Eutrophic hypnaceous swamp developed over limestone, with sluggishly-flowing waters (91 m a.s.l.) (northwest of Ayan Settlement).
	Part of: Bakalin V, Klimova K, Bakalin D, Choi SS (2021) Liverwort flora of Ayan – a gained link between subarctic and hemiboreal floras in West Okhotiya (Pacific Russia). Biodiversity Data Journal 9: e65199. https://doi.org/10.3897/BDJ.9.e65199