair of median marginal setae and two pairs of discal setae; T4 with a row of marginal setae and one pair of discal setae. Terminalia (Fig. 20): sternite 5 (Fig. 20c) with two small lobes and a wide U-shaped median cleft, 0.47X the length of the sternite from lobe to apex; inner margin covered by dense pollinosity, appearing darker than surrounding cuticle; entire lobe of sternite with short setae, of varying lengths. Cerci (Fig. 20b), in dorsal view, separated by a narrow gap not widened at apex; each cercus very slightly tapered from its already narrow base; cercus long, slender and with an upward bend when viewed laterally; setose along its entire length. Surstylus (Fig. 20a) slender and digitiform with a strong divot at tip, giving it an overall bent look; appearing 0.2X shorter than cercus when viewed laterally; with short setae along entire length; tip of surstylus slightly curved inwards when viewed dorsally.

Female (Fig. 19d, e, f). Length: 5–6mm. Fronto-orbital plate 2.2X wider than in male; thoracic vittae 0.5X as wide as in male.

 Diagnosis

Calolydella fredriksjobergi can be distinguished from all other species of Calolydella by the following combination of characters: parafacial mostly gold, anatergite with three or more hair-like setae arranged in a small tuft, two pairs of discal setae on T3 and one pair of discal setae on T4.

 Etymology

The specific epithet is in honor of Fredrik Sjoberg of Runmaro Island, Sweden, in recognition of his wonderful book "The Fly Trap" (Sjöberg 2014, translation by Thomas Teal); a book that explains much about Malaise traps, which have captured so many ACG Tachinidae.

 Distribution

Costa Rica, ACG, Alajuela and Guanacaste provinces, 105–1060m.

 Ecology

Calolydella fredriksjobergi has been reared six times from six different species belonging to various families of Lepidoptera (Nymphalidae; Erebidae; and Crambidae), in rain forest, cloud forest, and dry forest ecosystems.

 0AFB106C-F0B0-5C38-8C61-32EDC7DC9764Calolydella inflatipalpisurn:lsid:zoobank.org:act:BB67E463-DE23-474A-9900-9EC0193FC403 Fleming & Wood sp. n. Materials

Type status: Holotype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0016181; recordedBy: D.H. Janzen, W. Hallwachs & Jose Cortez; individualID: DHJPAR0016181; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-58499, BOLD:ABX6062, ASTAP210-06; Taxon: scientificName: Calolydella inflatipalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: inflatipalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Mundo Nuevo; locality: Area de Conservacion Guanacaste; verbatimLocality: Vado Licania; verbatimElevation: 470; verbatimLatitude: 10.7722; verbatimLongitude: -85.4122; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Dione juno; verbatimEventDate: 10-Oct-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017796; recordedBy: D.H. Janzen, W. Hallwachs & Mariano Pereira; individualID: DHJPAR0017796; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 02-SRNP-9198, BOLD:ABX6062, ASTAR507-07; Taxon: scientificName: Calolydella inflatipalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: inflatipalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Cacao; locality: Area de Conservacion Guanacaste; verbatimLocality: Estacion Cacao; verbatimElevation: 1150; verbatimLatitude: 10.9269; verbatimLongitude: -85.4682; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Dione juno; verbatimEventDate: 08-Jun-2002; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017806; recordedBy: D.H. Janzen, W. Hallwachs & Roster Moraga; individualID: DHJPAR0017806; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 00-SRNP-3991.32, BOLD:ABX6062, ASTAR517-07; Taxon: scientificName: Calolydella inflatipalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: inflatipalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector El Hacha; locality: Area de Conservacion Guanacaste; verbatimLocality: Casa Uno; verbatimElevation: 280; verbatimLatitude: 11.0056; verbatimLongitude: -85.5743; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Dione juno; verbatimEventDate: 21-Sep-2000; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0016179; recordedBy: D.H. Janzen, W. Hallwachs & Jose Cortez; individualID: DHJPAR0016179; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-58475, BOLD:ABX6062, ASTAP208-06; Taxon: scientificName: Calolydella inflatipalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: inflatipalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Mundo Nuevo; locality: Area de Conservacion Guanacaste; verbatimLocality: Vado Licania; verbatimElevation: 470; verbatimLatitude: 10.7722; verbatimLongitude: -85.4122; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Dione juno; verbatimEventDate: 10-Oct-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0016169; recordedBy: D.H. Janzen, W. Hallwachs & Jose Cortez; individualID: DHJPAR0016169; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-58472, BOLD:ABX6062, ASTAP198-06; Taxon: scientificName: Calolydella inflatipalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: inflatipalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Mundo Nuevo; locality: Area de Conservacion Guanacaste; verbatimLocality: Vado Licania; verbatimElevation: 470; verbatimLatitude: 10.7722; verbatimLongitude: -85.4122; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Dione juno; verbatimEventDate: 08-Oct-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017793; recordedBy: D.H. Janzen, W. Hallwachs & Mariano Pereira; individualID: DHJPAR0017793; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 02-SRNP-9197, BOLD:ABX6062, ASTAR504-07; Taxon: scientificName: Calolydella inflatipalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: inflatipalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Cacao; locality: Area de Conservacion Guanacaste; verbatimLocality: Estacion Cacao; verbatimElevation: 1150; verbatimLatitude: 10.9269; verbatimLongitude: -85.4682; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Dione juno; verbatimEventDate: 08-Jun-2002; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0016170; recordedBy: D.H. Janzen, W. Hallwachs & Jose Cortez; individualID: DHJPAR0016170; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-58481, BOLD:ABX6062, ASTAP199-06; Taxon: scientificName: Calolydella inflatipalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: inflatipalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Mundo Nuevo; locality: Area de Conservacion Guanacaste; verbatimLocality: Vado Licania; verbatimElevation: 470; verbatimLatitude: 10.7722; verbatimLongitude: -85.4122; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Dione juno; verbatimEventDate: 07-Oct-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017800; recordedBy: D.H. Janzen, W. Hallwachs & Roster Moraga; individualID: DHJPAR0017800; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 00-SRNP-3991.22, BOLD:ABX6062, ASTAR511-07; Taxon: scientificName: Calolydella inflatipalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: inflatipalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector El Hacha; locality: Area de Conservacion Guanacaste; verbatimLocality: Casa Uno; verbatimElevation: 280; verbatimLatitude: 11.0056; verbatimLongitude: -85.5743; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Dione juno; verbatimEventDate: 23-Sep-2000; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0016197; recordedBy: D.H. Janzen, W. Hallwachs & Roster Moraga; individualID: DHJPAR0016197; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-22670, BOLD:ABX6062, ASTAP226-06; Taxon: scientificName: Calolydella inflatipalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: inflatipalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste; verbatimLocality: San Antonio; verbatimElevation: 335; verbatimLatitude: 11.0353; verbatimLongitude: -85.4453; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Dione juno; verbatimEventDate: 28-Sep-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017801; recordedBy: D.H. Janzen, W. Hallwachs & Dunia Garcia; individualID: DHJPAR0017801; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 03-SRNP-25826, BOLD:ABX6062, ASTAR512-07; Taxon: scientificName: Calolydella inflatipalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: inflatipalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Santa Rosa; locality: Area de Conservacion Guanacaste; verbatimLocality: Bosque Humedo; verbatimElevation: 290; verbatimLatitude: 10.8514; verbatimLongitude: -85.608; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Dione juno; verbatimEventDate: 25-Sep-2003; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017802; recordedBy: D.H. Janzen, W. Hallwachs & gusaneros; individualID: DHJPAR0017802; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 95-SRNP-8266, BOLD:ABX6062, ASTAR513-07; Taxon: scientificName: Calolydella inflatipalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: inflatipalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Santa Rosa; locality: Area de Conservacion Guanacaste; verbatimLocality: Bosque San Emilio; verbatimElevation: 300; verbatimLatitude: 10.8439; verbatimLongitude: -85.6138; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Dione juno; verbatimEventDate: 09-Feb-1995; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017798; recordedBy: D.H. Janzen, W. Hallwachs & Roster Moraga; individualID: DHJPAR0017798; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 00-SRNP-3828, BOLD:ABX6062, ASTAR509-07; Taxon: scientificName: Calolydella inflatipalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: inflatipalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector El Hacha; locality: Area de Conservacion Guanacaste; verbatimLocality: Casa Uno; verbatimElevation: 280; verbatimLatitude: 11.0056; verbatimLongitude: -85.5743; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Dione juno; verbatimEventDate: 06-Sep-2000; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017804; recordedBy: D.H. Janzen, W. Hallwachs & Roster Moraga; individualID: DHJPAR0017804; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 00-SRNP-3991.03, BOLD:ABX6062, ASTAR515-07; Taxon: scientificName: Calolydella inflatipalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: inflatipalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector El Hacha; locality: Area de Conservacion Guanacaste; verbatimLocality: Casa Uno; verbatimElevation: 280; verbatimLatitude: 11.0056; verbatimLongitude: -85.5743; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Dione juno; verbatimEventDate: 20-Sep-2000; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0016098; recordedBy: D.H. Janzen, W. Hallwachs & Jose Cortez; individualID: DHJPAR0016098; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-58469, BOLD:ABX6062, ASTAP127-06; Taxon: scientificName: Calolydella inflatipalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: inflatipalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Mundo Nuevo; locality: Area de Conservacion Guanacaste; verbatimLocality: Vado Licania; verbatimElevation: 470; verbatimLatitude: 10.7722; verbatimLongitude: -85.4122; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Dione juno; verbatimEventDate: 03-Oct-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017803; recordedBy: D.H. Janzen, W. Hallwachs & Roster Moraga; individualID: DHJPAR0017803; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 00-SRNP-3842, BOLD:ABX6062, ASTAR514-07; Taxon: scientificName: Calolydella inflatipalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: inflatipalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector El Hacha; locality: Area de Conservacion Guanacaste; verbatimLocality: Casa Uno; verbatimElevation: 280; verbatimLatitude: 11.0056; verbatimLongitude: -85.5743; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Dione juno; verbatimEventDate: 06-Sep-2000; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017807; recordedBy: D.H. Janzen, W. Hallwachs & Roster Moraga; individualID: DHJPAR0017807; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 00-SRNP-3991.04, BOLD:ABX6062, ASTAR518-07; Taxon: scientificName: Calolydella inflatipalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: inflatipalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector El Hacha; locality: Area de Conservacion Guanacaste; verbatimLocality: Casa Uno; verbatimElevation: 280; verbatimLatitude: 11.0056; verbatimLongitude: -85.5743; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Dione juno; verbatimEventDate: 19-Sep-2000; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011704; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: DHJPAR0011704; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 05-SRNP-31118, BOLD:ABX6062, ASTAS430-06; Taxon: scientificName: Calolydella inflatipalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: inflatipalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Pasmompa; verbatimElevation: 440; verbatimLatitude: 11.0193; verbatimLongitude: -85.41; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Dryadula phaetusa; verbatimEventDate: 13-Apr-2005; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0016168; recordedBy: D.H. Janzen, W. Hallwachs & Jose Cortez; individualID: DHJPAR0016168; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-58471, BOLD:ABX6062, ASTAP197-06; Taxon: scientificName: Calolydella inflatipalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: inflatipalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Mundo Nuevo; locality: Area de Conservacion Guanacaste; verbatimLocality: Vado Licania; verbatimElevation: 470; verbatimLatitude: 10.7722; verbatimLongitude: -85.4122; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Dione juno; verbatimEventDate: 07-Oct-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0029990; recordedBy: D.H. Janzen, W. Hallwachs & Elda Araya; individualID: DHJPAR0029990; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 08-SRNP-5495, BOLD:ABX6062, ASHYB734-09; Taxon: scientificName: Calolydella inflatipalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: inflatipalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Buenos Aires; locality: Area de Conservacion Guanacaste; verbatimLocality: Finca Tomate; verbatimElevation: 360; verbatimLatitude: 10.9035; verbatimLongitude: -85.3092; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Dione juno; verbatimEventDate: 02-Nov-2008; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017797; recordedBy: D.H. Janzen, W. Hallwachs & Roster Moraga; individualID: DHJPAR0017797; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 00-SRNP-3829, BOLD:ABX6062, ASTAR508-07; Taxon: scientificName: Calolydella inflatipalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: inflatipalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector El Hacha; locality: Area de Conservacion Guanacaste; verbatimLocality: Casa Uno; verbatimElevation: 280; verbatimLatitude: 11.0056; verbatimLongitude: -85.5743; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Dione juno; verbatimEventDate: 05-Sep-2000; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0015983; recordedBy: D.H. Janzen, W. Hallwachs & Mariano Pereira; individualID: DHJPAR0015983; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-58420, BOLD:ABX6062, ASTAP012-06; Taxon: scientificName: Calolydella inflatipalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: inflatipalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Mundo Nuevo; locality: Area de Conservacion Guanacaste; verbatimLocality: Vado Licania; verbatimElevation: 470; verbatimLatitude: 10.7722; verbatimLongitude: -85.4122; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Dione juno; verbatimEventDate: 06-Oct-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0016196; recordedBy: D.H. Janzen, W. Hallwachs & Elieth Cantillano; individualID: DHJPAR0016196; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-22687, BOLD:ABX6062, ASTAP225-06; Taxon: scientificName: Calolydella inflatipalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: inflatipalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste; verbatimLocality: San Antonio; verbatimElevation: 335; verbatimLatitude: 11.0353; verbatimLongitude: -85.4453; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Dione juno; verbatimEventDate: 25-Sep-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0015972; recordedBy: D.H. Janzen, W. Hallwachs & Jose Cortez; individualID: DHJPAR0015972; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-58451, BOLD:ABX6062, ASTAP001-06; Taxon: scientificName: Calolydella inflatipalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: inflatipalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Mundo Nuevo; locality: Area de Conservacion Guanacaste; verbatimLocality: Vado Licania; verbatimElevation: 470; verbatimLatitude: 10.7722; verbatimLongitude: -85.4122; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Dione juno; verbatimEventDate: 09-Oct-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011691; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: DHJPAR0011691; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 04-SRNP-33986, BOLD:ABX6062, ASTAS417-06; Taxon: scientificName: Calolydella inflatipalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: inflatipalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Loaiciga; verbatimElevation: 445; verbatimLatitude: 11.0198; verbatimLongitude: -85.4134; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Dryas iulia; verbatimEventDate: 14-Aug-2004; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0015975; recordedBy: D.H. Janzen, W. Hallwachs & Jose Cortez; individualID: DHJPAR0015975; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-58470, BOLD:ABX6062, ASTAP004-06; Taxon: scientificName: Calolydella inflatipalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: inflatipalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Mundo Nuevo; locality: Area de Conservacion Guanacaste; verbatimLocality: Vado Licania; verbatimElevation: 470; verbatimLatitude: 10.7722; verbatimLongitude: -85.4122; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Dione juno; verbatimEventDate: 03-Oct-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0024555; recordedBy: D.H. Janzen, W. Hallwachs & Duvalier Briceno; individualID: DHJPAR0024555; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 08-SRNP-65294, BOLD:ABX6062, ASTAW665-08; Taxon: scientificName: Calolydella inflatipalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: inflatipalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Brasilia; locality: Area de Conservacion Guanacaste; verbatimLocality: Piedrona; verbatimElevation: 340; verbatimLatitude: 11.0162; verbatimLongitude: -85.359; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Heliconius charithonia; verbatimEventDate: 15-Apr-2008; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017805; recordedBy: D.H. Janzen, W. Hallwachs & Roster Moraga; individualID: DHJPAR0017805; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 00-SRNP-3991.10, BOLD:ABX6062, ASTAR516-07; Taxon: scientificName: Calolydella inflatipalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: inflatipalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector El Hacha; locality: Area de Conservacion Guanacaste; verbatimLocality: Casa Uno; verbatimElevation: 280; verbatimLatitude: 11.0056; verbatimLongitude: -85.5743; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Dione juno; verbatimEventDate: 20-Sep-2000; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0016185; recordedBy: D.H. Janzen, W. Hallwachs & Jose Cortez; individualID: DHJPAR0016185; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-58496, BOLD:ABX6062, ASTAP214-06; Taxon: scientificName: Calolydella inflatipalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: inflatipalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Mundo Nuevo; locality: Area de Conservacion Guanacaste; verbatimLocality: Vado Licania; verbatimElevation: 470; verbatimLatitude: 10.7722; verbatimLongitude: -85.4122; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Dione juno; verbatimEventDate: 12-Oct-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017794; recordedBy: D.H. Janzen, W. Hallwachs & Mariano Pereira; individualID: DHJPAR0017794; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 02-SRNP-9224, BOLD:ABX6062, ASTAR505-07; Taxon: scientificName: Calolydella inflatipalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: inflatipalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Cacao; locality: Area de Conservacion Guanacaste; verbatimLocality: Estacion Cacao; verbatimElevation: 1150; verbatimLatitude: 10.9269; verbatimLongitude: -85.4682; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Dione juno; verbatimEventDate: 05-Jun-2002; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017795; recordedBy: D.H. Janzen, W. Hallwachs & Mariano Pereira; individualID: DHJPAR0017795; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 02-SRNP-9199, BOLD:ABX6062, ASTAR506-07; Taxon: scientificName: Calolydella inflatipalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: inflatipalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Cacao; locality: Area de Conservacion Guanacaste; verbatimLocality: Estacion Cacao; verbatimElevation: 1150; verbatimLatitude: 10.9269; verbatimLongitude: -85.4682; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Dione juno; verbatimEventDate: 08-Jun-2002; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

 Description

Male (Fig. 21a, b, c). Length: 5–9mm. Head (Fig. 21b, e): frontal setae extending beyond base of postpedicel; fronto-orbital plate gold, sparsely setulose throughout; parafacial almost entirely gold (75-90% coverage). Thorax (Fig. 21a, c, d, f): pollinosity gold on both dorsal and lateral surfaces; or gold on dorsal surface and silver laterally (more than 50% coverage); thorax with outermost two vittae twice as wide as innermost two; postpronotum with three setae; 3:3 acrostichal setae; 3:3 dorsocentral setae; 2:3 intra-alar setae; 2:3 supra-alar setae; three katepisternal setae; anatergite with three or more hair-like setae, often in a small tuft; scutellar discal setae situated as wide apart as subapical scutellar setae. Wing vein R4+5 with at most 2–3 small setulae dorsally at base. Abdomen (Fig. 21a, d): ground color dark brown-orange; with uninterrupted transverse marginal pollinose bands; pollinosity gold dorsally, silver ventrally, and with an orange spot lateroventrally at base of ST1+2; T3 with one pair of median marginal setae and two pairs of discal setae; T4 with two pairs of discal setae. Terminalia (Fig. 22): sternite 5 (Fig. 22c) with two small lobes and a wide U-shaped median cleft, 0.37X the length of the sternite from lobe to apex; inner margin covered by dense pollinosity, appearing darker than surrounding cuticle; entire lobe of sternite with sparse short setae, of varying lengths. Cerci (Fig. 22b), in dorsal view, separated by a narrow gap widening at apex; slender and straight; cercus, in lateral view, long, very slightly tapered from its already narrow base, setose along its basal half. Surstylus (Fig. 22a) subequal to length of cercus, turned slightly inward at tip, appearing as a small apical lobe when viewed dorsally; surstylus slender and digitiform when viewed laterally; with short setae along entire length; tip of surstylus not curved inwards when viewed dorsally.

