Five-year overview of seasons, photoperiod, rainfall and samples of the four Pterostichus species. Dashed lines indicate the following dates: 1 - initiation of sampling for the project; 2 - beginning of the period for which samples are considered for the mating marks study (i.e. the beginning of the drought recovery period); and 3 - the official, State of California end of the drought. Seasons are noted as wi= winter, sp= spring, su= summer, au= autumn.