Biodiversity Data Journal : Taxonomic Paper
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Taxonomic Paper
Marine Bryozoa of Greece: an annotated checklist
expand article infoVasilis Gerovasileiou, Antonietta Rosso§,|
‡ Institute of Marine Biology, Biotechnology and Aquaculture, Hellenic Centre for Marine Research, Heraklion, Greece
§ Sezione di Scienze della Terra, Dipartimento di Scienze Biologiche, Geologiche e Ambientali, Università di Catania, Catania, Italy
| Unità di Ricerca di Catania, Consorzio Interuniversitario per le Scienze del Mare (CoNISMa), Rome, Italy
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Abstract

Background

Until today, a complete checklist of Bryozoa of the Greek seas had never been published and species records were scattered in several taxonomic and ecological studies. The aim of this paper is to produce a first checklist of marine bryozoan species of Greece, in the framework of the Greek Taxon Information System (GTIS) initiative of the LifeWatchGreece Research Infrastructure (ESFRI), by reviewing the existing literature and following the recent trends in the taxonomy of this group. 

New information

The marine bryozoan fauna of Greece comprises 237 species, classified in 127 genera, 66 families, 3 orders, and 2 classes. The vast majority belongs to the class Gymnolaemata (177 Cheilostomatida and 21 Ctenostomatida), while the remaining 39 species are Stenolaemata (all Cyclostomatida). Among these species, 12 are considered endemic to the eastern Mediterranean, while another 12 species are non-indigenous.

Keywords

Biodiversity, Aegean Sea, Sea of Crete, Levantine Sea, Ionian Sea, Eastern Mediterranean.

Introduction

The Mediterranean bryozoan fauna represents about 9.6% of the known global diversity of Bryozoa (Rosso and Di Martino 2016). However, current knowledge on the bryozoan fauna of this basin is highly biased due to differences in research effort for different taxa, marine areas and habitat types. Recent efforts in the south-eastern Mediterranean Sea have significantly increased our knowledge from areas which were previously considered “data-poor” spots for bryozoans, revealing a considerable proportion of non-indigenous taxa (e.g. Harmelin et al. 2016, Sokolover et al. 2016). Furthermore, information on Bryozoa from several Mediterranean areas is fragmented in old publications and grey literature; the collation of updated checklists (e.g. Koçak and Aydın Önen 2014) by experts could contribute to the mapping of diversity for understudied taxa and areas (Bailly et al. 2016, this special collection).

Sporadic records of bryozoans from the Aegean Sea can be found in old taxonomic studies, including the descriptions of the cheilostome species Calpensia nobilis, Hippaliosina depressa and Watersipora cucullata from this marine region (Busk 1854), based on material collected from shallow water macroalgae by E. Forbes. The first studies focusing on Bryozoa of the eastern Mediterranean basin, the Greek seas included, took place in the late 1960s (Harmelin 1969), based on material that was collected with dredges during the oceanographic expedition of the French research vessel “Jean Charcot” (1968). A total of 101 species belonging to all the 3 living bryozoan orders, were found, including several new records for the Mediterranean fauna. Later, the taxonomic study by Hayward (1974) on the cheilostome Bryozoa of Chios Island (north-eastern Aegean) yielded 101 species, including rare records and new species. Sharing only 47 cheilostomes with Harmelin’s list mentioned above, this study increased the eastern Mediterranean bryozoan diversity by 54 species.

More recently, few taxonomic and ecological studies have focused on the bryozoan fauna of Greece (e.g. Castritsi-Catharios and Marcopoulou-Diacantoni 1983, Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios and Kiortis 1985, Castritsi-Catharios et al. 1985a, Castritsi-Catharios et al. 1985b, Castritsi-Catharios et al. 1986a, Castritsi-Catharios et al. 1986b, Castritsi-Catharios and Ganias 1989, Ganias 1990), while a considerable number of records has been included in general ecological studies examining a variety of habitats (e.g. Simboura et al. 1995, Conides et al. 1999, Morri et al. 1999, Antoniadou and Chintiroglou 2005, Gerovasileiou et al. 2013, Gerovasileiou et al. 2015). A complete checklist of Bryozoa however had never been published from Greece. The aim of the present study is to give a first annotated checklist of Bryozoa from the Greek seas.

Materials and methods

The checklist presented in this paper was compiled in the framework of the Greek Taxon Information System (GTIS), an initiative of the LifeWatchGreece Research e-Infrastructure aiming to produce a complete inventory of the known biodiversity of Greece (Bailly et al. 2016, this special collection). As this is the first attempt to compile a checklist of bryozoan diversity of the Greek seas, we performed an exhaustive literature review of bryozoan records. All taxa were cross-checked for synonymies and dubious records against the World List of Bryozoa (recent and fossil), the World Register of Marine Species (Bock and Gordon 2016, WoRMS Editorial Board 2016), and the recent update of the Mediterranean bryozoan diversity by Rosso and Di Martino (2016). Taxa identified as cf., as well as those left in open nomenclature, and fossil records were not included in the list. The checklist is annotated with the literature references reporting the presence of each species from Greek waters, as well as with notes on the nomenclature of some taxa. Non-indigenous bryozoans are also marked (NIB). The checklist follows the alphabetical order.