Female (Fig. 21d, e, f). Length: 5–7mm. Fronto-orbital plate 1.6X as wide as in male; palpus significantly inflated apically.

 Diagnosis

Calolydella inflatipalpis can be distinguished from all other species of Calolydella by the following combination of traits: fine setulae interspersed among frontal setae, palps normal in male but significantly inflated in female, anatergite with three or more hair-like setae arranged in a small tuft, and T3 and T4 each with two pairs of discal setae.

 Etymology

The specific epithet is derived from the Latin adjective “inflatum”, meaning inflated, and "palpus", meaning palp, in reference to the inflated, bulbous palpi of females of this species.

 Distribution

Costa Rica, ACG, Alajuela and Guanacaste provinces, 290–1150m.

 Ecology

Calolydella inflatipalpis has been reared 31 times from four separate species of Heliconiini: Dione juno (Cramer, 1779), Dryas iulia (Fab., 1775), Dryadula phaetusa (Linn., 1758), and Heliconius charithonia (Linn., 1767) (Lepidoptera: Nymphalidae), in rain forest, dry forest, cloud forest and dry-rain lowland intergrade ecosystems.

 CB4AF84B-C274-5A99-A599-C86E5A64A657Calolydella interruptaurn:lsid:zoobank.org:act:1B9143F7-AA12-4105-9EF3-B5BA3B836D20 Fleming & Wood sp. n. Materials

Type status: Holotype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0029598; recordedBy: D.H. Janzen, W. Hallwachs & Dunia Garcia; individualID: DHJPAR0029598; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 08-SRNP-36564, BOLD:AAW8657, ASHYM1019-09; Taxon: scientificName: Calolydella interrupta; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: interrupta; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Cacao; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Arenales; verbatimElevation: 1080; verbatimLatitude: 10.9247; verbatimLongitude: -85.4674; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the limacodid moth, Acharia ophelians; verbatimEventDate: 17-Sep-2008; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: 08-SRNP-36564; recordedBy: D.H. Janzen, W. Hallwachs & Dunia Garcia; individualID: 08-SRNP-36564; individualCount: 1, sibling of holotype; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 08-SRNP-36564; Taxon: scientificName: Calolydella interrupta; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: interrupta; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Cacao; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Arenales; verbatimElevation: 1080; verbatimLatitude: 10.9247; verbatimLongitude: -85.4674; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the limacodid moth, Acharia ophelians; verbatimEventDate: 17-Sep-2008; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

 Description

Male (Fig. 23a, b, c). Length: 8mm. Head (Fig. 23b): frontal setae extending to base of postpedicel; fronto-orbital plate gold, ranging from bare to almost bare; parafacial silver along lower half. Thorax (Fig. 23a, c): gold on dorsal surface, silver laterally (>50% coverage); with two bold thoracic vittae; postpronotum with three setae; 2:3 acrostichal setae; 2:3 dorsocentral setae; 2:3 intra-alar setae; 1:3 supra-alar setae; three katepisternal setae; anatergite with three or more hair-like setae, often in a small tuft; scutellar discal setae slightly closer together than subapical scutellar setae. Wing vein R4+5 with 2–4 setulae dorsally, confined to base of wing vein, not extending up to crossvein R-M. Abdomen (Fig. 23a): ground color dark orange, with a median dark stripe breaking up pollinose marginal banding; abdominal pollinosity gold dorsally, silver ventrally; base of ST1+2 black lateroventrally, mostly gold pollinose dorsally; T3 with a row of marginal setae and one pair of discal setae; T4 with one pair of discal setae. Terminalia: not examined.

Female (Fig. 23d, e, f). Length: 6mm. Fronto-orbital plate 2X as wide as in male.

 Diagnosis

Calolydella interrupta can be distinguished from all other species of Calolydella by the following combination of traits: parafacial at least 50% silver pollinose, two bold thoracic vittae, abdominal pollinosity interrupted by median stripe, anatergite with three or more setae arranged in a small tuft, and T3 with a complete row of marginal setae.

 Etymology

The specific epithet is derived from the Latin adjective “interruptum", meaning breach or interruption, in reference to dark median stripe breaking up the gold marginal pollinosity on the abdomen of this species.

 Distribution

Costa Rica, ACG, Guanacaste Province, Sendero Arenales 1080m.

 Ecology

Calolydella interrupta has been reared once from Acharia ophelians (Dyar, 1927) (Lepidoptera: Limacodidae), in cloud forest.

 0411C249-CEBD-5B7E-8F60-AEA85739DD5ECalolydella nigripalpisurn:lsid:zoobank.org:act:612EE188-6B86-49E4-BD72-EBCAE03A2D91 Fleming & Wood sp. n. Materials

Type status: Holotype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017812; recordedBy: D.H. Janzen, W. Hallwachs & Carolina Cano; individualID: DHJPAR0017812; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 05-SRNP-1172, BOLD:ABY2592, ASTAR523-07; Taxon: scientificName: Calolydella nigripalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: nigripalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector San Cristobal; locality: Area de Conservacion Guanacaste; verbatimLocality: Puente Palma; verbatimElevation: 460; verbatimLatitude: 10.9163; verbatimLongitude: -85.3787; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the geometrid moth, Semaeopus Janzen08; verbatimEventDate: 04-Apr-2005; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0040836; recordedBy: D.H. Janzen, W. Hallwachs & Ricardo Calero; individualID: DHJPAR0040836; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 10-SRNP-73126, BOLD:ABY2592, ASHYF751-11; Taxon: scientificName: Calolydella nigripalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: nigripalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Calma; verbatimElevation: 412; verbatimLatitude: 11.0099; verbatimLongitude: -85.3921; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the geometrid moth, Semaeopus Janzen08; verbatimEventDate: 20-Nov-2010; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0052029; recordedBy: D.H. Janzen, W. Hallwachs & Edwin Apu; individualID: DHJPAR0052029; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 13-SRNP-69658, BOLD:ABY2592, ASHYH1141-13; Taxon: scientificName: Calolydella nigripalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: nigripalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Jacobo; verbatimElevation: 461; verbatimLatitude: 10.9408; verbatimLongitude: -85.3177; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the geometrid moth, Semaeopus Janzen08; verbatimEventDate: 26-May-2013; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017811; recordedBy: D.H. Janzen, W. Hallwachs & Yessenia Mendoza; individualID: DHJPAR0017811; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 05-SRNP-4251, BOLD:ABY2592, ASTAR522-07; Taxon: scientificName: Calolydella nigripalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: nigripalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector San Cristobal; locality: Area de Conservacion Guanacaste; verbatimLocality: Potrero Argentina; verbatimElevation: 520; verbatimLatitude: 10.8902; verbatimLongitude: -85.388; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the geometrid moth, Semaeopus Janzen08; verbatimEventDate: 20-Aug-2005; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0054198; recordedBy: D.H. Janzen, W. Hallwachs & Cirilo Umana; individualID: DHJPAR0054198; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 14-SRNP-75117, BOLD:ABY2592, ASHYD3366-14; Taxon: scientificName: Calolydella nigripalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: nigripalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Estacion Llanura; verbatimElevation: 135; verbatimLatitude: 10.9333; verbatimLongitude: -85.2533; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the geometrid moth, Semaeopus Janzen08; verbatimEventDate: 29-Jan-2014; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0054195; recordedBy: D.H. Janzen, W. Hallwachs & Cirilo Umana; individualID: DHJPAR0054195; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 14-SRNP-75120, BOLD:ABY2592, ASHYD3363-14; Taxon: scientificName: Calolydella nigripalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: nigripalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Estacion Llanura; verbatimElevation: 135; verbatimLatitude: 10.9333; verbatimLongitude: -85.2533; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the geometrid moth, Semaeopus Janzen08; verbatimEventDate: 01-Feb-2014; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

 Description

Male (Fig. 24a, b, c). Length: 5–6mm. Head (Fig. 24b): frontal setae extending to base of postpedicel; fronto-orbital plate gold, sparsely setulose throughout; parafacial silver throughout; palps black. Thorax (Fig. 24a, c): gold on both dorsal and lateral surfaces; four almost indistinct thoracic vittae barely visible except under certain angles of light; postpronotum with three setae; 3:3 acrostichal setae; 3:3 dorsocentral setae; 2:3 intra-alar setae; 2:3 supra-alar setae; three katepisternal setae; anatergite bare; scutellar discal setae situated as wide apart as subapical scutellar setae. Wing vein R4+5 with 6–7 setulae dorsally, reaching from the base almost to crossvein R-M. Abdomen (Fig. 24a): ground color black, with uninterrupted transverse marginal pollinose bands; pollinosity gold dorsally, silver ventrally; base of ST1+2 black lateroventrally; T3 with one pair of median marginal setae and two pairs of discal setae; T4 with a row of marginal setae and two pairs of discal setae. Terminalia (Fig. 25): sternite 5 (Fig. 25c) with two small lobes and a wide U-shaped median cleft, 0.62X the length of the sternite from lobe to apex; inner margin covered by dense pollinosity, appearing darker than surrounding cuticle; with 4–6 short setae, of varying lengths. Cerci (Fig. 25b), in dorsal view, separated by a narrow gap, divergent at apex; each cercus sharply tapered, 2.2X broader at base than at apex, digitiform when viewed dorsally; cercus with a very slight downward bend at tip when viewed laterally; setose along its entire length. Surstylus (Fig. 25a) digitiform, with a strong apical downward curve giving a hooked tip, appearing 0.2X longer than cercus when viewed laterally; with short setae along entire length; tip of surstylus appearing curved inwards when viewed dorsally.

Female: not known at this time.

 Diagnosis

Calolydella nigripalpis can be distinguished from all other species of Calolydella by the following combination of characters: frontal setae not extending beyond base of postpedicel, palps black in male, four barely visible thoracic vittae, anatergite bare, and abdominal ground color black.

 Etymology

The specific epithet is derived from the Latin adjective "nigrum", meaning black, and "palpus", meaning palp, in reference to the black palpi of males of this species.

 Distribution

Costa Rica, ACG, Alajuela and Guanacaste provinces, 135–520m.

 Ecology

Calolydella nigripalpis has been reared six times from Semaeopus Janzen08 (Lepidoptera: Geometridae), in rain forest.

 86255FEB-3ABB-5F18-A2AE-8F2B0BFB2383Calolydella omissaurn:lsid:zoobank.org:act:7E31F35E-1BFB-4A68-B246-F0E22EC4AEDD Fleming & Wood sp. n. Materials

Type status: Holotype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0016605; recordedBy: D.H. Janzen, W. Hallwachs & Jose Perez; individualID: DHJPAR0016605; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-43825, BOLD:AAD9874, ASTAP809-07; Taxon: scientificName: Calolydella omissa; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: omissa; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Cabanya; verbatimElevation: 340; verbatimLatitude: 10.877; verbatimLongitude: -85.2308; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the erebid moth, Letis buteoDHJ03; verbatimEventDate: 02-Nov-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: 06-SRNP-43825; recordedBy: D.H. Janzen, W. Hallwachs & Jose Perez; individualID: 06-SRNP-43825; individualCount: 5; sex: 3M, 2F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-43825; Taxon: scientificName: Calolydella omissa; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: omissa; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Cabanya; verbatimElevation: 340; verbatimLatitude: 10.877; verbatimLongitude: -85.2308; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the erebid moth, Letis buteoDHJ03; verbatimEventDate: 02-Nov-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

 Description

Male (Fig. 26a, b, c). Length: 7–8mm. Head (Fig. 26b): frontal setae extending to base of postpedicel; fronto-orbital plate gold, with a single row of fine setulae outside of frontal setae; parafacial silver throughout. Thorax (Fig. 26a, c): gold on dorsal surface, silver laterally (>50% coverage); with four regular thoracic vittae; postpronotum with four setae; 3:3 acrostichal setae; 2:3 dorsocentral setae; 2:3 intra-alar setae; 2:3 supra-alar setae; three katepisternal setae; anatergite bare or rarely with up to two hair-like setae; scutellar discal setae situated as wide apart as subapical scutellar setae. Wing vein R4+5 with 6–7 setulae dorsally, reaching from base almost to crossvein R-M. Abdomen (Fig. 26a): with uninterrupted transverse marginal pollinose bands; abdominal pollinosity gold dorsally, silver ventrally, and with an orange spot lateroventrally at base of ST1+2; T3 with one pair of median marginal setae, lacking discal setae; T4 with one pair of discal setae. Terminalia sternite 5 (Fig. 27c): with two small lobes and a wide U-shaped median cleft, 0.45X the length of the sternite from lobe to apex; inner margin covered by dense pollinosity, appearing darker than surrounding cuticle; entire lobe of sternite with short setae, of varying lengths. Cerci (Fig. 27b), in dorsal view, separated by a narrow gap but rejoining at apex; each cercus long and very slightly tapered, almost of equal width along entire length; cercus slender with an upward bend when viewed laterally; setose along its entire length. Surstylus (Fig. 27a) slender and digitiform with a slight upward curve, appearing 0.2X shorter than cercus when viewed laterally; with short setae along entire length; tip of surstylus very slightly curved inwards when viewed dorsally.

Female (Fig. 26d, e, f). Length: 5–6mm. Fronto-orbital plate 1.4X wider than in male.

 Diagnosis

Calolydella omissa can be distinguished from all other species of Calolydella by the following combination of traits: fronto-orbital plate gold, parafacial silver, abdomen with uninterrupted transverse silver pollinose bands across all tergites, anatergite bare (rarely setose), and T3 without discal setae.

 Etymology

The specific epithet is derived from the Latin adjective "omissa", meaning dropped or absent, in reference to the absence of discal setae on T3, a character state unique to this species.

 Distribution

Costa Rica, ACG, Alajuela Province, Cabanya, 340m.

 Ecology

Calolydella omissa has been reared once from Letis Janzen03 (Lepidoptera: Erebidae), in rain forest.

