lateral view. Arrows T10–T12 show pronotum (T10), lateral pronotal carina (T11), and hypomeron (T12);
	Part of: Montanaro G, Balhoff JP, Girón JC, Söderholm M, Tarasov S (2024) Computable species descriptions and nanopublications: applying ontology-based technologies to dung beetles (Coleoptera, Scarabaeinae). Biodiversity Data Journal 12: e121562. https://doi.org/10.3897/BDJ.12.e121562