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  <front>
    <journal-meta>
      <journal-id journal-id-type="publisher-id">1</journal-id>
      <journal-id journal-id-type="index">urn:lsid:arphahub.com:pub:F9B2E808-C883-5F47-B276-6D62129E4FF4</journal-id>
      <journal-id journal-id-type="aggregator">urn:lsid:zoobank.org:pub:245B00E9-BFE5-4B4F-B76E-15C30BA74C02</journal-id>
      <journal-title-group>
        <journal-title xml:lang="en">Biodiversity Data Journal</journal-title>
        <abbrev-journal-title xml:lang="en">BDJ</abbrev-journal-title>
      </journal-title-group>
      <issn pub-type="ppub">1314-2836</issn>
      <issn pub-type="epub">1314-2828</issn>
      <publisher>
        <publisher-name>Pensoft Publishers</publisher-name>
      </publisher>
    </journal-meta>
    <article-meta>
      <article-id pub-id-type="doi">10.3897/BDJ.14.e214835</article-id>
      <article-id pub-id-type="publisher-id">214835</article-id>
      <article-id pub-id-type="manuscript">32212</article-id>
      <article-categories>
        <subj-group subj-group-type="heading">
          <subject>Taxonomy &amp; Inventories</subject>
        </subj-group>
        <subj-group subj-group-type="Taxon classification">
          <subject>Agelenidae</subject>
        </subj-group>
        <subj-group subj-group-type="Subject classification">
          <subject>Taxonomy</subject>
        </subj-group>
        <subj-group subj-group-type="Geographical classification">
          <subject>China</subject>
        </subj-group>
      </article-categories>
      <title-group>
        <article-title>First description of the male of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Troglocoelotes">Troglocoelotes</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="doul">doul</tp:taxon-name-part></tp:taxon-name></italic> Lin &amp; Li, 2024 (<tp:taxon-name><tp:taxon-name-part taxon-name-part-type="order" reg="Araneae">Araneae</tp:taxon-name-part></tp:taxon-name>, <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="family" reg="Agelenidae">Agelenidae</tp:taxon-name-part></tp:taxon-name>, <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="subfamily" reg="Coelotinae">Coelotinae</tp:taxon-name-part></tp:taxon-name>)</article-title>
      </title-group>
      <contrib-group content-type="authors">
        <contrib contrib-type="author" corresp="no">
          <name name-style="western">
            <surname>Wang</surname>
            <given-names>Junshuo</given-names>
          </name>
          <xref ref-type="aff" rid="A1">1</xref>
        </contrib>
        <contrib contrib-type="author" corresp="no">
          <name name-style="western">
            <surname>Wei</surname>
            <given-names>Mian</given-names>
          </name>
          <uri content-type="orcid">https://orcid.org/0000-0001-7348-8885</uri>
          <xref ref-type="aff" rid="A2">2</xref>
        </contrib>
        <contrib contrib-type="author" corresp="no">
          <name name-style="western">
            <surname>Zhang</surname>
            <given-names>He</given-names>
          </name>
          <xref ref-type="aff" rid="A3">3</xref>
        </contrib>
        <contrib contrib-type="author" corresp="no">
          <name name-style="western">
            <surname>Liu</surname>
            <given-names>Jie</given-names>
          </name>
          <xref ref-type="aff" rid="A1">1</xref>
          <xref ref-type="aff" rid="A4">4</xref>
        </contrib>
        <contrib contrib-type="author" corresp="yes">
          <name name-style="western">
            <surname>Hu</surname>
            <given-names>Changhao</given-names>
          </name>
          <email xlink:type="simple">changhaohu1998@gmail.com</email>
          <uri content-type="orcid">https://orcid.org/0009-0007-5591-3121</uri>
          <xref ref-type="aff" rid="A1">1</xref>
          <xref ref-type="aff" rid="A4">4</xref>
          <xref ref-type="aff" rid="A5">5</xref>
        </contrib>
      </contrib-group>
      <aff id="A1">
        <label>1</label>
        <addr-line content-type="verbatim">Arachnid Resource Centre of Hubei Province &amp; Hubei Key Laboratory of Regional Development and Environmental Response, Faculty of Resources and Environmental Science, Hubei University, Wuhan, China</addr-line>
        <institution>Arachnid Resource Centre of Hubei Province &amp; Hubei Key Laboratory of Regional Development and Environmental Response, Faculty of Resources and Environmental Science, Hubei University</institution>