 D258E8B6-4502-5D9C-8FA2-504500E1A5B1Calolydella ordinalisurn:lsid:zoobank.org:act:410DAC20-265A-4703-B31D-228436174914 Fleming & Wood sp. n. Materials

Type status: Holotype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017787; recordedBy: D.H. Janzen, W. Hallwachs & Mariano Pereira; individualID: DHJPAR0017787; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 98-SRNP-15880, BOLD:ACF1517, ASTAR498-07; Taxon: scientificName: Calolydella ordinalis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: ordinalis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Cacao; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Maritza; verbatimElevation: 760; verbatimLatitude: 10.9364; verbatimLongitude: -85.4776; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the erebid moth, Eloria torrida; verbatimEventDate: 20-Jan-1999; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017792; recordedBy: D.H. Janzen, W. Hallwachs & Mariano Pereira; individualID: DHJPAR0017792; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 98-SRNP-15883, BOLD:ACF1517, ASTAR503-07; Taxon: scientificName: Calolydella ordinalis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: ordinalis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Cacao; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Maritza; verbatimElevation: 760; verbatimLatitude: 10.9364; verbatimLongitude: -85.4776; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the erebid moth, Eloria torrida; verbatimEventDate: 20-Jan-1999; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017791; recordedBy: D.H. Janzen, W. Hallwachs & Mariano Pereira; individualID: DHJPAR0017791; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 98-SRNP-15887, BOLD:ACF1517, ASTAR502-07; Taxon: scientificName: Calolydella ordinalis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: ordinalis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Cacao; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Maritza; verbatimElevation: 760; verbatimLatitude: 10.9364; verbatimLongitude: -85.4776; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the erebid moth, Eloria torrida; verbatimEventDate: 20-Jan-1999; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017789; recordedBy: D.H. Janzen, W. Hallwachs & Mariano Pereira; individualID: DHJPAR0017789; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 98-SRNP-15883, BOLD:ACF1517, ASTAR500-07; Taxon: scientificName: Calolydella ordinalis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: ordinalis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Cacao; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Maritza; verbatimElevation: 760; verbatimLatitude: 10.9364; verbatimLongitude: -85.4776; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the erebid moth, Eloria torrida; verbatimEventDate: 20-Jan-1999; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017790; recordedBy: D.H. Janzen, W. Hallwachs & Mariano Pereira; individualID: DHJPAR0017790; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 98-SRNP-15887, BOLD:ACF1517, ASTAR501-07; Taxon: scientificName: Calolydella ordinalis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: ordinalis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Cacao; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Maritza; verbatimElevation: 760; verbatimLatitude: 10.9364; verbatimLongitude: -85.4776; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the erebid moth, Eloria torrida; verbatimEventDate: 20-Jan-1999; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017788; recordedBy: D.H. Janzen, W. Hallwachs & Mariano Pereira; individualID: DHJPAR0017788; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 98-SRNP-15891, BOLD:ACF1517, ASTAR499-07; Taxon: scientificName: Calolydella ordinalis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: ordinalis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Cacao; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Maritza; verbatimElevation: 760; verbatimLatitude: 10.9364; verbatimLongitude: -85.4776; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the erebid moth, Eloria torrida; verbatimEventDate: 20-Jan-1999; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

 Description

Male (Fig. 28a, b, c). Length: 6–8mm. Head (Fig. 28b): frontal setae extending to base of postpedicel; fronto-orbital plate gold, sparsely setulose, throughout; parafacial almost entirely gold (75-90% coverage). Thorax (Fig. 28a, c): gold on both dorsal and lateral surfaces; outermost two vittae twice as wide as innermost two; postpronotum with three setae; 3:3 acrostichal setae; 2:3 dorsocentral setae; 2:3 intra-alar setae; 2:3 supra-alar setae; three katepisternal setae; anatergite bare; scutellar discal setae situated as wide apart as or slightly wider apart than subapical scutellar setae. Wing vein R4+5 with at most 2–3 small setulae dorsally at base. Abdomen (Fig. 28a): ground color dark brown-orange to orange with uninterrupted transverse marginal pollinose bands; pollinosity gold dorsally, silver ventrally, and with an orange spot lateroventrally at base of ST1+2; T3 with one pair of median marginal setae and one to more than two pairs of discal setae; T4 with a row of marginal setae and a row of discal setae. Terminalia (Fig. 29): sternite 5 (Fig. 29a) with two small lobes and a wide U-shaped median cleft, 0.34X the length of the sternite from lobe to apex; inner margin covered by dense pollinosity, appearing darker than surrounding cuticle; entire lobe of sternite with short setae, of varying lengths. Cerci (Fig. 29b), in dorsal view, separated by a narrow gap, rejoining at apex; each cercus broad and digitiform, setose along its entire length; cercus with a very slight upward bend when viewed laterally. Surstylus (Fig. 29a) spatulate with a downward curve, appearing subequal to length of cercus when viewed laterally; with short setae along entire length; tip of surstylus not appearing curved inwards when viewed dorsally.

Female (Fig. 28d, e, f). Length: 5–6mm. Fronto-orbital plate 2.1X as wide as in male.

 Diagnosis

Calolydella ordinalis can be distinguished from all other species of Calolydella by the following combination of characters: parafacial mostly gold, fronto-orbital plate with setulae sparsely interspersed among frontal setae, anatergite bare, and a complete row of discal setae on T4.

 Etymology

The specific epithet is derived from the Latin adjective "ordinarius", meaning ordinary or complete, in reference to the complete, regular row of discal setae on T4, a character state unique to this species.

 Distribution

Costa Rica, ACG, Guanacaste Province, Sendero Maritza, 760m.

 Ecology

Calolydella ordinalis has been reared six times from Eloria torrida Schaus, 1910 (Lepidoptera: Erebidae) in cloud forest.

 C3D6F82F-169D-5E14-A104-2321085E92DECalolydella renemalaiseiurn:lsid:zoobank.org:act:FA671FB0-0BC4-48CA-86CE-B183BB3AD479 Fleming & Wood sp. n. Materials

Type status: Holotype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011709; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: DHJPAR0011709; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 04-SRNP-31285, BOLD:AAD9853, ASTAS435-06; Taxon: scientificName: Calolydella renemalaisei; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: renemalaisei; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Mismo; verbatimElevation: 680; verbatimLatitude: 10.9876; verbatimLongitude: -85.4197; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the erebid moth, Eucereon aeolum; verbatimEventDate: 10-Apr-2004; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011710; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: DHJPAR0011710; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 04-SRNP-31280, BOLD:AAD9853, ASTAS436-06; Taxon: scientificName: Calolydella renemalaisei; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: renemalaisei; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Mismo; verbatimElevation: 680; verbatimLatitude: 10.9876; verbatimLongitude: -85.4197; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the erebid moth, Eucereon aeolum; verbatimEventDate: 08-Apr-2004; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0057608; recordedBy: D.H. Janzen, W. Hallwachs & Gilberth Ampie; individualID: DHJPAR0057608; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 15-SRNP-891, BOLD:AAD9853, ASTAX020-15; Taxon: scientificName: Calolydella renemalaisei; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: renemalaisei; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero al Crater; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the erebid moth, Eucereon aeolum; verbatimEventDate: 20-Apr-2015; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011705; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: DHJPAR0011705; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 04-SRNP-31286, BOLD:AAD9853, ASTAS431-06; Taxon: scientificName: Calolydella renemalaisei; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: renemalaisei; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Mismo; verbatimElevation: 680; verbatimLatitude: 10.9876; verbatimLongitude: -85.4197; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the erebid moth, Eucereon aeolum; verbatimEventDate: 09-Apr-2004; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011712; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: DHJPAR0011712; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 04-SRNP-31283, BOLD:AAD9853, ASTAS438-06; Taxon: scientificName: Calolydella renemalaisei; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: renemalaisei; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Mismo; verbatimElevation: 680; verbatimLatitude: 10.9876; verbatimLongitude: -85.4197; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the erebid moth, Eucereon aeolum; verbatimEventDate: 10-Apr-2004; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017809; recordedBy: D.H. Janzen, W. Hallwachs & Elieth Cantillano; individualID: DHJPAR0017809; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 03-SRNP-2236, BOLD:AAD9853, ASTAR520-07; Taxon: scientificName: Calolydella renemalaisei; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: renemalaisei; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste; verbatimLocality: Puente Mena; verbatimElevation: 280; verbatimLatitude: 11.0456; verbatimLongitude: -85.4574; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the erebid moth, Leucotmemis nexa; verbatimEventDate: 20-Jun-2003; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

 Description

Male (Fig. 30a, b, c). Length: 5–8mm. Head (Fig. 30b): frontal setae extending to base of postpedicel; fronto-orbital plate gold, sparsely setulose throughout; parafacial up to 50% silver pollinose or mostly gold (75-90%). Thorax (Fig. 30a, c): gold on both dorsal and lateral surfaces; with four regular thoracic vittae; postpronotum with three setae; 3:3 acrostichal setae; 3:3 dorsocentral setae; 1:3 intra-alar setae; 2:3 supra-alar setae; three katepisternal setae; anatergite with three or more hair-like setae, often in a small tuft; scutellar discal setae situated as wide apart as subapical scutellar setae or absent. Wing vein R4+5 with at most 2–3 setulae dorsally at base, not extending to crossvein R-M. Abdomen (Fig. 30a): ground color dark brown-orange with uninterrupted transverse marginal pollinose bands; gold dorsally, silver ventrally, and with an orange spot lateroventrally at base of ST1+2 or base of ST1+2 black lateroventrally; T3 with one pair of median marginal setae and one pair of discal setae; T4 with 1–2 pairs of discal setae. Terminalia (Fig. 31): sternite 5 (Fig. 31c) with two small lobes and a wide U-shaped median cleft, 0.47X the length of the sternite from lobe to apex; inner margin covered by dense pollinosity, appearing darker than surrounding cuticle; entire lobe of sternite with short setae, all of equal length. Cerci (Fig. 31b), in dorsal view, separated by a narrow gap widening at apex; each cercus long, slender and straight, very slightly tapered from its already narrow base; setose along basal half. Surstylus (Fig. 31a) subequal to length of cercus, broad and digitiform when viewed laterally; with short setae along entire length; tip of surstylus not curved inwards when viewed dorsally.

Female: not known at this time.

 Diagnosis

Calolydella renemalaisei can be distinguished from all other other species of Calolydella by the following combination of characters: frontal setae extending to base of postpedicel, fronto-orbital plate sparsely setose througout, with four thoracic vittae, the outermost two twice as wide as the innermost two, anatergite with three or more hair-like setae arranged in a small tuft, and up to two pairs of discal setae on T4.

 Etymology

The specific epithet is in honor of René Edmond Malaise (1892-1978) of Stockholm, Sweden, in recognition of his invention of the Malaise trap, which has captured so many ACG Tachinidae and other insects, as explained by Fredrik Sjöberg in his wonderful book "The Fly Trap" (Sjöberg 2014, translation by Thomas Teal), a book that explains much about Malaise traps.

 Distribution

Costa Rica, ACG, Guanacaste Province, 280–680m.

 Ecology

Calolydella renemalaisei has been reared six times from two species of Arctiinae: Euceron aeolum Hampson, 1898, and Leucotmemis nexa Herrich-Schäffer, 1854 (Lepidoptera: Erebidae), in rain forest and dry-rain lowland intergrade ecosystems.

 F0546958-903B-50E6-A9C0-4CB1258818C6Calolydella susanaroibasaeurn:lsid:zoobank.org:act:A67E6DB2-D3C1-47BB-B098-B7373D8BEBC4 Fleming & Wood sp. n. Materials

Type status: Holotype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0057699; recordedBy: D.H. Janzen, W. Hallwachs & Carolina Cano; individualID: DHJPAR0057699; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 15-SRNP-1269, BOLD:ACW6315, ASTAX072-15; Taxon: scientificName: Calolydella susanaroibasae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: susanaroibasae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector San Cristobal; locality: Area de Conservacion Guanacaste; verbatimLocality: Rio Blanco Abajo; verbatimElevation: 500; verbatimLatitude: 10.9004; verbatimLongitude: -85.3725; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the erebid moth, Isanthrene echemon; verbatimEventDate: 20-May-2015; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: 15-SRNP-1269; recordedBy: D.H. Janzen, W. Hallwachs & Carolina Cano; individualID: 15-SRNP-1269; individualCount: 2; sex: 2F, siblings to holotype; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 15-SRNP-1269; Taxon: scientificName: Calolydella susanaroibasae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: susanaroibasae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector San Cristobal; locality: Area de Conservacion Guanacaste; verbatimLocality: Rio Blanco Abajo; verbatimElevation: 500; verbatimLatitude: 10.9004; verbatimLongitude: -85.3725; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the erebid moth, Isanthrene echemon; verbatimEventDate: 20-May-2015; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

 Description

Male (Fig. 32a, b, c). Length: 9mm. Head (Fig. 32b): frontal setae extending to base of postpedicel; fronto-orbital plate gold, with a single row of fine setulae outside of frontal setae; parafacial silver along lower half. Thorax (Fig. 32a, c): gold on dorsal surface, silver laterally (>50% coverage); outermost two vittae twice as wide as innermost two; postpronotum with three setae; 3:3 acrostichal setae; 3:3 dorsocentral setae; 2:3 intra-alar setae; 2:3 supra-alar setae 2:3; three katepisternal setae; anatergite with three or more hair-like setae, often in a small tuft, or rarely bare; scutellar discal setae situated as wide apart as subapical scutellar setae. Wing vein R4+5 with 5–7 setulae dorsally, reaching from base almost to crossvein R-M. Abdomen (Fig. 32a): ground color dark brown-orange, with transverse marginal pollinose bands; gold pollinosity dorsally, silver ventrally; base of ST1+2 black lateroventrally; T3 with one pair of median marginal setae and one pair of discal setae; T4 with 1–2 pairs of discal setae. Terminalia: not examined.

Female (Fig. 32d, e, f) . Length: 6mm. Fronto-orbital plate 1.7X as wide as in male.

 Diagnosis

Calolydella susanaroibasae can be distinguished from all other species of Calolydella by the following combination of characters: fronto-orbital plate with a single row of fine setulae outside of frontal setae, thoracic pollinosity gold dorsally and on 50% of lateral surface, with outer two thoracic vittae twice as wide as inner two, anatergite with three or more hair-like setae arranged in a small tuft, and base of ST1+2 black lateroventrally.

 Etymology

The specific epithet is in honor of Susana Roibas of Lexington, Kentucky, in recognition of her company "Sante-Traps" (http://www.santetraps.com), which has long supplied the world with Malaise traps and other kinds of insect traps, many of which have been used to collect ACG Tachinidae and other insects.

 Distribution

Costa Rica, ACG, Alajuela Province, Rio Blanco Abajo, 500m.

 Ecology

Calolydella susanaroibasae has been reared once from Isanthrene echemon Druce, 1884 (Lepidoptera: Erebidae: Arctiinae), in rain forest.

 D21AE302-34E4-59D3-AD0B-E6865DC004EFCalolydella tanyadapkeyaeurn:lsid:zoobank.org:act:C7F91FFF-F269-4D02-BAA6-903F6B5BBB28 Fleming & Wood sp. n. Materials

Type status: Holotype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011701; recordedBy: D.H. Janzen, W. Hallwachs & Jose Perez; individualID: DHJPAR0011701; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 05-SRNP-41436, BOLD:AAW8655, ASTAS427-06; Taxon: scientificName: Calolydella tanyadapkeyae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: tanyadapkeyae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Camino Rio Francia; verbatimElevation: 410; verbatimLatitude: 10.9043; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the erebid moth, Napata flaviceps; verbatimEventDate: 25-Jun-2005; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

 Description

Female (Fig. 33a, b, c). Length: 6mm. Head (Fig. 33b): frontal setae extending to base of postpedicel; fronto-orbital plate gold throughout, and sparsely setulose; parafacial silver throughout. Thorax (Fig. 33a, c): gold on dorsal surface, silver laterally (>50% coverage); with four regular thoracic vittae; postpronotum with two setae (inner basal seta absent); 3:3 acrostichal setae; 2:3 dorsocentral setae; 2:3 intra-alar setae; 2:3 supra-alar setae; three katepisternal setae; anatergite bare; scutellar discal setae situated as wide apart as subapical scutellar setae. Wing vein R4+5 with 6–7 setulae dorsally, extending to crossvein R-M. Abdomen (Fig. 33a): ground color black, with uninterrupted transverse marginal pollinose bands and with both dorsal and ventral surfaces concolorous; base of ST1+2 black lateroventrally; T3 with one pair of median marginal setae and one pair of discal setae; T4 with one pair of discal setae.

Male: not known at this time.

 Diagnosis

Calolydella tanyadapkeyae can be distinguished from all other other species of Calolydella by the following combination of traits: parafacial all silver, frontal setae extending to base of postpedicel, thoracic pollinosity gold dorsally and over 50% of lateral surfaces, postpronotum with only two setae (inner basal seta absent), anatergite bare, and scutellar discal setae situated as wide apart as subapical scutellar setae.

 Etymology

The specific epithet is in honor of Tanya Dapkey James of Levittown, Pennsylvania, in recognition of her efforts curating and preparing ACG parasitoid flies for DNA barcoding.

 Distribution

Costa Rica, ACG, Alajuela Province, Camino Rio Francia, 410m.

 Ecology

Calolydella tanyadapkeyae has been reared once from Uranophora flaviceps (Hampson, 1901) (Lepidoptera: Erebidae), in rain forest.

 44E1EE70-BD0D-5392-A907-93555802F2D4Calolydella tenebrosaurn:lsid:zoobank.org:act:1D33CB25-2145-45FC-B696-5E25C44CD4BD Fleming & Wood sp. n. Materials

Type status: Holotype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017810; recordedBy: D.H. Janzen, W. Hallwachs & Jose Perez; individualID: DHJPAR0017810; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 05-SRNP-40421, BOLD:AAG2405, ASTAR521-07; Taxon: scientificName: Calolydella tenebrosa; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: tenebrosa; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Vochysia; verbatimElevation: 320; verbatimLatitude: 10.8667; verbatimLongitude: -85.2453; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larvae of the geometrid moth, Cyclophora Janzen14; verbatimEventDate: 01-Mar-2005; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0053996; recordedBy: D.H. Janzen, W. Hallwachs & Ricardo Calero; individualID: DHJPAR0053996; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 13-SRNP-71761, BOLD:AAG2405, ASHYD3164-14; Taxon: scientificName: Calolydella tenebrosa; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: tenebrosa; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Medrano; verbatimElevation: 380; verbatimLatitude: 11.016; verbatimLongitude: -85.3805; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larvae of the geometrid moth, Semaeopus Janzen08; verbatimEventDate: 27-Nov-2013; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

 Description

Male (Fig. 34a, b, c). Length: 8mm. Head (Fig. 34b): frontal setae extending beyond base of postpedicel; fronto-orbital plate gold, sparsely setulose throughout; parafacial color all silver; or mostly gold (75-90%); palps black. Thorax (Fig. 34a, c): gold on both dorsal and lateral surfaces; with four regular thoracic vittae; postpronotum with three setae; 3:3 acrostichal setae; 3:3 dorsocentral setae; 2:3 intra-alar setae; 2:3 supra-alar setae; three katepisternal setae; anatergite bare; scutellar discal setae slightly closer together than subapical scutellar setae, or absent. Wing vein R4+5 with at most 2–3 setulae dorsally at base, not extending to crossvein R-M. Abdomen (Fig. 34a): ground color dark brown-orange, with uninterrupted transverse marginal pollinose bands and T5 black apically; pollinosity gold dorsally, silver ventrally; base of ST1+2 black lateroventrally; T3 with one pair of median marginal setae and two pairs of discal setae; T4 with two pairs of discal setae. Terminalia: not examined.

