Corresponding authors: Recep Sulhi Özkütük (
Academic editor: Jason Bond
A new species of
The goal of this article is to describe a recently discovered species of
Specimens were collected from Antalya Province in the Mediterranean region of Turkey, using a sifter. The specimens were preserved in 70% ethanol and deposited in the Anadolu University Zoology Museum. Digital images of the copulatory organs were taken with a Leica DFC295 digital camera attached to a Leica S8AP0 stereomicroscope and 5-15 photographs were taken in different focal planes and combined using automontage software. SEM microphotographs were made from dried and sputter coated (by gold) organs by use of a Zeiss Ultra Plus SEM device (Anadolu University, Eskişehir). All measurements are in mm, with methods as per (
The following abbreviations are used in the text:
No apparent dimorphism between the sexes except in body sizes. Carapace hexagonal, reddish brown, dull and smooth. Carapace covered with very short, tiny and sparsely distributed setae. AME, PLE and PME closely grouped; AME separated (Fig.
Cheliceral groove with four teeth; at retromargin, with a small tooth located at the base of the groove, and with a more developed second tooth a little above the second quarter. Both retromarginal teeth conical and tubercular. Promarginal teeth more strongly developed; the one closer to the base of the cheliceral groove larger and almost twice the size of the other (Fig.
Legs yellowish brown, covered with tiny brownish hairs on all surface. Leg formula IV, I, II, III. Tarsi with three claws. Paired claws toothed. Paired claws of leg I and II with 7 teeth; leg III and IV with 4 teeth. Scopula weak and in ventral position on the first distal half on 3rd and 4th tarsi and on 4th metatarsus; relatively strongly developed on 3rd metatarsus in ventral position at the first distal half. Coxae III with 1 prolateral spine; coxae IV with 1-2 prolateral spines. Patellae III with 1 spine dorsally. Further details on leg spination are given in (Table
Bulbus almost oval, yellowish brown, with embolus and two apophyses. An apparent membranous part between the bulb and the distal appendages; embolus hook-shaped, black, almost homogenously sclerotized. Bent anteriorly following the same course as apophysisb. Apophysisa L-shaped, apically well sclerotized, short and strong. Apophysisb separated from the membranous part of the bulbus with a wide angle from the base and orientates anteriorly. Apically blunt, conical and in the shape of a triangular apophysis in retrolateral view (Fig.
Distal expansion of spermatheca tenuously developed between distal crest and rod-shaped part of the anterior spermatheca. Nevertheless, it is more sclerotized at the surface compared to other parts of the anterior spermatheca. Distal crest gradually tapers through the tip. Rod-shaped part of the anterior spermatheca almost one and a half times the length of the distal crest. Basal transverse part of the anterior spermatheca widens through laterals and narrows through the tips, in the shape of open wings. A ring-shaped strongly sclerotized structure is apparent at the juncture of the rod-shaped part of the anterior spermatheca and the basal transverse part of the anterior spermatheca. Transverse bar in the shape of a lip. Posterior diverticulum prominent, and in the form of a broad membranous sac (Fig.
The specific name refers to the type locality (Alanya District, Antalya Province, Turkey).
All specimens were collected from slopes and ridges lying parallel to the coastline, from sea level up to a maximum of around 1200 m on peaks of the Taurides Mountain range, by sifting tree litter of mixed forest with the following species
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We are deeply indebted to Dr. Fulvio Gasparo (Trieste, Italy) who discussed differences between males of the new species and
Habitus of holotype male
Carapace, male
Ditto, female
Ditto, lateral view
Cheliceral teeth (arrow indicates position of retromarginal small tooth)
Ditto
Distal appendages of male bulb, nearly posterior view. Abbreviations:
Male palp, prolateral view
Ditto, retrolateral view
Ditto, nearly retrolateral view
Vulva, dorsal view
Ditto
Ditto
Ditto, ventral view
Male palp, prolateral view
Ditto, retrolateral view
Vulva, dorsal view.
Leg measurements of
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1.48 / 1.88 | 0.83 / 1.14 | 1.18 / 1.55 | 1.00 / 1.35 | 0.40 / 0.45 | 4.89 / 6.37 |
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1.35 / 1.75 | 0.75 / 1.13 | 1.13 / 1.45 | 1.10 / 1.38 | 0.38 / 0.38 | 4.71 / 6.09 |
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1.07 / 1.45 | 0.57 / 0.73 | 0.83 / 0.78 | 1.10 / 1.38 | 0.34 / 0.48 | 3.91 / 4.82 |
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1.56 / 2.00 | 0.75 / 0.98 | 1.38 / 1.53 | 1.41 / 1.78 | 0.50 / 0.53 | 5.60 / 6.82 |
Leg spination of
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0 | 0 | 1 Pl | 1 Pl |
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2 Pl | 1, 1 Pl | 1, 1 D 1, 1, 1 Rl | 1, 2 D |
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0 | 0 | 1 D | 0 |
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0 | 0 | 1, 1 Pl 1, 1, 1 Rl 1, 1, 2 V | 1, 1, 1 Pl 1, 1, 1 Rl 1, 1, 2 V |
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0 | 0 | 1, 1 Pl 1, 1, 1 Rl 2, 1, 2 V | 1, 1, 1 Pl 1, 1, 1, 1 Rl 1, 1, 2 V |
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0 | 0 | 1 Pl | 1-2 Pl |
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2 Pl | 1, 1 Pl | 1, 1 D 1, 1 Rl | 1, 1 Pl 1, 1 D |
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0 | 0 | 1 D | 0 |
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0 | 0 | 1, 1 Pl 1, 1, 1 Rl 1, 1, 2 V | 1, 1, 1 Pl 1, 1, 1 Rl 1, 1, 2 V |
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0 | 0 | 1, 1 Pl 1, 1, 1 Rl 2, 1, 2 V | 1, 1, 1 Pl 1, 1, 1, 1 Rl 1, 1, 2 V |