Checklist of Bryozoa known to occur in the Greek seas

Class Gymnolaemata Allman, 1856

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Order Cheilostomatida Busk, 1852

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Family Adeonidae Busk, 1884

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Genus Adeonella Busk, 1884

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Adeonella calveti (Canu & Bassler, 1930)

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Notes: 

Gautier 1962

Adeonella pallasii (Heller, 1867)

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Notes: 

Castritsi-Catharios et al. 1985a, Castritsi-Catharios et al. 1985b, Castritsi-Catharios et al. 1986a, Ganias 1990, Rosso and Novosel 2010, Chimenz Gusso et al. 2014, Gerovasileiou et al. 2015

Genus Reptadeonella Busk, 1884

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Reptadeonella violacea (Johnston, 1847)

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Notes: 

Harmelin 1969, Hayward 1974, Castritsi-Catharios and Marcopoulou-Diacantoni 1983, Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios et al. 1986a (as Reptoporellina, probably a mispelling), Ganias 1990, Morri et al. 1999, Antoniadou and Chintiroglou 2005

Family Aeteidae Smitt, 1868

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Genus Aetea Lamouroux, 1812

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Aetea anguina (Linnaeus, 1758)

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Notes: 

Harmelin 1969, Hayward 1974, Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios and Kiortis 1985, Ganias 1990

Aetea lepadiformis Waters, 1906

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Notes: 

Ganias 1990, Morri et al. 1999

Aetea sica (Couch, 1844)

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Notes: 

Harmelin 1969, Hayward 1974, Castritsi-Catharios and Marcopoulou-Diacantoni 1983, Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios and Kiortis 1985, Castritsi-Catharios et al. 1985a, Castritsi-Catharios et al. 1986a, Castritsi-Catharios et al. 1986b, Castritsi-Catharios and Ganias 1989, Ganias 1990, Morri et al. 1999

Aetea truncata (Landsborough, 1852)

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Notes: 

Harmelin 1969, Hayward 1974, Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios and Kiortis 1985, Castritsi-Catharios et al. 1985a, Castritsi-Catharios et al. 1986a, Castritsi-Catharios and Ganias 1989, Ganias 1990, Morri et al. 1999

Family Antroporidae Vigneaux, 1949

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Genus Rosseliana Jullien, 1888

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Rosseliana rosselii (Audouin, 1826)

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Notes: 

Harmelin 1969, Ganias 1990

Family Beaniidae Canu & Bassler, 1927

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Genus Beania Johnston, 1840

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Beania cylindrica (Hincks, 1886)

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Notes: 

Harmelin 1969, Ganias 1990, Morri et al. 1999

Beania hirtissima (Heller, 1867)

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Notes: 

Harmelin 1969, Hayward 1974, Castritsi-Catharios and Marcopoulou-Diacantoni 1983, Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios et al. 1986a, Castritsi-Catharios and Ganias 1989, Ganias 1990, Simboura et al. 1995, Morri et al. 1999

Beania magellanica (Busk, 1852)

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Notes: 

Harmelin 1969, Hayward 1974, Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios and Ganias 1989, Ganias 1990

Beania mirabilis Johnston, 1840

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Notes: 

Hayward 1974, Castritsi-Catharios and Ganias 1989, Ganias 1990

Family Bitectiporidae MacGillivray, 1895

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Genus Metroperiella Canu & Bassler, 1917

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Metroperiella lepralioides (Calvet, 1903)

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Notes: 

Harmelin 1969, Hayward 1974, Ganias 1990, Morri et al. 1999

Genus Pentapora Fischer, 1807

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Pentapora fascialis (Pallas, 1766)

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Notes: 

Ganias 1990, Antoniadou and Chintiroglou 2005, Chimenz Gusso et al. 2014

Genus Schizomavella Canu & Bassler, 1917

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Schizomavella rudis (Manzoni, 1869)

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Notes: 

In the absence of type material, the validity of S. rudis has been questioned by Reverter-Gil et al. (2012). Two new species, Stephanotheca watersi Reverter-Gil, Souto & Fernandez-Pulpeiro, 2012 and Schizomavella adriatica Reverter-Gil, Souto, Novosel & Tilbrook, 2016 have been erected to allocate some records from the western Basin and the Adriatic, respectively. Nevertheless, records and material from the Aegean were not considered and hence they are provisionally still reported as S. rudis (see also Rosso and Di Martino 2016).

Schizomavella (Calvetomavella) discoidea (Busk, 1859)

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Notes: 

Harmelin 1969, Hayward 1974, Morri et al. 1999; the status of these records needs to be checked (see Rosso and Di Martino 2016​).

Schizomavella (Schizomavella) asymetrica (Calvet, 1927)

Nomenclature: 

Synonym valid for Calyptotheca triarmata Hayward, 1974 (see Rosso et al. 2010​).