Female (Fig. 34d, e, f). Length: 6mm. As male, except for the following characters: fronto-orbital plate 1.41X as wide as in male; palps dark grayish-orange.

 Diagnosis

Calolydella tenebrosa can be distinguished from all other species of Calolydella by the following combination of characters: frontal setae extending beyond base of postpedicel, palps black in males and dark grayish-orange in females, abdominal ground color dark brown-orange, anatergite bare, and abdominal T5 black apically in both sexes.

 Etymology

The specific epithet Calolydella tenebrosa is derived from the Latin adjective "tenebrosum", meaning black or dark, in reference to the dark-colored palpi and dark-colored legs, character states found in both sexes of this species.

 Distribution

Costa Rica, ACG, Alajuela and Guanacaste provinces, 320–380m.

 Ecology

Calolydella tenebrosa has been reared once from Cyclophora Janzen14 and once from Semaeopus Janzen08 (Lepidoptera: Geometridae), in rain forest.

 0D75A381-069D-52BF-B9C0-091C1ECA39D9Calolydella timjamesiurn:lsid:zoobank.org:act:19044401-2286-439E-B788-B906DED6FBD5 Fleming & Wood sp. n. Materials

Type status: Holotype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017781; recordedBy: D.H. Janzen, W. Hallwachs & Freddy Quesada; individualID: DHJPAR0017781; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 99-SRNP-12851, BOLD:AAA1932, ASTAR492-07; Taxon: scientificName: Calolydella timjamesi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: timjamesi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector San Cristobal; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Corredor; verbatimElevation: 620; verbatimLatitude: 10.8787; verbatimLongitude: -85.3896; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the notodontid moth, Dunama jessiehillae; verbatimEventDate: 09-Dec-1999; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0016130; recordedBy: D.H. Janzen, W. Hallwachs & Petrona Rios; individualID: DHJPAR0016130; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-33809, BOLD:AAA1932, ASTAP159-06; Taxon: scientificName: Calolydella timjamesi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: timjamesi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Laguna; verbatimElevation: 680; verbatimLatitude: 10.9888; verbatimLongitude: -85.4234; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the notodontid moth, Tithraustes lambertae; verbatimEventDate: 11-Sep-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0016143; recordedBy: D.H. Janzen, W. Hallwachs & Calixto Moraga; individualID: DHJPAR0016143; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-33843, BOLD:AAA1932, ASTAP172-06; Taxon: scientificName: Calolydella timjamesi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: timjamesi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Laguna; verbatimElevation: 680; verbatimLatitude: 10.9888; verbatimLongitude: -85.4234; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the notodontid moth, Dunama janewaldronae; verbatimEventDate: 19-Sep-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0036611; recordedBy: D.H. Janzen, W. Hallwachs & Calixto Moraga; individualID: DHJPAR0036611; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 09-SRNP-32524, BOLD:AAA1932, ASHYE1522-09; Taxon: scientificName: Calolydella timjamesi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: timjamesi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Rotulo; verbatimElevation: 510; verbatimLatitude: 11.0135; verbatimLongitude: -85.4241; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the notodontid moth, Dioptis longipennis; verbatimEventDate: 16-Sep-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0016132; recordedBy: D.H. Janzen, W. Hallwachs & Calixto Moraga; individualID: DHJPAR0016132; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-33839, BOLD:AAA1932, ASTAP161-06; Taxon: scientificName: Calolydella timjamesi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: timjamesi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Laguna; verbatimElevation: 680; verbatimLatitude: 10.9888; verbatimLongitude: -85.4234; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the notodontid moth, Dunama janewaldronae; verbatimEventDate: 25-Sep-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0044914; recordedBy: D.H. Janzen, W. Hallwachs & Freddy Quesada; individualID: DHJPAR0044914; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 11-SRNP-71657, BOLD:AAA1932, ACGAZ138-11; Taxon: scientificName: Calolydella timjamesi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: timjamesi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Charia; verbatimElevation: 530; verbatimLatitude: 10.9934; verbatimLongitude: -85.4027; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the notodontid moth, Dottia viridifusca; verbatimEventDate: 26-Aug-2011; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017778; recordedBy: D.H. Janzen, W. Hallwachs & Freddy Quesada; individualID: DHJPAR0017778; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 99-SRNP-12846, BOLD:AAA1932, ASTAR489-07; Taxon: scientificName: Calolydella timjamesi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: timjamesi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector San Cristobal; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Corredor; verbatimElevation: 620; verbatimLatitude: 10.8787; verbatimLongitude: -85.3896; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the notodontid moth, Dunama jessiehillae; verbatimEventDate: 09-Nov-1999; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0053460; recordedBy: D.H. Janzen, W. Hallwachs & Elda Araya; individualID: DHJPAR0053460; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 13-SRNP-4277, BOLD:AAA1932, ASHYM2814-13; Taxon: scientificName: Calolydella timjamesi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: timjamesi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector San Cristobal; locality: Area de Conservacion Guanacaste; verbatimLocality: Rio Blanco Abajo; verbatimElevation: 500; verbatimLatitude: 10.9004; verbatimLongitude: -85.3725; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the notodontid moth, Tithraustes noctilucesICG02; verbatimEventDate: 06-Sep-2013; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0035882; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: DHJPAR0035882; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 09-SRNP-31957, BOLD:AAA1932, ASHYD1263-09; Taxon: scientificName: Calolydella timjamesi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: timjamesi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Memos; verbatimElevation: 740; verbatimLatitude: 10.9817; verbatimLongitude: -85.4278; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the notodontid moth, Dunama jessiehillae; verbatimEventDate: 11-Jul-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0016152; recordedBy: D.H. Janzen, W. Hallwachs & Petrona Rios; individualID: DHJPAR0016152; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-33804, BOLD:AAA1932, ASTAP181-06; Taxon: scientificName: Calolydella timjamesi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: timjamesi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Laguna; verbatimElevation: 680; verbatimLatitude: 10.9888; verbatimLongitude: -85.4234; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the notodontid moth, Tithraustes lambertae; verbatimEventDate: 21-Sep-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017783; recordedBy: D.H. Janzen, W. Hallwachs & Freddy Quesada; individualID: DHJPAR0017783; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 99-SRNP-5596, BOLD:AAA1932, ASTAR494-07; Taxon: scientificName: Calolydella timjamesi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: timjamesi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector San Cristobal; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Carmona; verbatimElevation: 670; verbatimLatitude: 10.8762; verbatimLongitude: -85.3863; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the notodontid moth, Dunama jessiehillae; verbatimEventDate: 07-Mar-1999; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017780; recordedBy: D.H. Janzen, W. Hallwachs & Freddy Quesada; individualID: DHJPAR0017780; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 99-SRNP-12853, BOLD:AAA1932, ASTAR491-07; Taxon: scientificName: Calolydella timjamesi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: timjamesi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector San Cristobal; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Corredor; verbatimElevation: 620; verbatimLatitude: 10.8787; verbatimLongitude: -85.3896; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the notodontid moth, Dunama jessiehillae; verbatimEventDate: 09-Dec-1999; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0016139; recordedBy: D.H. Janzen, W. Hallwachs & Calixto Moraga; individualID: DHJPAR0016139; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-33840, BOLD:AAA1932, ASTAP168-06; Taxon: scientificName: Calolydella timjamesi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: timjamesi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Laguna; verbatimElevation: 680; verbatimLatitude: 10.9888; verbatimLongitude: -85.4234; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the notodontid moth, Dunama janewaldronae; verbatimEventDate: 19-Sep-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011690; recordedBy: D.H. Janzen, W. Hallwachs & Yessenia Mendoza; individualID: DHJPAR0011690; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 04-SRNP-3964, BOLD:AAA1932, ASTAS416-06; Taxon: scientificName: Calolydella timjamesi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: timjamesi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector San Cristobal; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Corredor; verbatimElevation: 620; verbatimLatitude: 10.8787; verbatimLongitude: -85.3896; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the notodontid moth, Dottia viridifusca; verbatimEventDate: 05-Sep-2004; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0016137; recordedBy: D.H. Janzen, W. Hallwachs & Petrona Rios; individualID: DHJPAR0016137; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-33659, BOLD:AAA1932, ASTAP166-06; Taxon: scientificName: Calolydella timjamesi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: timjamesi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Evangelista; verbatimElevation: 660; verbatimLatitude: 10.9868; verbatimLongitude: -85.4208; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the notodontid moth, Dioptis longipennis; verbatimEventDate: 07-Sep-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0016129; recordedBy: D.H. Janzen, W. Hallwachs & Calixto Moraga; individualID: DHJPAR0016129; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-33813, BOLD:AAA1932, ASTAP158-06; Taxon: scientificName: Calolydella timjamesi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: timjamesi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Laguna; verbatimElevation: 680; verbatimLatitude: 10.9888; verbatimLongitude: -85.4234; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the notodontid moth, Tithraustes lambertae; verbatimEventDate: 25-Sep-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017779; recordedBy: D.H. Janzen, W. Hallwachs & Freddy Quesada; individualID: DHJPAR0017779; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 99-SRNP-12844, BOLD:AAA1932, ASTAR490-07; Taxon: scientificName: Calolydella timjamesi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: timjamesi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector San Cristobal; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Corredor; verbatimElevation: 620; verbatimLatitude: 10.8787; verbatimLongitude: -85.3896; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the notodontid moth, Dunama jessiehillae; verbatimEventDate: 09-Oct-1999; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017772; recordedBy: D.H. Janzen, W. Hallwachs & Carolina Cano; individualID: DHJPAR0017772; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 03-SRNP-8648, BOLD:AAA1932, ASTAR483-07; Taxon: scientificName: Calolydella timjamesi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: timjamesi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector San Cristobal; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Corredor; verbatimElevation: 620; verbatimLatitude: 10.8787; verbatimLongitude: -85.3896; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the notodontid moth, Dunama jessiehillae; verbatimEventDate: 19-Oct-2003; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017777; recordedBy: D.H. Janzen, W. Hallwachs & Freddy Quesada; individualID: DHJPAR0017777; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 99-SRNP-12847, BOLD:AAA1932, ASTAR488-07; Taxon: scientificName: Calolydella timjamesi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: timjamesi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector San Cristobal; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Corredor; verbatimElevation: 620; verbatimLatitude: 10.8787; verbatimLongitude: -85.3896; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the notodontid moth, Dunama jessiehillae; verbatimEventDate: 09-Dec-1999; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0007021; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: DHJPAR0007021; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 05-SRNP-70526, BOLD:AAA1932, ASTAV263-06; Taxon: scientificName: Calolydella timjamesi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: timjamesi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Colocho; verbatimElevation: 375; verbatimLatitude: 11.0237; verbatimLongitude: -85.4188; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the notodontid moth, Dunama jessiehillae; verbatimEventDate: 05-Feb-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0016128; recordedBy: D.H. Janzen, W. Hallwachs & Calixto Moraga; individualID: DHJPAR0016128; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-33929, BOLD:AAA1932, ASTAP157-06; Taxon: scientificName: Calolydella timjamesi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: timjamesi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Laguna; verbatimElevation: 680; verbatimLatitude: 10.9888; verbatimLongitude: -85.4234; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the notodontid moth, Dunama jessiehillae; verbatimEventDate: 25-Sep-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0016576; recordedBy: D.H. Janzen, W. Hallwachs & Calixto Moraga; individualID: DHJPAR0016576; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-34841, BOLD:AAA1932, ASTAP780-07; Taxon: scientificName: Calolydella timjamesi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: timjamesi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Laguna; verbatimElevation: 680; verbatimLatitude: 10.9888; verbatimLongitude: -85.4234; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the notodontid moth, Tithraustes lambertae; verbatimEventDate: 10-Nov-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0036621; recordedBy: D.H. Janzen, W. Hallwachs & Petrona Rios; individualID: DHJPAR0036621; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 09-SRNP-32404, BOLD:AAA1932, ASHYE1532-09; Taxon: scientificName: Calolydella timjamesi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: timjamesi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Cuestona; verbatimElevation: 640; verbatimLatitude: 10.9945; verbatimLongitude: -85.4146; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the notodontid moth, Dioptis longipennis; verbatimEventDate: 15-Sep-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0016131; recordedBy: D.H. Janzen, W. Hallwachs & Calixto Moraga; individualID: DHJPAR0016131; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-33838, BOLD:AAA1932, ASTAP160-06; Taxon: scientificName: Calolydella timjamesi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: timjamesi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Laguna; verbatimElevation: 680; verbatimLatitude: 10.9888; verbatimLongitude: -85.4234; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the notodontid moth, Dunama janewaldronae; verbatimEventDate: 21-Sep-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0027866; recordedBy: D.H. Janzen, W. Hallwachs & Jose Perez; individualID: DHJPAR0027866; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 08-SRNP-41163, BOLD:AAA1932, ASHYE103-08; Taxon: scientificName: Calolydella timjamesi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: timjamesi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Anonas; verbatimElevation: 405; verbatimLatitude: 10.9053; verbatimLongitude: -85.2788; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the notodontid moth, Dunama jessiehillae; verbatimEventDate: 30-Jun-2008; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0016192; recordedBy: D.H. Janzen, W. Hallwachs & Roster Moraga; individualID: DHJPAR0016192; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-22485, BOLD:AAA1932, ASTAP221-06; Taxon: scientificName: Calolydella timjamesi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: timjamesi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste; verbatimLocality: San Antonio; verbatimElevation: 335; verbatimLatitude: 11.0353; verbatimLongitude: -85.4453; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the notodontid moth, Dunama jessiehillae; verbatimEventDate: 13-Sep-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

 Description

Male (Fig. 35a, b, c). Length: 5–7mm. Head (Fig. 35b): frontal setae extending beyond base of postpedicel; fronto-orbital plate gold, sparsely setulose, these setulae being confined to the upper half; parafacial silver throughout. Thorax (Fig. 35a, c): gold on dorsal surface, silver laterally (>50% coverage); with four regular thoracic vittae presuturally, these fusing together postsuturally; postpronotum with two setae (inner basal seta absent); 3:3 acrostichal setae; 3:3 dorsocentral setae; 1:3 intra-alar setae; 1:3 supra-alar setae; three katepisternal setae; anatergite with three or more hair-like setae, often in a small tuft; scutellar discal setae slightly closer together than subapical scutellar setae. Wing vein R4+5 with 6–7 setulae dorsally from base to crossvein R-M. Abdomen (Fig. 35a): ground color black, with uninterrupted transverse marginal pollinose bands; abdomen gold dorsally, silver ventrally; base of ST1+2 black lateroventrally; T3 with one pair of median marginal setae and one pair of discal setae; T4 with one pair of discal setae. Terminalia (Fig. 36): sternite 5 (Fig. 36c) with two small lobes and a wide U-shaped median cleft; 0.47X the length of the sternite from lobe to apex; inner margin covered by dense pollinosity, appearing darker than surrounding cuticle; entire lobe of sternite with short stout setae, longer along margins of sternite. Cerci (Fig. 36b), in dorsal view, separated by a gap widening at apex; each cercus long, slender and pointed, very slightly tapered from its already narrow base; cercus slightly apically lobed, setose along its basal half. Surstylus (Fig. 36a) subequal to length of cercus, stout and spatulate, appearing rounded at apex when viewed laterally; apical half with short setulae; tip of surstylus angled (almost hooked) inwards when viewed dorsally.

Female (Fig. 35d, e, f). Length: 4–6mm. Fronto-orbital plate 1.73X wider than in male.

 Diagnosis

Calolydella timjamesi can be distinguished from all other species of Calolydella by the following combination of traits: fronto-orbital plate sparsely setulose to bare, frontal setae extending beyond base of postpedicel, abdominal ground color black, anatergite with three or more setae often in a small tuft, and scutellar discal setae slightly closer togetherthan subapical scutellar setae.

 Etymology

The specific epithet is in honor of Tim James of Levittown, Pennsylvania, in recognition of the moral and family support of Tanya Dapkey in her efforts curating and preparing ACG parasitoid flies for DNA barcoding.

 Distribution

Costa Rica, ACG, Alajuela and Guanacaste provinces, 335–740m.

 Ecology

Calolydella timjamesi has been reared 26 times from six species of Notodontidae: Dunama janewaldronae Chacon, 2013, Dunama jessiehillae Chacon, 2013, Tithraustes noctilucesICG02, Tithraustes lambertae Miller, 2008, Dioptis longipennis (Schaus, 1913), and Dottia viridifusca (Schaus, 1906), in rain forest and dry-rain lowland intergrade.