        <addr-line content-type="city">Wuhan</addr-line>
        <country>China</country>
        <uri content-type="ror">https://ror.org/03a60m280</uri>
      </aff>
      <aff id="A2">
        <label>2</label>
        <addr-line content-type="verbatim">College of Life Sciences, Sichuan University, Chengdu, China</addr-line>
        <institution>College of Life Sciences, Sichuan University</institution>
        <addr-line content-type="city">Chengdu</addr-line>
        <country>China</country>
        <uri content-type="ror">https://ror.org/011ashp19</uri>
      </aff>
      <aff id="A3">
        <label>3</label>
        <addr-line content-type="verbatim">College of Physics and Electronic Engineering, Xingtai University, Xingtai, China</addr-line>
        <institution>College of Physics and Electronic Engineering, Xingtai University</institution>
        <addr-line content-type="city">Xingtai</addr-line>
        <country>China</country>
        <uri content-type="ror">https://ror.org/05c1r5z64</uri>
      </aff>
      <aff id="A4">
        <label>4</label>
        <addr-line content-type="verbatim">Centre for Behavioral Ecology and Evolution, School of Life Sciences, Hubei University, Wuhan, China</addr-line>
        <institution>Centre for Behavioral Ecology and Evolution, School of Life Sciences, Hubei University</institution>
        <addr-line content-type="city">Wuhan</addr-line>
        <country>China</country>
        <uri content-type="ror">https://ror.org/03a60m280</uri>
      </aff>
      <aff id="A5">
        <label>5</label>
        <addr-line content-type="verbatim">College of Life Sciences, Hunan Normal University, Changsha, China</addr-line>
        <institution>College of Life Sciences, Hunan Normal University</institution>
        <addr-line content-type="city">Changsha</addr-line>
        <country>China</country>
        <uri content-type="ror">https://ror.org/053w1zy07</uri>
      </aff>
      <author-notes>
        <fn fn-type="corresp">
          <p>Corresponding author: Changhao Hu (<email xlink:type="simple">changhaohu1998@gmail.com</email>).</p>
        </fn>
        <fn fn-type="edited-by">
          <p>Academic editor: Dragomir Dimitrov</p>
        </fn>
      </author-notes>
      <pub-date pub-type="collection">
        <year>2026</year>
      </pub-date>
      <pub-date pub-type="epub">
        <day>02</day>
        <month>10</month>
        <year>2026</year>
      </pub-date>
      <volume>14</volume>
      <elocation-id>e214835</elocation-id>
      <uri content-type="arpha" xlink:href="http://openbiodiv.net/84668B50-96FC-518C-9D12-0C953F3B6038">84668B50-96FC-518C-9D12-0C953F3B6038</uri>
      <history>
        <date date-type="received">
          <day>08</day>
          <month>09</month>
          <year>2026</year>
        </date>
        <date date-type="accepted">
          <day>25</day>
          <month>09</month>
          <year>2026</year>
        </date>
      </history>
      <permissions>
        <copyright-statement>Junshuo Wang, Mian Wei, He Zhang, Jie Liu, Changhao Hu</copyright-statement>
        <license license-type="creative-commons-attribution" xlink:href="http://creativecommons.org/licenses/by/4.0/" xlink:type="simple">
          <license-p>This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.</license-p>
        </license>
      </permissions>
      <abstract>
        <label>Abstract</label>
        <sec sec-type="background">
          <title>Background</title>
          <p><italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Troglocoelotes">Troglocoelotes</tp:taxon-name-part></tp:taxon-name></italic> Zhao &amp; Li, 2019 is currently known from Guangxi and Guizhou, China and comprises 12 species. Of these, only four are known from both sexes, whereas the remaining eight are known only from females. <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Troglocoelotes">Troglocoelotes</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="doul">doul</tp:taxon-name-part></tp:taxon-name></italic> Lin &amp; Li, 2024 was originally described from a female specimen collected in Guangxi and its male has remained unknown.</p>
        </sec>
        <sec sec-type="new information">
          <title>New information</title>
          <p>Based on specimens collected from the type locality of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Troglocoelotes">Troglocoelotes</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="doul">doul</tp:taxon-name-part></tp:taxon-name></italic> Lin &amp; Li, 2024, the male of this species is described for the first time.</p>