 0962604B-89E7-5B54-B1DE-0C0B9ED946D0Calolydella virginiajamesaeurn:lsid:zoobank.org:act:58B44ECE-D7A6-49F6-B050-D118409D499A Fleming & Wood sp. n. Materials

Type status: Holotype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017770; recordedBy: D.H. Janzen, W. Hallwachs & Gloria Sihezar; individualID: DHJPAR0017770; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 99-SRNP-13019, BOLD:ABY7242, ASTAR481-07; Taxon: scientificName: Calolydella virginiajamesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: virginiajamesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector San Cristobal; locality: Area de Conservacion Guanacaste; verbatimLocality: Quebrada Cementerio; verbatimElevation: 700; verbatimLatitude: 10.8712; verbatimLongitude: -85.3875; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the sphingid moth, Manduca sextaDHJ03; verbatimEventDate: 13-Oct-1999; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0054010; recordedBy: D.H. Janzen, W. Hallwachs & Keiner Aragon; individualID: DHJPAR0054010; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 13-SRNP-45063, BOLD:ABY7242, ASHYD3178-14; Taxon: scientificName: Calolydella virginiajamesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: virginiajamesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Palomo; verbatimElevation: 96; verbatimLatitude: 10.9619; verbatimLongitude: -85.2804; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the gelechiid, gelJanzen01 Janzen11; verbatimEventDate: 03-Dec-2013; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0034486; recordedBy: D.H. Janzen, W. Hallwachs & Duvalier Briceno; individualID: DHJPAR0034486; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 09-SRNP-65345, BOLD:ABY7242, ASHYC1138-09; Taxon: scientificName: Calolydella virginiajamesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: virginiajamesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Brasilia; locality: Area de Conservacion Guanacaste; verbatimLocality: Moga; verbatimElevation: 320; verbatimLatitude: 11.0123; verbatimLongitude: -85.3493; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the hesperiid moth, Saliana antoninus; verbatimEventDate: 12-Mar-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0039336; recordedBy: D.H. Janzen, W. Hallwachs & Calixto Moraga; individualID: DHJPAR0039336; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 10-SRNP-30911, BOLD:ABY7242, ASTAV899-10; Taxon: scientificName: Calolydella virginiajamesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: virginiajamesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Estacion Pitilla; verbatimElevation: 675; verbatimLatitude: 10.9893; verbatimLongitude: -85.4258; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the crambid moth, Herpetogramma phaeopteralis; verbatimEventDate: 26-Apr-2010; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0053994; recordedBy: D.H. Janzen, W. Hallwachs & Ricardo Calero; individualID: DHJPAR0053994; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 13-SRNP-71701, BOLD:ABY7242, ASHYD3162-14; Taxon: scientificName: Calolydella virginiajamesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: virginiajamesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Medrano; verbatimElevation: 380; verbatimLatitude: 11.016; verbatimLongitude: -85.3805; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid butterfly Thisbe irenea; verbatimEventDate: 02-Nov-2013; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0007071; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: DHJPAR0007071; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-30377, BOLD:ABY7242, ASTAB408-10; Taxon: scientificName: Calolydella virginiajamesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: virginiajamesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Pasmompa; verbatimElevation: 440; verbatimLatitude: 11.0193; verbatimLongitude: -85.41; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the hesperiid butterfly, Perichares poaceaphaga; verbatimEventDate: 10-Feb-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017786; recordedBy: D.H. Janzen, W. Hallwachs & Jose Perez; individualID: DHJPAR0017786; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 00-SRNP-20847, BOLD:ABY7242, ASTAR497-07; Taxon: scientificName: Calolydella virginiajamesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: virginiajamesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Rincon; verbatimElevation: 430; verbatimLatitude: 10.8962; verbatimLongitude: -85.2777; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Danaus plexippus; verbatimEventDate: 28-Nov-2000; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017769; recordedBy: D.H. Janzen, W. Hallwachs & Dunia Garcia; individualID: DHJPAR0017769; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 02-SRNP-15501, BOLD:ABY7242, ASTAR480-07; Taxon: scientificName: Calolydella virginiajamesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: virginiajamesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector El Hacha; locality: Area de Conservacion Guanacaste; verbatimLocality: Finca Araya; verbatimElevation: 295; verbatimLatitude: 11.0154; verbatimLongitude: -85.5113; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Mechanitis isthmia; verbatimEventDate: 26-Jun-2002; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0052421; recordedBy: D.H. Janzen, W. Hallwachs & Cirilo Umana; individualID: DHJPAR0052421; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 13-SRNP-77065, BOLD:ABY7242, ASHYM1775-13; Taxon: scientificName: Calolydella virginiajamesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: virginiajamesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Quebrada Bambu; verbatimElevation: 109; verbatimLatitude: 10.9301; verbatimLongitude: -85.2521; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the erebid moth, Anycles adusta; verbatimEventDate: 09-Aug-2013; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011696; recordedBy: D.H. Janzen, W. Hallwachs & Eilyn Camacho; individualID: DHJPAR0011696; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 05-SRNP-60644, BOLD:ABY7242, ASTAS422-06; Taxon: scientificName: Calolydella virginiajamesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: virginiajamesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Santa Rosa; locality: Area de Conservacion Guanacaste; verbatimLocality: Quebrada Puercos; verbatimElevation: 155; verbatimLatitude: 10.8591; verbatimLongitude: -85.5709; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the noctuid moth, Cropia cedica; verbatimEventDate: 21-Sep-2005; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0045648; recordedBy: D.H. Janzen, W. Hallwachs & Minor Carmona; individualID: DHJPAR0045648; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 11-SRNP-66015, BOLD:ABY7242, ACGAZ837-11; Taxon: scientificName: Calolydella virginiajamesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: virginiajamesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Brasilia; locality: Area de Conservacion Guanacaste; verbatimLocality: Gallinazo; verbatimElevation: 360; verbatimLatitude: 11.0183; verbatimLongitude: -85.372; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the noctuid moth, Perigea agnonia; verbatimEventDate: 01-Oct-2011; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017767; recordedBy: D.H. Janzen, W. Hallwachs & Ruth Franco; individualID: DHJPAR0017767; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 02-SRNP-32494, BOLD:ABY7242, ASTAR478-07; Taxon: scientificName: Calolydella virginiajamesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: virginiajamesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Santa Rosa; locality: Area de Conservacion Guanacaste; verbatimLocality: Quebrada Puercos; verbatimElevation: 155; verbatimLatitude: 10.8591; verbatimLongitude: -85.5709; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the pierid butterfly Colias cesonia; verbatimEventDate: 24-Nov-2002; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0057673; recordedBy: D.H. Janzen, W. Hallwachs & Elissa Arroyo; individualID: DHJPAR0057673; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 15-SRNP-30814, BOLD:ABY7242, ASTAX046-15; Taxon: scientificName: Calolydella virginiajamesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: virginiajamesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Ingas; verbatimElevation: 580; verbatimLatitude: 11.0031; verbatimLongitude: -85.4204; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the crambid moth, Rhectocraspeda periusalis; verbatimEventDate: 25-May-2015; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0007075; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: DHJPAR0007075; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-31177, , ASTAV317-06; Taxon: scientificName: Calolydella virginiajamesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: virginiajamesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Pasmompa; verbatimElevation: 440; verbatimLatitude: 11.0193; verbatimLongitude: -85.41; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the lycaenid butterfly, Arawacus togarna; verbatimEventDate: 04-Apr-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0046613; recordedBy: D.H. Janzen, W. Hallwachs & Ricardo Calero; individualID: DHJPAR0046613; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 11-SRNP-72723, BOLD:ABY7242, ACGBA786-12; Taxon: scientificName: Calolydella virginiajamesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: virginiajamesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Medrano; verbatimElevation: 380; verbatimLatitude: 11.016; verbatimLongitude: -85.3805; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the erebid moth, noctJanzen01 Janzen102; verbatimEventDate: 04-Jan-2012; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0023071; recordedBy: D.H. Janzen, W. Hallwachs & Roster Moraga; individualID: DHJPAR0023071; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 07-SRNP-24114, BOLD:ABY7242, ASTAW235-08; Taxon: scientificName: Calolydella virginiajamesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: virginiajamesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste; verbatimLocality: Uncaria; verbatimElevation: 370; verbatimLatitude: 11.0175; verbatimLongitude: -85.4741; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the limacodid moth, Epiperola vafera; verbatimEventDate: 08-Nov-2007; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0010317; recordedBy: D.H. Janzen, W. Hallwachs & Lucia Vargas; individualID: DHJPAR0010317; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-17555, BOLD:ABY7242, ASTAS148-06; Taxon: scientificName: Calolydella virginiajamesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: virginiajamesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Horizontes; locality: Area de Conservacion Guanacaste; verbatimLocality: Vado Esperanza; verbatimElevation: 85; verbatimLatitude: 10.7894; verbatimLongitude: -85.551; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the crambid moth, Spoladea recurvalis; verbatimEventDate: 18-Jul-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0050664; recordedBy: D.H. Janzen, W. Hallwachs & Ricardo Calero; individualID: DHJPAR0050664; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 12-SRNP-72398, BOLD:ABY7242, ACGBA3256-13; Taxon: scientificName: Calolydella virginiajamesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: virginiajamesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Medrano; verbatimElevation: 380; verbatimLatitude: 11.016; verbatimLongitude: -85.3805; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the crambid moth, Hileithia Solis01DHJ02; verbatimEventDate: 08-Dec-2012; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0021000; recordedBy: D.H. Janzen, W. Hallwachs & Jose Perez; individualID: DHJPAR0021000; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 07-SRNP-42208, BOLD:ABY7242, ASTA1343-07; Taxon: scientificName: Calolydella virginiajamesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: virginiajamesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Camino Porvenir; verbatimElevation: 383; verbatimLatitude: 10.9038; verbatimLongitude: -85.2596; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid butterfly, Euselasia cheles; verbatimEventDate: 18-Aug-2007; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0035672; recordedBy: D.H. Janzen, W. Hallwachs & Mariano Pereira; individualID: DHJPAR0035672; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 09-SRNP-56981, BOLD:ABY7242, ASHYD1053-09; Taxon: scientificName: Calolydella virginiajamesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: virginiajamesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Mundo Nuevo; locality: Area de Conservacion Guanacaste; verbatimLocality: Camino Pozo Tres; verbatimElevation: 733; verbatimLatitude: 10.7708; verbatimLongitude: -85.3742; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Chlosyne rosita; verbatimEventDate: 06-Aug-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0046436; recordedBy: D.H. Janzen, W. Hallwachs & Jose Perez; individualID: DHJPAR0046436; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 11-SRNP-26137, BOLD:ABY7242, ACGBA609-12; Taxon: scientificName: Calolydella virginiajamesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: virginiajamesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Venado; verbatimElevation: 420; verbatimLatitude: 10.8968; verbatimLongitude: -85.27; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the crambid moth, Undulambia BioLep248; verbatimEventDate: 12-Jan-2012; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017052; recordedBy: D.H. Janzen, W. Hallwachs & Calixto Moraga; individualID: DHJPAR0017052; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 07-SRNP-30522, BOLD:ABY7242, ASTAP490-07; Taxon: scientificName: Calolydella virginiajamesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: virginiajamesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Nacho; verbatimElevation: 710; verbatimLatitude: 10.9845; verbatimLongitude: -85.4248; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the erebid moth, Coreura phoenicides; verbatimEventDate: 15-Feb-2007; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0046433; recordedBy: D.H. Janzen, W. Hallwachs & Jose Perez; individualID: DHJPAR0046433; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 11-SRNP-26139, BOLD:ABY7242, ACGBA606-12; Taxon: scientificName: Calolydella virginiajamesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: virginiajamesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Venado; verbatimElevation: 420; verbatimLatitude: 10.8968; verbatimLongitude: -85.27; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the noctuid moth, Callopistria floridensis; verbatimEventDate: 14-Jan-2012; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0018404; recordedBy: D.H. Janzen, W. Hallwachs & Petrona Rios; individualID: DHJPAR0018404; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 05-SRNP-30122, BOLD:ABY7242, ASTAI1051-07; Taxon: scientificName: Calolydella virginiajamesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: virginiajamesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Pasmompa; verbatimElevation: 440; verbatimLatitude: 11.0193; verbatimLongitude: -85.41; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the sphingid moth, Manduca muscosa; verbatimEventDate: 10-Feb-2005; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011713; recordedBy: D.H. Janzen, W. Hallwachs & Guillermo Pereira; individualID: DHJPAR0011713; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 04-SRNP-13310, BOLD:ABY7242, ASTAS439-06; Taxon: scientificName: Calolydella virginiajamesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: virginiajamesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Santa Rosa; locality: Area de Conservacion Guanacaste; verbatimLocality: Quebrada San Pancho; verbatimElevation: 90; verbatimLatitude: 10.7477; verbatimLongitude: -85.5858; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the crambid moth, Omiodes cuniculalis; verbatimEventDate: 13-Aug-2004; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0016740; recordedBy: D.H. Janzen, W. Hallwachs & Jose Alberto Sanchez; individualID: DHJPAR0016740; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-59934, BOLD:ABY7242, ASTAP850-07; Taxon: scientificName: Calolydella virginiajamesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: virginiajamesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Mundo Nuevo; locality: Area de Conservacion Guanacaste; verbatimLocality: Porton Rivas; verbatimElevation: 570; verbatimLatitude: 10.7586; verbatimLongitude: -85.3727; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the crambid moth, Omiodes cuniculalis; verbatimEventDate: 06-Dec-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0046439; recordedBy: D.H. Janzen, W. Hallwachs & Jose Perez; individualID: DHJPAR0046439; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 11-SRNP-26138, BOLD:ABY7242, ACGBA612-12; Taxon: scientificName: Calolydella virginiajamesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: virginiajamesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Venado; verbatimElevation: 420; verbatimLatitude: 10.8968; verbatimLongitude: -85.27; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the noctuid moth, Callopistria floridensis; verbatimEventDate: 10-Jan-2012; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0040787; recordedBy: D.H. Janzen, W. Hallwachs & Dunia Garcia; individualID: DHJPAR0040787; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 10-SRNP-36111, BOLD:ABY7242, ASHYE2923-11; Taxon: scientificName: Calolydella virginiajamesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: virginiajamesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Cacao; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero a Maritza, 1 km NW Estacion Cacao; verbatimElevation: 1150; verbatimLatitude: 10.9592; verbatimLongitude: -85.4682; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the hesperiid butterfly, Mylon lassia; verbatimEventDate: 22-Sep-2010; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0003459; recordedBy: D.H. Janzen, W. Hallwachs & Petrona Rios; individualID: DHJPAR0003459; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 05-SRNP-30122, , ASTA366-05; Taxon: scientificName: Calolydella virginiajamesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: virginiajamesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Pasmompa; verbatimElevation: 440; verbatimLatitude: 11.0193; verbatimLongitude: -85.41; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the sphingid moth, Manduca muscosa; verbatimEventDate: 10-Feb-2005; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017785; recordedBy: D.H. Janzen, W. Hallwachs & Jose Perez; individualID: DHJPAR0017785; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 00-SRNP-20847, BOLD:ABY7242, ASTAR496-07; Taxon: scientificName: Calolydella virginiajamesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: virginiajamesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Rincon; verbatimElevation: 430; verbatimLatitude: 10.8962; verbatimLongitude: -85.2777; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the nymphalid butterfly, Danaus plexippus; verbatimEventDate: 28-Nov-2000; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0029640; recordedBy: D.H. Janzen, W. Hallwachs & Gloria Sihezar; individualID: DHJPAR0029640; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 08-SRNP-5217, BOLD:ABY7242, ASHYM1061-09; Taxon: scientificName: Calolydella virginiajamesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: virginiajamesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector San Cristobal; locality: Area de Conservacion Guanacaste; verbatimLocality: Rio Areno; verbatimElevation: 460; verbatimLatitude: 10.9141; verbatimLongitude: -85.3817; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the erebid moth, Hypena coatilis; verbatimEventDate: 07-Oct-2008; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0030167; recordedBy: D.H. Janzen, W. Hallwachs & Petrona Rios; individualID: DHJPAR0030167; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 08-SRNP-32592, BOLD:ABY7242, ASHYB911-09; Taxon: scientificName: Calolydella virginiajamesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: virginiajamesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Amonias; verbatimElevation: 390; verbatimLatitude: 11.0425; verbatimLongitude: -85.4034; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the geometrid moth, Eois Janzen04; verbatimEventDate: 05-Nov-2008; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0016580; recordedBy: D.H. Janzen, W. Hallwachs & Manuel Rios; individualID: DHJPAR0016580; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-65089, BOLD:ABY7242, ASTAP784-07; Taxon: scientificName: Calolydella virginiajamesae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: virginiajamesae; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Coneja; verbatimElevation: 415; verbatimLatitude: 11.0153; verbatimLongitude: -85.3977; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the riodinid butterfly, Ancyluris inca; verbatimEventDate: 22-Nov-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

 Description

Male (Fig. 37a, b, c). Length: 5–8mm. Head (Fig. 37b): frontal setae extending beyond base of postpedicel; fronto-orbital plate gold, with a single row of fine setulae outside of frontal setae; parafacial silver throughout. Thorax (Fig. 37a, c): gold on both dorsal and lateral surfaces; outermost two vittae twice as wide as innermost two; postpronotum with three setae; 3:3 acrostichal setae; 3:3 dorsocentral setae; 2:3 intra-alar setae; 2:3 supra-alar setae; three katepisternal setae; anatergite with three or more hair-like setae, often in a small tuft; scutellar discal setae situated as wide apart as subapical scutellar setae. Wing vein R4+5 with 7–9 setulae from base to crossvein R-M. Abdomen (Fig. 37a): with uninterrupted transverse marginal pollinose bands, and with an orange spot lateroventrally at base of ST1+2; T3 with one pair of median marginal setae and one pair of discal setae; T4 with one pair of discal setae. Terminalia (Fig. 38): sternite 5 (Fig. 38c) with two small lobes and a wide U-shaped median cleft, 0.48X the length of the sternite from lobe to apex; inner margin covered by dense pollinosity, appearing darker than surrounding cuticle; entire lobe of sternite with short setae, of varying lengths. Cerci (Fig. 38b), in dorsal view, separated by a narrow gap widening at apex, and with separated tips; each cercus long, slender and straight, very slightly tapered from its already narrow base; setose along its entire length. Surstylus (Fig. 38a) slender and digitiform, appearing 0.2X shorter than cercus when viewed laterally; with short setae along entire length; tip of surstylus curved inwards when viewed dorsally.