        </sec>
      </abstract>
      <kwd-group>
        <label>Keywords</label>
        <kwd>cave</kwd>
        <kwd>morphology</kwd>
        <kwd>spider</kwd>
        <kwd>taxonomy</kwd>
        <kwd>troglobite</kwd>
      </kwd-group>
      <counts>
        <fig-count count="5"/>
        <table-count count="0"/>
        <ref-count count="7"/>
      </counts>
    </article-meta>
  </front>
  <body>
    <sec sec-type="Introduction">
      <title>Introduction</title>
      <p><italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Troglocoelotes">Troglocoelotes</tp:taxon-name-part></tp:taxon-name></italic> Zhao &amp; Li, 2019 is currently the only genus of <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="subfamily" reg="Coelotinae">Coelotinae</tp:taxon-name-part></tp:taxon-name> F. O. Pickard-Cambridge, 1893 composed entirely of troglobitic species (<xref ref-type="bibr" rid="B14451343">Zhou et al. 2023</xref>). Members of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Troglocoelotes">Troglocoelotes</tp:taxon-name-part></tp:taxon-name></italic> exhibit typical cave-associated morphological characteristics, including reduction or loss of pigmentation and regression or loss of eyes (<xref ref-type="bibr" rid="B14451331">Mammola and Isaia 2017</xref>, <xref ref-type="bibr" rid="B14451290">Li et al. 2019</xref>). They inhabit the deep zones of limestone caves in Guangxi and Guizhou, China, where they build webs in rock crevices, shallow depressions on the cave floor and beneath loose stones (personal observation). To date, 12 extant species have been described in the genus, eight of which are known only from females (<xref ref-type="bibr" rid="B14451352">World Spider Catalog 2026</xref>).</p>
      <p>Shenlonggong Cave is located in Huanjiang, Guangxi, an area characterised by extensive karst landscapes and numerous limestone caves. The cave is the type locality of several troglobitic species, including <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Coecobrya">Coecobrya</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="draconis">draconis</tp:taxon-name-part></tp:taxon-name></italic> Zhang &amp; Dong, 2014 (<tp:taxon-name><tp:taxon-name-part taxon-name-part-type="class" reg="Collembola">Collembola</tp:taxon-name-part></tp:taxon-name> Lubbock, 1871), <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Desmoxytes">Desmoxytes</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="scolopendroides">scolopendroides</tp:taxon-name-part></tp:taxon-name></italic> Golovatch, Geoffroy &amp; Mauriès, 2010 (<tp:taxon-name><tp:taxon-name-part taxon-name-part-type="class" reg="Diplopoda">Diplopoda</tp:taxon-name-part></tp:taxon-name> Blainville in Gervais, 1844), <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Troglocoelotes">T.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="doul">doul</tp:taxon-name-part></tp:taxon-name></italic> Lin &amp; Li, 2024 (<tp:taxon-name><tp:taxon-name-part taxon-name-part-type="order" reg="Araneae">Araneae</tp:taxon-name-part></tp:taxon-name> Clerck, 1757), amongst others (<xref ref-type="bibr" rid="B14451281">Golovatch et al. 2010</xref>, <xref ref-type="bibr" rid="B14451322">Zhang and Dong 2014</xref>, <xref ref-type="bibr" rid="B14451299">Lin et al. 2024</xref>).</p>
      <p><italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Troglocoelotes">Troglocoelotes</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="doul">doul</tp:taxon-name-part></tp:taxon-name></italic> was originally described, based on a female specimen and its male has remained unknown (<xref ref-type="bibr" rid="B14451299">Lin et al. 2024</xref>). During the examination of recently collected spider specimens from the cave, one female was identified as <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Troglocoelotes">T.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="doul">doul</tp:taxon-name-part></tp:taxon-name></italic> and two males were considered conspecific, based on their co-occurrence and morphological similarities, particularly their similarly degenerated eyes. The male of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Troglocoelotes">T.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="doul">doul</tp:taxon-name-part></tp:taxon-name></italic> is, therefore, described herein for the first time.</p>
    </sec>
    <sec sec-type="Materials and methods">