Female (Fig. 37d, e, f). Length: 4–7mm. Fronto-orbital plate 2.1X wider than in male.

 Diagnosis

Calolydella virginiajamesae can be distinguished from all other species of Calolydella by the following combination of traits: fronto-orbital plate with a single row of small black setulae outside of frontal setae, parafacial silver-pollinose, anatergite with three or more setae arranged in a small tuft, and T3–T4 each with a single pair of discal setae.

 Etymology

The specific epithet is in honor of Virginia James of Levittown, Pennsylvania, in recognition of the moral and family support of her mother, Tanya Dapkey, in her efforts curating and preparing ACG parasitoid flies for DNA barcoding.

 Distribution

Costa Rica, ACG, Alajuela and Guanacaste provinces, 85–1150m.

 Ecology

Calolydella virginiajamesae has been reared 34 times from various species of Lepidoptera from several families (Nymphalidae; Hesperiidae; Lycaenidae; Geometridae; Crambidae; Erebidae; Riodinidae; Sphingidae; and Pieridae), in rain forest, cloud forest, dry forest, and dry-rain lowland intergrade ecosystems.

 Identification Keys Key to the species of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Calolydella">Calolydella</tp:taxon-name-part></tp:taxon-name></italic> of the Mesoamerican region
1 Fronto-orbital plate with gold pollinosity; thorax with some gold on both lateral and dorsal surfaces 2
– Fronto-orbital plate entirely silver; thorax silver on both lateral and dorsal surfaces 25
2 Fronto-orbital plate entirely gold 3
– Fronto-orbital plate with both silver and gold pollinosity (Fig. 15b, d) Calolydella erasmocoronadoi sp. n.
3 Abdomen with transverse marginal pollinose bands interrupted by a median dark stripe 4
– Abdomen with uninterrupted transverse marginal pollinose bands 8
4 Parafacial pollinosity either entirely silver or up to 50% silver pollinose 5
– Parafacial pollinosity entirely gold Calolydella gentica (Walker, 1860)
5 Fronto-orbital plate with setulae outside of frontal seta; thorax with four distinct dorsal vittae; anatergite with up to two fine hair-like seta; T3 with one pair of median marginal setae 6
– Fronto-orbital plate bare; thorax with two, prominent, dark dorsal vittae; anatergite with three or more hair-like setae; T3 with a complete row of marginal setae and one pair of discal setae 7
6 Fronto-orbital plate sparsely setulose; frontal setae not extending beyond base of postpedicel; anatergite bare; wing vein R4+5 with 4–5 small setulae dorsally, near base; T3 with one pair of discal setae (Fig. 9a) Calolydella crocata sp. n.
– Fronto-orbital plate with a row of fine setulae outside of frontal setae; frontal setae extending beyond base of postpedicel; anatergite with up to two hair-like setae; wing vein R4+5 with at most three small setulae dorsally, near base; T3 with two pairs of discal setae Calolydella trifasciata (Walker, 1837)
7 Thorax with two presutural acrostichal setae; abdominal ST1+2 mostly black dorsally, with small spots of gold pollinosity laterally (Fig. 23a, d) Calolydella interrupta sp. n.
– Thorax with three presutural acrostichal setae; abdominal ST1+2 with a thick median black stripe but still mostly gold dorsally (Fig. 6a, c) Calolydella bicolor sp. n.
8 T3 with at least one pair of median marginal setae 9
– T3 without median marginal setae (Fig. 11a, d) Calolydella destituta sp. n.
9 T3 with at least one pair of discal setae 10
– T3 without discal setae (Fig. 26a, d) Calolydella omissa sp. n.
10 Palpus dark gray to black 11
– Palpus yellow-orange 12
11 Frontal setae extending to base of postpedicel but not beyond; thoracic vittae only faintly visible; abdominal ground color black (Fig. 24a, b) Calolydella nigripalpis sp. n.
– Frontal setae extending beyond base of postpedicel; thoracic vittae prominent and easily visible; abdominal ground color dark brown-orange (Fig. 34a, b, d, e) Calolydella tenebrosa sp. n.
12 One or more pairs of discal setae on abdominal T4 13
– A complete row of discal setae on abdominal T4 (Fig. 28a, b, d, e) Calolydella ordinalis sp. n.
13 Frontal setae extending beyond base of postpedicel 14
– Frontal setae extending to base of postpedicel 17
14 Parafacial silver; palpus normal; T3 and T4 each with one pair of discal setae 15
– Parafacial gold; palpus normal in male but significantly enlarged in female (Fig. 21b, e); T3 and T4 each with two pairs of discal setae (Fig. 21c, f) Calolydella inflatipalpis sp. n.
15 Four distinct thoracic vittae postsuturally 16
– Thoracic vittae fused postsuturally (Fig. 35a, d) Calolydella timjamesi sp. n.
16 Anatergite with up to two hair-like setae or, rarely, bare (Fig. 17a, d) Calolydella felipechavarriai sp. n.
– Anatergite with a small tuft of three or more hair-like setae (Fig. 37a, d) Calolydella virginiajamesae sp. n.
17 Abdominal pollinosity concolorous on both ventral and dorsal surfaces 18
– Abdominal pollinosity gold dorsally, silver ventrally 19
18 Parafacial all gold pollinose; palpus black basally; thorax with four regular thoracic vittae; scutellum without discal setae; abdomen silver pollinose on both dorsal and ventral surfaces (Fig. 5a, b) Calolydella aureofacies sp. n.
– Parafacial with some silver pollinosity; palpus orange; thorax with two bold and prominent thoracic vittae; scutellum with discal setae; abdomen gold pollinose on both dorsal and ventral surfaces (Fig. 7a, d) Calolydella bifissus sp. n.
19 Base of ST1+2 with an orange spot lateroventrally 20
– Base of ST1+2 black lateroventrally (Fig. 33a, b) Calolydella tanyadapkeyae sp. n.
20 Parafacial silver-pollinose throughout 21
– Parafacial with both silver and gold pollinosity 22
21 Anatergite with three or more hair-like setae arranged in a tuft; thoracic chaetotaxy as follows: 3:3 acrostichal setae; 3:3 dorsocentral setae; 2:3 intra-alar setae; 2:3 supra-alar setae; abdominal T4 with one pair of discal setae (Fig. 19a, b, d, e) Calolydella fredriksjobergi sp. n.
– Anatergite bare; thoracic chaetotaxy as follows: 3:3 acrostichal setae (2:3 in females); 3:4 dorsocentral setae; 1:3 intra-alar setae; 1:3 supra-alar setae; abdominal T4 with two pairs of discal setae(Fig. 1a, b, d, e) Calolydella adelinamoralesae sp. n.
22 Outermost thoracic vittae 2X as wide as innermost, these becoming fused postsuturally 23
– Four thoracic vittae of sub-equal width, not fused postsuturally 24
23 Fronto-orbital plate with a single row of setulae outside of frontal setae; postpronotum with three setae; wing vein R4+5 with 5–7 setulae dorsally, reaching from base almost to crossvein R-M; abdominal T4 with two pairs of discal setae (Fig. 32a, b, d, e) Calolydella susanaroibasae sp. n.
– Fronto-orbital plate with setulae interspersed among frontal setae; postpronotum with two setae, inner basal seta absent; wing vein R4+5 with at most three small setulae dorsally at base; T4 with one pair of discal setae (Fig. 3a, b) Calolydella alexanderjamesi sp. n.
24 Fronto-orbital plate with a single row of setulae outside of frontal setae; thoracic chaetotaxy as follows: 3:3 acrostichal setae; 2:3 dorsocentral setae; 3:3 intra-alar setae; 2:3 supra-alar setae; wing vein R4+5 with more than 4–5 setulae dorsally, which may or may not extend to crossvein R-M; T3 and T4 both with up to two pairs of discal setae (Fig. 13a, b) Calolydella discalis sp. n.
– Fronto-orbital plate with setulae interspersed among frontal setae; thoracic chaetotaxy as follows: 3:3 acrostichal setae; 3:3 dorsocentral setae; 1:3 intra-alar setae; 2:3 supra-alar setae; wing vein R4+5 with at most 2–3 setulae dorsally at base, never extending to crossvein R-M; T3 with 1 pair of discal setae; T4 with up to two pairs of discal setae (Fig. 30a, b) Calolydella renemalaisei sp. n.
25 Thorax with two longitudinal vittae 26
– Thorax with four distinct longitudinal vittae 28
26 T3 with only one pair of discal setae; abdominal pollinosity appearing as thick uninterrupted bands across tergites 27
– T3 with two pairs of discal setae; abdominal pollinosity across tergites interrupted by a dark median stripe Calolydella summatis Reinhard, 1974
27 Frontal setae extending to base of postpedicel; three supravibrissal setae; 3:3 acrostichal setae; anatergite with three or more hair-like setae in a small tuft; anterior 2/3 of abdominal tergites silver pollinose; wing vein R4+5 with 6–7 setulae dorsally, extending almost to crossvein R-M (Fig. 4a, b) Calolydella argentea sp. n.
– Frontal setae extending beyond base of postpedicel; six or more supravibrissal setae; 2:3 acrostichal setae; anatergite bare; anterior 1/3 of abdominal tergites silver pollinose; wing vein R4+5 with two setulae dorsally, not extending to crossvein R-M Calolydella cylindriventris (Wulp, 1890)
28 Fronto-orbital plate bare; frontal setae reaching base of postpedicel; thoracic pollinosity silver throughout; T3 with one pair of discal setae and one pair of median marginal setae Calolydella triangulifera (Bigot, 1889)
– Fronto-orbital plate with sparse setulae interspersed among frontal setae; frontal setae reaching beyond base of postpedicel; thoracic pollinosity gold throughout; T3 with two pairs of discal setae and a row of marginal setae Calolydella blandita (Wulp, 1890)

This key is confined to species of Calolydella occurring in the Mesoamerican biogeographical region. It is applicable to both males and females.

 Analysis DNA Barcoding

A phylogenetic tree based on DNA barcodes was used to visually demonstrate the variation within and between species, and is presented in Fig. 39. Interested readers can consult the Barcode of Life Data System (BOLD) (Ratnasingham and Hebert 2007) for all information associated with each sequence (including GenBank accession numbers), derived from each individual specimen using the persistent DOI: dx.doi.org/10.5883/DS-ASCALOL.

 Acknowledgements

We gratefully acknowledge the unflagging support of the team of ACG parataxonomists (Janzen et al 2009, Janzen & Hallwachs 2011) who found and reared the specimens used in this study, and the team of biodiversity managers who protect and manage the ACG forests that are home to these tachinids and their caterpillar hosts. The study has been supported by U.S. National Science Foundation grants BSR 9024770 and DEB 9306296, 9400829, 9705072, 0072730, 0515699, and grants from the Wege Foundation, International Conservation Fund of Canada, Jessie B. Cox Charitable Trust, Blue Moon Fund, Guanacaste Dry Forest Conservation Fund, Area de Conservación Guanacaste, Permian Global and University of Pennsylvania (DHJ & WH). This study has been supported by the Government of Canada through its ongoing support of the Canadian National Collection, Genome Canada, the Biodiversity Institute of Ontario, and the Ontario Genomics Institute (2008–0GI–ICI–03) (MAS), and by a Discovery Grant from Natural Sciences and Engineering Research Council of Canada (MAS).

 References 1907 The synonymy and generic position of certain species of Muscidae (sens. lat.) in the collection of the British Museum, described by the late Francis Walker The annals and magazine of natural history including zoology, botany and geology XIX 326 2009 Adult morphology and terminology. Pp. 9–50 Manual of Central American Diptera 1 NRC Research Press, Ottawa, Canada, xi + 714 pp pp. Ottawa, Canada xi + 714 pp pp. 2015 Seven new species of Spathidexia Townsend (Diptera: Tachinidae) reared from caterpillars in Area de Conservación Guanacaste, Costa Rica Biodiversity Data Journal 3 e4597 10.3897/bdj.3.e4597 2015 Three new species of Trigonospila Pokorny (Diptera: Tachinidae), from Area de Conservacion Guanacaste, northwestern Costa Rica, with a key for their identification Biodiversity Data Journal 3 4595 4595 10.3897/BDJ.3.e4595 2014 Revision of the New World species of Houghia Coquillett (Diptera, Tachinidae) reared from caterpillars in Area de Conservación Guanacaste, Costa Rica Zootaxa 3858 1 1 10.11646/zootaxa.3858.1.1 2015 Nine new species of Itaplectops (Diptera: Tachinidae) reared from caterpillars in Area de Conservación Guanacaste, northwestern Costa Rica, with a key to Itaplectops species Biodiversity Data Journal 3 e4596 10.3897/bdj.3.e4596 2015 Three new species of Ametadoria Townsend (Diptera: Tachinidae) from Area de Conservación Guanacaste, Costa Rica Biodiversity Data Journal 3 e5039 10.3897/bdj.3.e5039 2014 A new species of Cordyligaster Macquart, reared from caterpillars in Area de Conservacion Guanacaste, northwestern Costa Rica Biodiversity Data Journal 2 e4174 10.3897/bdj.2.e4174 2016 Two new species of Erythromelana Townsend, 1919 (Diptera: Tachinidae) from Area de Conservación Guanacaste in northwestern Costa Rica Biodiversity Data Journal 4 e7386 10.3897/bdj.4.e7386 2016 A new species of Phosocephala Townsend, 1908 (Diptera: Tachinidae) from Area de Conservación Guanacaste in northwestern Costa Rica Biodiversity Data Journal 4 e7863 10.3897/bdj.4.e7863 2004 Ten species in one: DNA barcoding reveals cryptic species in the neotropical skipper butterfly Astraptes fulgerator Proceedings of the National Academy of Sciences 101 41 14812 14817 10.1073/pnas.0406166101 2006 An inexpensive, automation-friendly protocol for recovering high-quality DNA Molecular Ecology Notes 6 4 998 1002 10.1111/j.1471-8286.2006.01428.x 2011 Joining inventory by parataxonomists with DNA barcoding of a large complex tropical conserved wildland in northwestern Costa Rica PLoS ONE 6 8 e18123 10.1371/journal.pone.0018123 Area de Conservación Guanacaste (ACG), northwestern Costa Rica: Caterpillars, pupae, butterflies & moths http://janzen.sas.upenn.edu 2016-06-21T00:00:00+03:00 2016 DNA barcoding the Lepidoptera inventory of a large complex tropical conserved wildland, Area de Conservacion Guanacaste, northwestern Costa Rica 1 Genome 59 9 641 660 10.1139/gen-2016-0005 2009 Integration of DNA barcoding into an ongoing inventory of complex tropical biodiversity Molecular Ecology Resources 9 1 26 10.1111/j.1755-0998.2009.02628.x Nei M Kumar S 2000 Molecular Evolution and Phylogenetics Oxford University Press New York 333 1983 Classification, phylogeny and zoogeography of the North American species of Siphona Meigen (Diptera: Tachinidae) Quaestiones Entomologicae 18 1982 261 380 2007 BOLD: The Barcode of Life Data System (http://www.barcodinglife.org). Molecular ecology notes 7 3 355 364 10.1111/j.1471-8286.2007.01678.x Sjöberg Fredrik 2014 The Fly Trap Pantheon 288 110187015X 2007 DNA barcodes affirm that 16 species of apparently generalist tropical parasitoid flies (Diptera, Tachinidae) are not all generalists Proceedings of the National Academy of Sciences 104 12 4967 4972 10.1073/pnas.0700050104 2006 DNA barcodes reveal cryptic host-specificity within the presumed polyphagous members of a genus of parasitoid flies (Diptera: Tachinidae) Proceedings of the National Academy of Sciences 103 10 3657 3662 10.1073/pnas.0511318103 2008 Extreme diversity of tropical parasitoid wasps exposed by iterative integration of natural history, DNA barcoding, morphology, and collections Proceedings of the National Academy of Sciences 105 34 12359 12364 10.1073/pnas.0805319105 2012 Wolbachia and DNA Barcoding Insects: Patterns, Potential, and Problems PLoS ONE 7 5 e36514 10.1371/journal.pone.0036514 2013 MEGA6: Molecular Evolutionary Genetics Analysis Version 6.0 Molecular Biology and Evolution 30 12 2725 2729 10.1093/molbev/mst197 1985 A taxonomic conspectus of the Blondeliini of North and Central America and the West Indies (Diptera: Tachinidae) Memoirs of the Entomological Society of Canada 117 132 1 130 10.4039/entm117132fv

General morphology of Calolydella adelinamoralesae sp. n.; a–c: male holotype, voucher n. DHJPAR0017775; d–f: female paratype, voucher n. DHJPAR0038669.