      <title>Materials and methods</title>
      <p>The specimens examined in this study are deposited in the Centre for Behavioral Ecology and Evolution (CBEE), School of Life Sciences, Hubei University in Wuhan, China (curator: Jie Liu). Specimens were examined using an OLYMPUS SZX7 stereomicroscope. The male palps were examined and photographed after dissection. The epigyne was dissected from the body, immersed in 0.1 mg/ml Protease K solution, heated in a water bath at 56℃ and stained with Amido black 10B. All measurements were taken using a Leica M205 C stereomicroscope. Eye diameters were taken at the widest point. Leg measurements are given as the total length of leg (femur, patella, tibia, metatarsus, tarsus). All measurements are in millimetres (mm). Photographs were taken using Leica M205 C stereomicroscope and OLYMPUS BX51 microscope and multifocal images were produced using Helicon Focus (Version 8.1.0).</p>
      <p>Abbreviations: ALE–anterior lateral eye; AME–anterior median eye; PLE–posterior lateral eye; PME–posterior median eye; I, II, III, IV–legs I to IV.</p>
    </sec>
    <sec sec-type="Taxon treatments">
      <title>Taxon treatments</title>
      <tp:taxon-treatment>
        <tp:nomenclature>
          <tp:taxon-name><object-id content-type="arpha">5ADAF947-46FA-53B8-8790-98F8856B7E12</object-id><tp:taxon-name-part taxon-name-part-type="genus">Troglocoelotes</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">doul</tp:taxon-name-part></tp:taxon-name>
          <tp:taxon-authority>Lin &amp; Li, 2024</tp:taxon-authority>
          <tp:nomenclature-citation-list>
            <tp:nomenclature-citation>
              <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Troglocoelotes">Troglocoelotes</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="doul">doul</tp:taxon-name-part></tp:taxon-name>
              <comment> Lin &amp; Li, in <xref ref-type="bibr" rid="B14451299">Lin et al. (2024)</xref>: 7, figs. 1A, B, 3A and 4.</comment>
            </tp:nomenclature-citation>
          </tp:nomenclature-citation-list>
        </tp:nomenclature>
        <tp:treatment-sec sec-type="materials">
          <title>Materials</title>
          <list list-type="alpha-lower" list-content="occurrences">
            <list-item>
              <p><bold>Type status:</bold> <named-content content-type="dwc:typeStatus" xlink:type="simple">Holotype</named-content>. <bold>Occurrence:</bold> recordedBy: <named-content content-type="dwc:recordedBy" xlink:type="simple">Haolin Mo and Shanmi Zheng</named-content>; individualCount: <named-content content-type="dwc:individualCount" xlink:type="simple">1</named-content>; sex: <named-content content-type="dwc:sex" xlink:type="simple">female</named-content>; lifeStage: <named-content content-type="dwc:lifeStage" xlink:type="simple">adult</named-content>; <bold>Location:</bold> country: <named-content content-type="dwc:country" xlink:type="simple">China</named-content>; stateProvince: <named-content content-type="dwc:stateProvince" xlink:type="simple">Guangxi Zhuang Autonomous Region</named-content>; county: <named-content content-type="dwc:county" xlink:type="simple">Hechi City, Huanjiang Maonan Autonomous County</named-content>; locality: <named-content content-type="dwc:locality" xlink:type="simple">Dacai Township, Shenlonggong Cave</named-content>; verbatimLatitude: <named-content content-type="dwc:verbatimLatitude" xlink:type="simple">108.3898°E</named-content>; verbatimLongitude: <named-content content-type="dwc:verbatimLongitude" xlink:type="simple">24.7707°N</named-content>; <bold>Identification:</bold> identificationRemarks: <named-content content-type="dwc:identificationRemarks" xlink:type="simple">not examined</named-content>; <bold>Event:</bold> year: <named-content content-type="dwc:year" xlink:type="simple">2023</named-content>; month: <named-content content-type="dwc:month" xlink:type="simple">8</named-content>; day: <named-content content-type="dwc:day" xlink:type="simple">1</named-content>; <bold>Record Level:</bold> institutionCode: <named-content content-type="dwc:institutionCode" xlink:type="simple">IZCAS-Ar44790</named-content></p>
            </list-item>
            <list-item>
              <p><bold>Type status:</bold> <named-content content-type="dwc:typeStatus" xlink:type="simple">Other material</named-content>. <bold>Occurrence:</bold> recordedBy: <named-content content-type="dwc:recordedBy" xlink:type="simple">Changhao Hu, Jian Chang, Jinhui Wang, and Mingwei Li</named-content>; individualCount: <named-content content-type="dwc:individualCount" xlink:type="simple">3</named-content>; sex: <named-content content-type="dwc:sex" xlink:type="simple">2 males, 1 female</named-content>; lifeStage: <named-content content-type="dwc:lifeStage" xlink:type="simple">adult</named-content>; <bold>Location:</bold> country: <named-content content-type="dwc:country" xlink:type="simple">China</named-content>; stateProvince: <named-content content-type="dwc:stateProvince" xlink:type="simple">Guangxi Zhuang Autonomous Region</named-content>; county: <named-content content-type="dwc:county" xlink:type="simple">Hechi City, Huanjiang Maonan Autonomous County</named-content>; locality: <named-content content-type="dwc:locality" xlink:type="simple">Dacai Township, Shenlonggong Cave</named-content>; verbatimElevation: <named-content content-type="dwc:verbatimElevation" xlink:type="simple">368 m</named-content>; verbatimLatitude: <named-content content-type="dwc:verbatimLatitude" xlink:type="simple">24.7707°N</named-content>; verbatimLongitude: <named-content content-type="dwc:verbatimLongitude" xlink:type="simple">108.3898°E</named-content>; <bold>Event:</bold> year: <named-content content-type="dwc:year" xlink:type="simple">2026</named-content>; month: <named-content content-type="dwc:month" xlink:type="simple">8</named-content>; day: <named-content content-type="dwc:day" xlink:type="simple">10</named-content></p>
            </list-item>
          </list>
        </tp:treatment-sec>
        <tp:treatment-sec sec-type="Description">
          <title>Description</title>
          <p><bold>Male</bold>: Total length 7.19, carapace 4.19 long, 2.74 wide, opisthosoma 3.12 long, 2.05 wide. Eyes vestigial (Fig. <xref ref-type="fig" rid="F14451377">1</xref>A). Eye sizes and interdistances: AME absent, ALE 0.07, PME 0.06, PLE 0.09, PME–PME 0.16, PME–PLE 0.19, ALE–PME 0.16, ALE–PLE 0.07. Leg measurements: I 17.25 (4.30, 1.06, 4.53, 4.37, 2.99), II 15.98 (4.23, 1.03, 3.94, 4.10, 2.68), III 15.30 (3.96, 0.92, 3.58, 4.24, 2.60), IV 19.94 (4.85, 1.12, 4.80, 6.15, 3.02). Leg formula IV-I-II-III. Chelicerae with three promarginal and two retromarginal teeth (Fig. <xref ref-type="fig" rid="F14451377">1</xref>C). Opisthosoma and legs with long hairs (Fig. <xref ref-type="fig" rid="F14451379">2</xref>A–C).</p>
          <p>Colouration (Fig. <xref ref-type="fig" rid="F14451379">2</xref>A–C). Carapace yellowish-brown, head region light brown, thoracic region pale yellow, fovea black. Chelicerae dark brown. Endites and labium brown, with white distal parts. Sternum yellowish-brown. Legs light brown. Opisthosoma and spinnerets grey.</p>
          <p>Palp (Fig. <xref ref-type="fig" rid="F14451381">3</xref>). Patellar apophysis digitiform. Retrolateral tibial apophysis trapeziform, almost 2/3 the length of tibia, with its distal end slightly extending beyond the tibia. Lateral tibial apophysis short and rounded. Cymbium slender, more than twice as long as bulb; cymbial furrow almost 1/3 the length of cymbium. Median apophysis spoon-shaped, almost half the length of bulb. Conductor strong and sclerotised, curved ventrally, with weak basal lamellar and blunt end; dorsal apophysis of conductor triangular, distal tip with three denticles. Embolus filform and curved, arising from embolic base at 8:30 o’clock position.</p>
          <p><bold>Female</bold>: See <xref ref-type="bibr" rid="B14451299">Lin et al. (2024)</xref> (Fig. <xref ref-type="fig" rid="F14451377">1</xref>B, D, Fig. <xref ref-type="fig" rid="F14451379">2</xref>D–F and Fig. <xref ref-type="fig" rid="F14451383">4</xref>).</p>
        </tp:treatment-sec>
        <tp:treatment-sec sec-type="Diagnosis">
          <title>Diagnosis</title>
          <p>The male of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Troglocoelotes">T.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="doul">doul</tp:taxon-name-part></tp:taxon-name></italic> can be distinguished from those of all other male-known congeners by the following combination of characters: 1. cymbium more than twice as long as bulb (less than twice as long as bulb in all other male-known congeners); 2. patella apophysis digitiform (pointed in <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Troglocoelotes">T.