5804208A-6D09-5950-9F1D-AA8DDACB356C 10.3897/BDJ.6.e11223.figure1a

habitus in dorsal view

https://binary.pensoft.net/fig/90672 9293741D-EE26-5917-B9F1-8A9366A38DA1 10.3897/BDJ.6.e11223.figure1b

head in frontal view

https://binary.pensoft.net/fig/90671 81569C4C-6EC9-5817-B876-64A2B5E8C8FA 10.3897/BDJ.6.e11223.figure1c

habitus in lateral view

https://binary.pensoft.net/fig/90674 906085C5-9A04-5C4A-A85D-3D46E2432286 10.3897/BDJ.6.e11223.figure1d

habitus in dorsal view

https://binary.pensoft.net/fig/90675 908FC69C-5560-54DA-8B03-78189662526F 10.3897/BDJ.6.e11223.figure1e

head in frontal view

https://binary.pensoft.net/fig/90673 D6476C77-1DF1-572F-87D0-64695476AEE4 10.3897/BDJ.6.e11223.figure1f

habitus in lateral view

https://binary.pensoft.net/fig/90676

Male terminalia of Calolydella adelinamoralesae sp. n.; a–c: paratype, voucher n. DHJPAR0019750.

0A4346EB-7A77-5F27-A9D8-7A9C77479178 10.3897/BDJ.6.e11223.figure2a

lateral view

https://binary.pensoft.net/fig/117741 1BFA140C-FBED-5712-8DA3-7FF8202E55CE 10.3897/BDJ.6.e11223.figure2b

dorsal view

https://binary.pensoft.net/fig/95836 CDFA124C-014F-5082-BF39-8CC557CF7708 10.3897/BDJ.6.e11223.figure2c

sternite 5 in ventral view

https://binary.pensoft.net/fig/95837

General morphology of Calolydella alexanderjamesi sp. n.; a–c: male holotype, voucher n. DHJPAR0017782.

C24CC551-204F-5648-9F3F-E998CDE33192 10.3897/BDJ.6.e11223.figure3a

habitus in dorsal view

https://binary.pensoft.net/fig/90710 74026052-DE5D-59AE-A2F8-A591C7B05E40 10.3897/BDJ.6.e11223.figure3b

head in frontal view

https://binary.pensoft.net/fig/90709 EF849679-BCA6-5CA5-93EA-A126E7FCB3AB 10.3897/BDJ.6.e11223.figure3c

habitus in lateral view

https://binary.pensoft.net/fig/90711

General morphology of Calolydella argentea sp. n.; a–c: female holotype, voucher n. DHJPAR0042289.

DF9677E0-2E27-5312-BAAF-D553668202A7 10.3897/BDJ.6.e11223.figure4a

habitus in dorsal view

https://binary.pensoft.net/fig/90501 1D922B08-8FB4-5F44-BE90-1F7BD2112436 10.3897/BDJ.6.e11223.figure4b

head in frontal view

https://binary.pensoft.net/fig/90499 D3399DA8-DC05-531F-AA09-9C7179CD157D 10.3897/BDJ.6.e11223.figure4c

habitus in lateral view

https://binary.pensoft.net/fig/90500

General morphology of Calolydella aureofacies sp. n.; a–c: female holotype, voucher n. DHJPAR0029582.

CC301E4E-77DA-52D0-A0C3-CBB602CAC154 10.3897/BDJ.6.e11223.figure5a

habitus in dorsal view

https://binary.pensoft.net/fig/94708 D7AE7415-4267-56F3-998C-4A8D001B6CAE 10.3897/BDJ.6.e11223.figure5b

head in frontal view

https://binary.pensoft.net/fig/94707 7D7C43DD-78B0-5C54-A49A-15B163DDCE55 10.3897/BDJ.6.e11223.figure5c

habitus in lateral view

https://binary.pensoft.net/fig/94709

General morphology of Calolydella bicolor sp. n.; a–c: female holotype, voucher n. DHJPAR0054050.

455C4CD3-7A01-557C-9622-0E4033D2CA74 10.3897/BDJ.6.e11223.figure6a

habitus in dorsal view

https://binary.pensoft.net/fig/94711 869D96B9-13B6-5476-89BF-4A8FBE671D41 10.3897/BDJ.6.e11223.figure6b

head in frontal view

https://binary.pensoft.net/fig/94710 B5985117-8C8B-5230-85BF-1E7303F94E7B 10.3897/BDJ.6.e11223.figure6c

habitus in lateral view

https://binary.pensoft.net/fig/94712

General morphology of Calolydella bifissus sp. n.; a–c: male holotype, voucher n. DHJPAR0017132; d–f: female paratype, voucher n. 07-SRNP-40315.

70C4174A-296A-5836-94AA-81D5677B6A3D 10.3897/BDJ.6.e11223.figure7a

habitus in dorsal view

https://binary.pensoft.net/fig/90666 4842CB29-FBCD-5A92-BDA1-26D8449B7463 10.3897/BDJ.6.e11223.figure7b

head in frontal view

https://binary.pensoft.net/fig/90665 1C10FA95-CF50-523D-B936-D976A75FE530 10.3897/BDJ.6.e11223.figure7c

habitus in lateral view

https://binary.pensoft.net/fig/90669 8B9D41F2-C882-5B4D-91EE-9A3AEDF792DA 10.3897/BDJ.6.e11223.figure7d

habitus in dorsal view

https://binary.pensoft.net/fig/90667 6C9544B9-CE6E-5CC1-8DAE-E3BD528BEC73 10.3897/BDJ.6.e11223.figure7e

head in frontal view

https://binary.pensoft.net/fig/90668 9CF59D18-593A-5766-957D-C3770263C29B 10.3897/BDJ.6.e11223.figure7f

habitus in lateral view

https://binary.pensoft.net/fig/90670

Male terminalia of Calolydella bifissus sp. n.; a–c: paratype, voucher n. 07-SRNP-40315.

217A9050-3698-5931-8898-E2F20FA2C114 10.3897/BDJ.6.e11223.figure8a

lateral view

https://binary.pensoft.net/fig/117735 66DD7BF7-A1E3-5400-8F30-0DC6879BF2FB 10.3897/BDJ.6.e11223.figure8b

dorsal view

https://binary.pensoft.net/fig/96579 7DEFBFAC-4F25-5228-955B-D53E6690E21B 10.3897/BDJ.6.e11223.figure8c

sternite 5 in ventral view

https://binary.pensoft.net/fig/96580

General morphology of Calolydella crocata sp. n.; a–c: male holotype, voucher n. DHJPAR0053232.

03A102C3-C9CA-5D41-B06A-5EA26021611E 10.3897/BDJ.6.e11223.figure9a

habitus in dorsal view

https://binary.pensoft.net/fig/94714 2CE37857-1028-5FAC-BA59-6B5D483F6430 10.3897/BDJ.6.e11223.figure9b

head in frontal view

https://binary.pensoft.net/fig/94713 15F0415F-396D-504F-8F64-9A2BA8612C65 10.3897/BDJ.6.e11223.figure9c

habitus in lateral view

https://binary.pensoft.net/fig/94715

Male terminalia of Calolydella crocata sp. n.; a–c: paratype, voucher n. DHJPAR0055883.

6AAE188F-15E7-59C3-A8D3-A6ECCB3EB39D 10.3897/BDJ.6.e11223.figure10a

lateral view

https://binary.pensoft.net/fig/117742 7A5EEC79-250A-50E6-930A-BF999D47F0D0 10.3897/BDJ.6.e11223.figure10b

dorsal view

https://binary.pensoft.net/fig/95833 A81F1743-7652-5947-98A4-B8CFA672523B 10.3897/BDJ.6.e11223.figure10c

sternite 5 in ventral view

https://binary.pensoft.net/fig/95834

General morphology of Calolydella destituta sp. n.; a–c: male holotype, voucher n. DHJPAR0017799; d–f: female paratype, voucher n. DHJPAR0017771.

5D2A8AE9-9365-5568-BDE7-041330FBE9E9 10.3897/BDJ.6.e11223.figure11a

habitus in dorsal view

https://binary.pensoft.net/fig/90840 3CDC2F0D-6217-5497-BD0B-9BFFECCD7D58 10.3897/BDJ.6.e11223.figure11b

head in frontal view

https://binary.pensoft.net/fig/90839 7A414671-82CE-5E75-9BA2-5B572D05B477 10.3897/BDJ.6.e11223.figure11c

habitus in lateal view

https://binary.pensoft.net/fig/90846 109FC9E2-4014-5432-8195-AB6825218F43 10.3897/BDJ.6.e11223.figure11d

habitus in dorsal view

https://binary.pensoft.net/fig/90844 899FE5DE-1C1C-5DF4-93C9-DE684CC7B927 10.3897/BDJ.6.e11223.figure11e

head in frontal view

https://binary.pensoft.net/fig/90842 EEDC26AB-F979-5DF7-81EA-28C7B9E6AB5A 10.3897/BDJ.6.e11223.figure11f

habitus in lateral view

https://binary.pensoft.net/fig/90841

Male terminalia of Calolydella destituta sp. n.; a–c: paratype, voucher n. DHJPAR0017768.

20E5009A-5603-5AC4-85EF-D4F0D3477804 10.3897/BDJ.6.e11223.figure12a

lateral view

https://binary.pensoft.net/fig/117779 732C53D3-7A04-5C85-8A42-6D680E34C49D 10.3897/BDJ.6.e11223.figure12b

dorsal view

https://binary.pensoft.net/fig/152184 EDC58F06-452A-555D-B5E9-4AF9369871D4 10.3897/BDJ.6.e11223.figure12c

sternite 5 in ventral view

https://binary.pensoft.net/fig/95844

General morphology of Calolydella discalis sp. n.; a–c: male holotype, voucher n. DHJPAR0017774.

540EBC0D-AD7F-5D18-97C7-931FD782F013 10.3897/BDJ.6.e11223.figure13a

habitus in dorsal view

https://binary.pensoft.net/fig/90706 DF97BCAC-BFA0-5872-9F56-FDE68491CB8C 10.3897/BDJ.6.e11223.figure13b

head in frontal view

https://binary.pensoft.net/fig/90705 52C0463E-0DBB-5964-9260-523F8920618D 10.3897/BDJ.6.e11223.figure13c

habitus in lateral view

https://binary.pensoft.net/fig/90708

Male terminalia of Calolydella discalis sp. n.; a–c: paratype, voucher n. DHJPAR0017773

8787B712-DF7B-5BD0-9646-AA96B53F9F5D 10.3897/BDJ.6.e11223.figure14a

lateral view

https://binary.pensoft.net/fig/173114 26F28D6D-590C-5FC2-BCE9-EB2306F32000 10.3897/BDJ.6.e11223.figure14b

dorsal view

https://binary.pensoft.net/fig/173115 9AE5B44E-315C-53C8-9EC4-5915B5EBDC4F 10.3897/BDJ.6.e11223.figure14c

sternite 5 in ventral view

https://binary.pensoft.net/fig/95847

General morphology of Calolydella erasmocoronadoi sp. n.; a–c: male holotype, voucher n. DHJPAR0039246; d–f: female paratype, voucher n. 04-SRNP-55202.

EB20825A-EF52-5F1B-AAEA-7767EE13EE23 10.3897/BDJ.6.e11223.figure15a

habitus in dorsal view

https://binary.pensoft.net/fig/94697 AD31E19B-4537-5E96-B578-1CD0BFE2E5F5 10.3897/BDJ.6.e11223.figure15b

head in frontal view

https://binary.pensoft.net/fig/94679 AC40ED06-42CD-5E8B-8245-2F64DE4183B9 10.3897/BDJ.6.e11223.figure15c

habitus in lateral view

https://binary.pensoft.net/fig/94698 543CE242-C990-5B2D-8226-FBF85FDFBD15 10.3897/BDJ.6.e11223.figure15d

habitus in dorsal view

https://binary.pensoft.net/fig/94699 6D89CB2D-13C1-52F8-967F-C06314B764C4 10.3897/BDJ.6.e11223.figure15e

head in frontal view

https://binary.pensoft.net/fig/94684 83D6A4D9-2087-5602-842C-DD9471D40468 10.3897/BDJ.6.e11223.figure15f

habitus in lateral view

https://binary.pensoft.net/fig/94700

Male terminalia of Calolydella erasmocoronadoi sp. n.; a–c: paratype, voucher n. DHJPAR0011690.

0351B032-FCD3-50EE-AA7E-A5DE7A5847EE 10.3897/BDJ.6.e11223.figure16a

lateral view

https://binary.pensoft.net/fig/117746 B81B29CE-62F3-5211-8CE9-6C563F19CEC6 10.3897/BDJ.6.e11223.figure16b

dorsal view

https://binary.pensoft.net/fig/95825 5559B710-D851-54DC-8C83-25F0504D32A2 10.3897/BDJ.6.e11223.figure16c

sternite 5 in ventral view

https://binary.pensoft.net/fig/95827 CFFEE930-E385-59E2-AB21-C3E9D8629347 10.3897/BDJ.6.e11223.figure16d https://binary.pensoft.net/fig/104838

General morphology of Calolydella felipechavarriai sp. n.; a–c: male holotype, voucher n. DHJPAR0050491; d–f: female paratype, voucher n. DHJPAR0050489.

1D75F1BD-66B3-5209-8665-76E55739049D 10.3897/BDJ.6.e11223.figure17a

habitus in dorsal view

https://binary.pensoft.net/fig/94692 A7311CC5-136E-568A-8005-D931E64F7106 10.3897/BDJ.6.e11223.figure17b

head in frontal view

https://binary.pensoft.net/fig/94687 042B8A39-80B2-5981-AC23-5DF354AAA276 10.3897/BDJ.6.e11223.figure17c

habitus in lateral view

https://binary.pensoft.net/fig/94693 E7C82220-B57C-5C80-8DFC-38F647935A9D 10.3897/BDJ.6.e11223.figure17d

habitus in dorsal view

https://binary.pensoft.net/fig/94694 3A8517E2-7804-566D-9D5E-AE957792EC14 10.3897/BDJ.6.e11223.figure17e

head in frontal view

https://binary.pensoft.net/fig/94689 5EB790F4-BC04-5261-8FF7-1D2A97587251 10.3897/BDJ.6.e11223.figure17f

habitus in lateral view

https://binary.pensoft.net/fig/94696

Male terminalia of Calolydella felipechavarriai sp. n.; a–c: paratype, voucher n. 12-SRNP-79034

EDD00BC6-931F-5783-B4B4-4877C31113BF 10.3897/BDJ.6.e11223.figure18a

lateral view

https://binary.pensoft.net/fig/117747 8688B983-F8EF-5C9B-AC8B-2EC1322DD5FB 10.3897/BDJ.6.e11223.figure18b

dorsal view

https://binary.pensoft.net/fig/95831 C2FA7A90-03AB-509F-AE2A-4823156C87F2 10.3897/BDJ.6.e11223.figure18c

sternite 5 in ventral view

https://binary.pensoft.net/fig/95830

General morphology of Calolydella fredriksjobergi sp. n.; a–c: male holotype, voucher n. DHJPAR0011711; d–f: female paratype, voucher n. DHJPAR0054013.

7B308781-A00A-58FE-83E3-82981EBB46F5 10.3897/BDJ.6.e11223.figure19a

habitus in dorsal view

https://binary.pensoft.net/fig/90494 A28768FC-368F-5FD7-BE01-8721673CB7F8 10.3897/BDJ.6.e11223.figure19b

head in frontal view

https://binary.pensoft.net/fig/90493 4587A05B-C78D-5BCC-9611-46462D7499FB 10.3897/BDJ.6.e11223.figure19c

habitus in lateral view

https://binary.pensoft.net/fig/90495 6A83F395-E1CF-5813-A837-7FA5B2C75533 10.3897/BDJ.6.e11223.figure19d

habitus in dorsal view

https://binary.pensoft.net/fig/90496 A8417D51-6B03-54A1-A10F-9F57A92018B5 10.3897/BDJ.6.e11223.figure19e

head in frontal view

https://binary.pensoft.net/fig/90497 7CB5811C-FEA3-5AE3-A759-C5CB57AFEA2C 10.3897/BDJ.6.e11223.figure19f

habitus in lateral view

https://binary.pensoft.net/fig/90498

Male terminalia of Calolydella fredriksjobergi sp. n.; a–c: paratype, voucher n. DHJPAR0050643.

2F552903-99ED-583C-ABCD-00BF20B81C04 10.3897/BDJ.6.e11223.figure20a

lateral view

https://binary.pensoft.net/fig/117748 6B83ED61-6DE9-5CCE-A6C6-D0FA3EA86CBB 10.3897/BDJ.6.e11223.figure20b

dorsal view

https://binary.pensoft.net/fig/94671 FA5BE910-F678-5EB6-8AAF-495D6BC5261B 10.3897/BDJ.6.e11223.figure20c

sternite 5 in ventral view

https://binary.pensoft.net/fig/94669

General morphology of Calolydella inflatipalpis sp. n.; a–c: male holotype, voucher n. DHJPAR0016181; d–f: female paratype, voucher n. DHJPAR0016169.