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="tortus">tortus</tp:taxon-name-part></tp:taxon-name></italic>); 3. median apophysis large, almost half the length of bulb (almost 1/3 the length of bulb in all other male-known congeners); 4. conductor curved, with blunt distal end (straight and pointed in all other male-known congeners); 5. dorsal apophysis of conductor triangular and pointed (rounded and blunt in <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Troglocoelotes">T.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="qixianensis">qixianensis</tp:taxon-name-part></tp:taxon-name></italic> and <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Troglocoelotes">T.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="tortus">tortus</tp:taxon-name-part></tp:taxon-name></italic>, absent in <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Troglocoelotes">T.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="liangensis">liangensis</tp:taxon-name-part></tp:taxon-name></italic> and <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Troglocoelotes">T.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="yumiganensis">yumiganensis</tp:taxon-name-part></tp:taxon-name></italic>); 6. embolus arising from embolic base at 8:30 o’clock position (6 o’clock position in all other male-known congeners) (cf. Fig. <xref ref-type="fig" rid="F14451381">3</xref> and figs. 5, 8, 10 and 12 in <xref ref-type="bibr" rid="B14451290">Li et al. (2019)</xref>).</p>
          <p>The female of this species is similar to those of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Troglocoelotes">T.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="sinanensis">sinanensis</tp:taxon-name-part></tp:taxon-name></italic> Zhou &amp; Zhao, 2023 in having wide copulatory opening, the posterior margin of atrium almost as wide as atrium and the epigynal teeth situated laterally on atrium (cf. Fig. <xref ref-type="fig" rid="F14451383">4</xref> and figs. 3A–C in <xref ref-type="bibr" rid="B14451343">Zhou et al. (2023)</xref>), but can be distinguished by: 1. lateral part of copulatory duct with a projection (absent in <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Troglocoelotes">T.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="sinanensis">sinanensis</tp:taxon-name-part></tp:taxon-name></italic>); and 2. spermatheca spherical, narrower than copulatory duct (oval and laterally elongated, wider than copulatory duct in <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Troglocoelotes">T.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="sinanensis">sinanensis</tp:taxon-name-part></tp:taxon-name></italic>).</p>
        </tp:treatment-sec>
        <tp:treatment-sec sec-type="Distribution">
          <title>Distribution</title>
          <p>Known only from type locality (Shenlonggong Cave in Guangxi, China) (Fig. <xref ref-type="fig" rid="F14451385">5</xref>).</p>
        </tp:treatment-sec>
      </tp:taxon-treatment>
    </sec>
  </body>
  <back>
    <ack>
      <title>Acknowledgements</title>
      <p>We are very grateful to Dr. Jinhui Wang, Mr. Jian Chang and Mr. Mingwei Li (CBEE) for their assistance with fieldwork. This manuscript benefitted greatly from the comments of the subject editor Dr. Dragomir Dimitrov (National Museum of Natural History, Bulgaria) and the reviewer. This research was supported by the National Science and Technology Fundamental Resources Investigation Program of China (Grant No. 2023FY100200), the National Natural Sciences Foundation of China (NSFC-32570541/32270495/32400360/32300378), the Hebei Natural Science Foundation (C2025108010) and the Science Research Project of Hebei Education Department (BJ2026157).</p>
    </ack>
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  </back>
  <floats-group>
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      <label>Figure 1.</label>
      <caption>
        <p><italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Troglocoelotes">Troglocoelotes</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="doul">doul</tp:taxon-name-part></tp:taxon-name></italic> Lin &amp; Li, 2024. <bold>A, B</bold> eye areas, frontal view; <bold>C, D</bold> chelicerae, retrolateral view (red lines: promarginal teeth; blue lines: retromarginal teeth); <bold>A, C</bold> male; <bold>B, D</bold> female. Scale bars: 0.5 mm.</p>
      </caption>
      <graphic xlink:href="bdj-14-e214835-g001.png" position="float" id="oo_1765653.png" orientation="portrait" xlink:type="simple">
        <uri content-type="original_file">https://binary.pensoft.net/fig/1765653</uri>
      </graphic>
    </fig>
    <fig id="F14451379" position="float" orientation="portrait">
      <object-id content-type="arpha">1B974D1F-D283-5A5A-8D9B-40EB11D4F3D5</object-id>
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      <label>Figure 2.</label>
      <caption>
        <p>Habitus of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Troglocoelotes">Troglocoelotes</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="doul">doul</tp:taxon-name-part></tp:taxon-name></italic> Lin &amp; Li, 2024. <bold>A–C</bold> male; <bold>D–F</bold> female; <bold>A, D</bold> dorsal view; <bold>B, E</bold> ventral view; <bold>C, F</bold> lateral view. Scale bars: 2 mm.</p>
      </caption>
      <graphic xlink:href="bdj-14-e214835-g002.png" position="float" id="oo_1765655.png" orientation="portrait" xlink:type="simple">
        <uri content-type="original_file">https://binary.pensoft.net/fig/1765655</uri>
      </graphic>
    </fig>
    <fig id="F14451381" position="float" orientation="portrait">
      <object-id content-type="arpha">EE9A628E-875F-5347-81F4-5A746D03DFD4</object-id>
      <object-id content-type="doi">10.3897/BDJ.14.e214835.figure1</object-id>
      <label>Figure 3.</label>
      <caption>
        <p>Male palp of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Troglocoelotes">Troglocoelotes</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="doul">doul</tp:taxon-name-part></tp:taxon-name></italic> Lin &amp; Li, 2024. <bold>A–C</bold> left palp (<bold>A</bold> prolateral view; <bold>B</bold> ventral view; <bold>C</bold> retrolateral view, inset showing the denticles on the distal tip of CDA); <bold>D–F</bold> right bulb, horizontally flipped (<bold>D</bold> apical view; <bold>E</bold> dorsal view; <bold>F</bold> retrolateral view). Abbreviations: C: conductor; CDA: dorsal apophysis of conductor; CF: cymbial furrow; E: embolus; EB: embolic base; LTA: lateral tibial apophysis; MA: median apophysis; PA: patellar apophysis; RTA: retrolateral tibial apophysis; ST: subtegulum; T: tegulum; TS: tegular sclerite. Scale bars: 0.5 mm (A–C), 0.1 mm (D–F).</p>
      </caption>
      <graphic xlink:href="bdj-14-e214835-g003.png" position="float" id="oo_1765656.png" orientation="portrait" xlink:type="simple">
        <uri content-type="original_file">https://binary.pensoft.net/fig/1765656</uri>
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    </fig>
    <fig id="F14451383" position="float" orientation="portrait">
      <object-id content-type="arpha">83322071-069D-5EB9-B231-B52835D5B45F</object-id>
      <object-id content-type="doi">10.3897/BDJ.14.e214835.figure5</object-id>
      <label>Figure 4.</label>
      <caption>
        <p>Female genitalia of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Troglocoelotes">Troglocoelotes</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="doul">doul</tp:taxon-name-part></tp:taxon-name></italic> Lin &amp; Li, 2024. <bold>A, B</bold> epigyne, ventral view (<bold>A</bold> uncleaned; <bold>B</bold> cleaned); <bold>C, D</bold> vulva, dorsal view (<bold>C</bold> unstained; <bold>D</bold> stained); <bold>E, F</bold> vulva, anterodorsal view (<bold>E</bold> unstained; <bold>F</bold> stained); <bold>G</bold> epigyne, anteroventral view. Abbreviations: A: atrium; CD: copulatory duct; CO: copulatory opening; ET: epigynal tooth; FD: fertilisation duct; S: spermatheca. Scale bars: 0.2 mm.</p>
      </caption>
      <graphic xlink:href="bdj-14-e214835-g004.png" position="float" id="oo_1765657.png" orientation="portrait" xlink:type="simple">
        <uri content-type="original_file">https://binary.pensoft.net/fig/1765657</uri>
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    </fig>
    <fig id="F14451385" position="float" orientation="portrait">
      <object-id content-type="arpha">6ECE1DAC-5921-522E-8158-C3F5A5C62E77</object-id>
      <object-id content-type="doi">10.3897/BDJ.14.e214835.figure1</object-id>
      <label>Figure 5.</label>
      <caption>
        <p>Distribution map of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Troglocoelotes">Troglocoelotes</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="doul">doul</tp:taxon-name-part></tp:taxon-name></italic> Lin &amp; Li, 2024.</p>
      </caption>
      <graphic xlink:href="bdj-14-e214835-g005.png" position="float" id="oo_1765658.png" orientation="portrait" xlink:type="simple">
        <uri content-type="original_file">https://binary.pensoft.net/fig/1765658</uri>
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    </fig>
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