256A2391-7BA9-5F7F-983D-0B1B2BE8F88E 10.3897/BDJ.6.e11223.figure21a

habitus in dorsal view

https://binary.pensoft.net/fig/90517 2E3D2549-5894-58A1-90AC-943ACC846153 10.3897/BDJ.6.e11223.figure21b

head in frontal view

https://binary.pensoft.net/fig/90516 31E73E48-3CAE-5E88-8E3B-81929F9C987F 10.3897/BDJ.6.e11223.figure21c

habitus in lateral view

https://binary.pensoft.net/fig/90519 E0787F38-173C-5238-AFDD-B5D304F52EB4 10.3897/BDJ.6.e11223.figure21d

habitus in dorsal view

https://binary.pensoft.net/fig/90520 A4BDF2C4-E8F9-5138-8925-B01CAF48370F 10.3897/BDJ.6.e11223.figure21e

head in frontal view

https://binary.pensoft.net/fig/90518 4BB5C313-2650-5622-A6E8-91D19FC58DA0 10.3897/BDJ.6.e11223.figure21f

habitus in lateral view

https://binary.pensoft.net/fig/90521

Male terminalia of Calolydella inflatipalpis sp. n.; a–c: paratype, voucher n. DHJPAR0016170.

FF0C12E9-4AB3-59EA-9D72-80E4605575F3 10.3897/BDJ.6.e11223.figure22a

lateral view

https://binary.pensoft.net/fig/117737 16341EF9-D399-59E2-B8C5-E4E73ADE5E66 10.3897/BDJ.6.e11223.figure22b

dorsal view

https://binary.pensoft.net/fig/95849 D4D2D7A4-859F-5B5B-BDBC-BE947A1C0167 10.3897/BDJ.6.e11223.figure22c

sternite 5 in ventral view

https://binary.pensoft.net/fig/95850

General morphology of Calolydella interrupta sp. n.; a–c: male holotype, voucher n. DHJPAR0029598; d–f: female paratype, voucher n. 08-SRNP-36564.

B6341AFA-0E5F-56FB-9301-841BF80FAFB0 10.3897/BDJ.6.e11223.figure23a

habitus in dorsal view

https://binary.pensoft.net/fig/94702 72732486-B005-5956-B6C6-043EF137A3EE 10.3897/BDJ.6.e11223.figure23b

head in frontal view

https://binary.pensoft.net/fig/94701 7D4824D8-ABB9-5FB3-8FAC-3C3F654E2D9A 10.3897/BDJ.6.e11223.figure23c

habitus in lateral view

https://binary.pensoft.net/fig/94704 DCF89F06-009F-541A-ABD9-487262D0F813 10.3897/BDJ.6.e11223.figure23d

habitus in frontal view

https://binary.pensoft.net/fig/94705 0F92EACB-F5A3-56E0-9BE4-FE406DEE7DC6 10.3897/BDJ.6.e11223.figure23e

head in frontal view

https://binary.pensoft.net/fig/94703 8F0DA641-6F42-52DF-9B29-7E2954579C4D 10.3897/BDJ.6.e11223.figure23f

habitus in lateral view

https://binary.pensoft.net/fig/94706

General morphology of Calolydella nigripalpis sp. n.; a–c: male holotype, voucher n. DHJPAR0017812.

948B0402-185E-5D6A-ADA4-72EB2E5FAF60 10.3897/BDJ.6.e11223.figure24a

habitus in dorsal view

https://binary.pensoft.net/fig/90523 A866D21E-FB06-55B3-BC96-D386D6BD306D 10.3897/BDJ.6.e11223.figure24b

head in frontal view

https://binary.pensoft.net/fig/90522 05A3C355-C134-5820-BD47-97D434E0E96D 10.3897/BDJ.6.e11223.figure24c

habitus in lateral view

https://binary.pensoft.net/fig/90524

Male terminalia of Calolydella nigripalpis sp. n.; a–c: paratype, voucher n. DHJPAR0017811.

6850425D-EC79-53D0-A90D-056288215562 10.3897/BDJ.6.e11223.figure25a

lateral view

https://binary.pensoft.net/fig/117739 77BC84BA-C00E-5423-A3AC-04AB07E7C9E0 10.3897/BDJ.6.e11223.figure25b

dorsal view

https://binary.pensoft.net/fig/95852 4F31CD8B-1952-5F86-9A69-531D39BD15C2 10.3897/BDJ.6.e11223.figure25c

sternite 5 in ventral view

https://binary.pensoft.net/fig/95853

General morphology of Calolydella omissa sp. n.; a–c: male holotype, voucher n. DHJPAR0016605; d–f: female paratype, voucher n. 06-SRNP-43825.

9D82470A-F52E-57FA-A77F-846B81F85698 10.3897/BDJ.6.e11223.figure26a

habitus in dorsal view

https://binary.pensoft.net/fig/90505 68BD88B5-D9A6-5BE7-B138-0B328B1B0B36 10.3897/BDJ.6.e11223.figure26b

head in frontal view

https://binary.pensoft.net/fig/90504 B094F838-45A5-5075-9C93-ED4CE1B076B5 10.3897/BDJ.6.e11223.figure26c

habitus in lateral view

https://binary.pensoft.net/fig/90508 06648A40-A328-5677-B369-038AA7BB2219 10.3897/BDJ.6.e11223.figure26d

habitus in dorsal view

https://binary.pensoft.net/fig/90507 A9FDF067-2AD6-5E57-87A9-8D0A6E08616C 10.3897/BDJ.6.e11223.figure26e

head in frontal view

https://binary.pensoft.net/fig/90506 ADBF149F-B5C7-5B7E-B07A-0ABB6DDF9E5F 10.3897/BDJ.6.e11223.figure26f

habitus in lateral view

https://binary.pensoft.net/fig/90509

Male terminalia of Calolydella omissa sp. n.; a–c: paratype, voucher n. 06-SRNP-43825.

44EC723C-5C5B-5ACB-927B-5F44F66F93DB 10.3897/BDJ.6.e11223.figure27a

lateral view

https://binary.pensoft.net/fig/117762 551E4CDA-6BF0-557D-AAE1-BEEAC6BAA6F1 10.3897/BDJ.6.e11223.figure27b

dorsal view

https://binary.pensoft.net/fig/94673 2D2E949B-1D40-55DA-8E6F-B592A0EA94FA 10.3897/BDJ.6.e11223.figure27c

sternite 5 in ventral view

https://binary.pensoft.net/fig/94674

General morphology of Calolydella ordinalis sp. n.; a–c: male holotype, voucher n. DHJPAR0017787; d–f: female paratype, voucher n. DHJPAR0017792.

7DA59D00-F335-5CDA-BCDE-551B5B62724F 10.3897/BDJ.6.e11223.figure28a

habitus in dorsal view

https://binary.pensoft.net/fig/90511 3780FCF9-1D2F-5F3A-AD76-8484BB0C3E06 10.3897/BDJ.6.e11223.figure28b

head in frontal view

https://binary.pensoft.net/fig/90510 817C57F5-1487-5644-AD11-C666B80DC2B4 10.3897/BDJ.6.e11223.figure28c

habitus in lateral view

https://binary.pensoft.net/fig/90514 EC37FB2E-5EAD-5B7E-B371-A89F0D4A80D7 10.3897/BDJ.6.e11223.figure28d

habitus in dorsal view

https://binary.pensoft.net/fig/90513 91CF8DF0-B33D-50EC-89F9-5D2374884CFA 10.3897/BDJ.6.e11223.figure28e

head in frontal view

https://binary.pensoft.net/fig/90512 A112A0BB-4B04-5D72-A4C2-DDB2F90E0034 10.3897/BDJ.6.e11223.figure28f

habitus in lateral view

https://binary.pensoft.net/fig/90515

Male terminalia of Calolydella ordinalis sp. n.; a–c: paratype, voucher n. DHJPAR0017791.

3178A412-3C91-52B3-932E-89190D00FDB5 10.3897/BDJ.6.e11223.figure29a

lateral view

https://binary.pensoft.net/fig/117771 22801627-FD08-51DF-AFDF-39F627EB8639 10.3897/BDJ.6.e11223.figure29b

dorsal view

https://binary.pensoft.net/fig/94676 66E06B05-4A1A-50DF-A0F0-2677D7214B81 10.3897/BDJ.6.e11223.figure29c

sternite 5 in ventral view

https://binary.pensoft.net/fig/94677

General morphology of Calolydella renemalaisei sp. n.; a–c: male holotype, voucher n. DHJPAR0011709.

C5068533-08BA-5F08-909A-84572C2D93AE 10.3897/BDJ.6.e11223.figure30a

habitus in lateral view

https://binary.pensoft.net/fig/90530 23AA7D62-E5BF-59F3-8BCF-A9B9500BA3F3 10.3897/BDJ.6.e11223.figure30b

head in frontal view

https://binary.pensoft.net/fig/90529 86FA2816-FFC1-5976-B9FA-D459DB1CD414 10.3897/BDJ.6.e11223.figure30c

habitus in lateral view

https://binary.pensoft.net/fig/90531

Male terminalia of Calolydella renemalaisei sp. n.; a–c: paratype, voucher n. DHJPAR0057608.

11290F88-FC71-54FC-ACE3-BA132F45DB01 10.3897/BDJ.6.e11223.figure31a

lateral view

https://binary.pensoft.net/fig/117778 88ECF57F-1E1D-582E-BE0A-A655A18E27E3 10.3897/BDJ.6.e11223.figure31b

dorsal view

https://binary.pensoft.net/fig/96582 8D4AF0CB-F469-5082-A9E6-6491227C513B 10.3897/BDJ.6.e11223.figure31c

sternite 5 in ventral view

https://binary.pensoft.net/fig/96583

General morphology of Calolydella susanaroibasae sp. n.; a–c: male holotype, voucher n. DHJPAR0057699; d–f: female paratype, voucher n. 15-SRNP-1269.

35733779-AC66-5314-9FC9-819EBDB2833C 10.3897/BDJ.6.e11223.figure32a

habitus in dorsal view

https://binary.pensoft.net/fig/90533 C89213D9-5620-500C-88B5-D0E263178798 10.3897/BDJ.6.e11223.figure32b

head in frontal view

https://binary.pensoft.net/fig/90532 A1B09DC2-38F2-506B-AA18-993B8072D3FB 10.3897/BDJ.6.e11223.figure32c

habitus in lateral view

https://binary.pensoft.net/fig/90535 CA6DE361-6446-582F-BFC0-A2AEC110CF3A 10.3897/BDJ.6.e11223.figure32d

habitus in dorsal view

https://binary.pensoft.net/fig/90536 9B33974A-AACA-57AD-8CA4-26E373828293 10.3897/BDJ.6.e11223.figure32e

head in frontal view

https://binary.pensoft.net/fig/90534 15D8F639-FCD0-5A24-AEED-F14A2F62B670 10.3897/BDJ.6.e11223.figure32f

habitus in lateral view

https://binary.pensoft.net/fig/90537

General morphology of Calolydella tanyadapkeyae sp. n.; a–c: female holotype, voucher n. DHJPAR0011701.

CD45A994-9ED8-5D14-88F1-C3E14C54CD22 10.3897/BDJ.6.e11223.figure33a

habitus in lateral view

https://binary.pensoft.net/fig/90694 7226811C-CDC1-5DA0-887C-CE1F7C3553E0 10.3897/BDJ.6.e11223.figure33b

head in frontal view

https://binary.pensoft.net/fig/90693 5D90840B-73B4-58E2-BC95-112C89A177AD 10.3897/BDJ.6.e11223.figure33c

habitus in lateral view

https://binary.pensoft.net/fig/90696

General morphology of Calolydella tenebrosa sp. n.; a–c: male holotype, voucher n. DHJPAR0017810; d–f: female paratype, voucher n. DHJPAR0053996

7463D206-46F9-5091-ACFE-D95EACC5EB82 10.3897/BDJ.6.e11223.figure34a

habitus in dorsal view

https://binary.pensoft.net/fig/90658 DEE09917-BFBE-5D6A-B8B9-F2A0C059E85D 10.3897/BDJ.6.e11223.figure34b

head in frontal view

https://binary.pensoft.net/fig/90657 774B6C6E-3404-52D1-A660-6A1D01441276 10.3897/BDJ.6.e11223.figure34c

habitus in lateral view

https://binary.pensoft.net/fig/90661 5965FDBE-629D-5E0D-A86C-0E97957A4CC4 10.3897/BDJ.6.e11223.figure34d

habitus in dorsal view

https://binary.pensoft.net/fig/90659 2DFC4DDF-9444-5AEB-AFEB-E219C91A6B5B 10.3897/BDJ.6.e11223.figure34e

head in frontal view

https://binary.pensoft.net/fig/90660 9AA40CB6-95D7-58D9-906B-B4BCD5E2B94B 10.3897/BDJ.6.e11223.figure34f

habitus in lateral view

https://binary.pensoft.net/fig/90664

General morphology of Calolydella timjamesi sp. n.; a–c: male holotype, voucher n. DHJPAR0017781; d–f: female paratype, voucher n. DHJPAR0016192.

52F28CE3-7BD3-5CFA-BBE3-E3E18D34DC38 10.3897/BDJ.6.e11223.figure35a

habitus in dorsal view

https://binary.pensoft.net/fig/90698 2669ED45-49C5-59B0-B595-21116AA34C75 10.3897/BDJ.6.e11223.figure35b

head in frontal view

https://binary.pensoft.net/fig/90697 03DBF841-927D-5CFF-8320-3796A26B2C23 10.3897/BDJ.6.e11223.figure35c

habitus in lateral view

https://binary.pensoft.net/fig/90699 C6749717-546D-5843-AA52-2D4B043044D3 10.3897/BDJ.6.e11223.figure35d

habitus in dorsal view

https://binary.pensoft.net/fig/90701 9354B1D5-A04F-5BDC-B9BB-25499AB3A589 10.3897/BDJ.6.e11223.figure35e

head in frontal view

https://binary.pensoft.net/fig/90700 C3712494-1AA8-5233-9720-DE90CE70500E 10.3897/BDJ.6.e11223.figure35f

habitus in lateral

https://binary.pensoft.net/fig/90702

Male terminalia of Calolydella timjamesi sp. n.; a–c: paratype, voucher n. DHJPAR0017783.

1E83E2D5-5CE7-5EDD-8A8B-EE04AB6E1287 10.3897/BDJ.6.e11223.figure36a

lateral view

https://binary.pensoft.net/fig/117589 AB6A6603-19F2-51BF-9545-5288681A7CF5 10.3897/BDJ.6.e11223.figure36b

dorsal view

https://binary.pensoft.net/fig/94662 07A6C06E-776B-5088-9BE1-CF6D54EAC5EC 10.3897/BDJ.6.e11223.figure36c

sternite 5 in ventral view

https://binary.pensoft.net/fig/94663

General morphology of Calolydella virginiajamesae sp. n.; a–c: male holotype, voucher n. DHJPAR0017770; d–f: female paratype, voucher n. DHJPAR0045648.

0586C1B4-CB72-5621-97C4-0C7B28AFCCE2 10.3897/BDJ.6.e11223.figure37a

habitus in dorsal view

https://binary.pensoft.net/fig/90682 3C95B7E2-627E-5FFD-8A54-E52E61E77B16 10.3897/BDJ.6.e11223.figure37b

head in frotnal view

https://binary.pensoft.net/fig/90683 D89294E8-A229-5949-99BE-CC44FDA2912A 10.3897/BDJ.6.e11223.figure37c

habitus in lateral view

https://binary.pensoft.net/fig/90688 9D441E7F-F11E-561F-9AAF-453CC5E84D4D 10.3897/BDJ.6.e11223.figure37d

habitus in dorsal view

https://binary.pensoft.net/fig/90689 ABF10BBC-74F3-5740-9795-1B1ADC328C77 10.3897/BDJ.6.e11223.figure37e

head in frontal view

https://binary.pensoft.net/fig/90684 9924F981-D389-553D-8D24-7FBEEFD7C3B8 10.3897/BDJ.6.e11223.figure37f

habitus in lateral view

https://binary.pensoft.net/fig/90691

Male terminalia of Calolydella virginiajamesae sp. n.; a–c: paratype, voucher n. DHJPAR0046433.

B7D9E41F-01B3-54E9-B7B3-8E799F1DBF71 10.3897/BDJ.6.e11223.figure38a

lateral view

https://binary.pensoft.net/fig/117740 60E3AE97-02B5-5159-A7D7-051F1A918470 10.3897/BDJ.6.e11223.figure38b

dorsal view

https://binary.pensoft.net/fig/96304 515412C2-8005-584D-A439-3B8CFA432BA3 10.3897/BDJ.6.e11223.figure38c

sternite 5 in ventral view

https://binary.pensoft.net/fig/96303 F817ACAF-C029-541F-813C-DB68CF39A54A 10.3897/BDJ.6.e11223.figure39 1223073

Maximum Likelihood tree showing DNA barcode variation within ACG species of Calolydella. The inter- and intra-specific variation within the DNA barcode region for the 23 species of Calolydella from ACG is illustrated using a Maximum Likelihood tree based on the General Time Reversible model (GTR) (Nei and Kumar 2000). GTR was selected due to the lowest BIC scores (Bayesian Information Criterion) of the Maximum Likelihood fits of 23 different nucleotide substitution models run in MEGA6 (Tamura et al. 2013). Tip labels include species name|specimen code|host family of each species. A habitus image in lateral view of each holotype is presented with the tree.

https://binary.pensoft.net/fig/117247