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Notes: 

Hayward 1974

Schizomavella (Schizomavella) cornuta (Heller, 1867)

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Notes: 

Harmelin 1969, Hayward 1974, Castritsi-Catharios and Ganias 1989, Ganias 1990, Simboura et al. 1995, Morri et al. 1999; Mediterranean colonies recorded as S. auriculata, S. auriculata cuspidata and S. cuspidata possibly belong to this species.

Schizomavella (Schizomavella) hastata (Hincks, 1862)

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Notes: 

Morri et al. 1999

Schizomavella (Schizomavella) linearis (Hassall, 1841)

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Notes: 

Gautier 1962, Harmelin 1969, Hayward 1974, Castritsi-Catharios et al. 1985a, Ganias 1990

Schizomavella (Schizomavella) mamillata (Hincks, 1880)

Nomenclature: 

Schizomavella linearis mamillata

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Notes: 

Gautier 1962, Harmelin 1969, Hayward 1974, Morri et al. 1999

Family Bryocryptellidae Vigneaux, 1949

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Genus Bryocryptella Cossmann, 1906

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Bryocryptella tubulata (Busk, 1861)

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Notes: 

Harmelin 1969

Genus Palmiskenea Bishop & Hayward, 1989

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Palmiskenea gautieri Madurell, Zabala, Domínguez-Carrió & Gili, 2013

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Notes: 

Harmelin 1969, Castritsi-Catharios et al. 1985a (as Palmiskenea aff. aviculifera), Castritsi-Catharios et al. 1986a. These records have been provisionally reported as P. gautieri following Madurell et al. (2013) and Rosso and Di Martino (2016).

Palmiskenea skenei (Ellis & Solander, 1786)

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Notes: 

Ganias 1990

Genus Porella Gray, 1848

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Porella concinna (Busk, 1854)

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Notes: 

Harmelin 1969

Porella minuta (Norman, 1868)

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Notes: 

Harmelin 1969

Family Bugulidae Gray, 1848

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Genus Bicellariella Levinsen, 1909

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Bicellariella ciliata (Linnaeus, 1758)

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Notes: 

Antoniadou and Chintiroglou 2005

Genus Bugula Oken, 1815

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Bugula calathus (Norman, 1868)

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Notes: 

Hayward 1974

Bugula neritina (Linnaeus, 1758)

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Distribution: 
NIB
Notes: 

Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios and Kiortis 1985, Castritsi-Catharios and Ganias 1989

Genus Bugulina Gray, 1848

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Bugulina avicularia (Linnaeus, 1758)

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Notes: 

Ganias 1990

Bugulina fulva (Ryland, 1960)

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Distribution: 
NIB
Notes: 

Morri et al. 1999

Bugulina simplex (Hincks, 1886)

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Notes: 

Castritsi-Catharios and Kiortis 1984

Bugulina spicata (Hincks, 1886)

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Notes: 

Castritsi-Catharios and Marcopoulou-Diacantoni 1983

Bugulina stolonifera (Ryland, 1960)

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Notes: 

Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios and Kiortis 1985, Ganias 1990, Antoniadou and Chintiroglou 2005

Bugulina turbinata (Alder, 1857)

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Notes: 

Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios and Kiortis 1985

Genus Crisularia Gray, 1848

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Crisularia aperta (Hincks, 1886)

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Notes: 

Castritsi-Catharios and Marcopoulou-Diacantoni 1983

Crisularia plumosa (Pallas, 1766)

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Distribution: 
NIB
Notes: 

Hayward 1974

Crisularia serrata (Lamarck, 1816)

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Distribution: 
NIB
Notes: 

Harmelin 1969, Ganias 1990

Family Calloporidae Norman, 1903

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Genus Alderina Norman, 1903

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Alderina imbellis (Hincks, 1860)

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Notes: 

Hayward 1974

Genus Aplousina Canu & Bassler, 1927

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Aplousina capriensis (Waters, 1898)

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Notes: 

Hayward 1974

Genus Callopora Gray, 1848

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Callopora dumerilii (Audouin, 1826)

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Notes: 

Hayward 1974, Ganias 1990

Callopora lineata (Linnaeus, 1767)

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Notes: 

Harmelin 1969, Ganias 1990

Genus Copidozoum Harmer, 1926

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Copidozoum planum (Hincks, 1880)

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Notes: 

Hayward 1974, Castritsi-Catharios et al. 1986b, Castritsi-Catharios and Ganias 1989, Ganias 1990

Copidozoum tenuirostre (Hincks, 1880)

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Notes: 

Hayward 1974, Castritsi-Catharios et al. 1986a, Castritsi-Catharios and Ganias 1989, Ganias 1990

Genus Corbulella Gordon, 1984

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Corbulella maderensis (Waters, 1898)

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Notes: 

Hayward 1974, Ganias 1990, Morri et al. 1999

Genus Crassimarginatella Canu, 1900

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Crassimarginatella crassimarginata (Hincks, 1880)

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Notes: 

Gautier 1962

Crassimarginatella solidula (Hincks, 1860)

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Notes: 

Ganias 1990

Genus Ellisina Norman, 1903

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Ellisina gautieri Fernandez Pulpeiro & Reverter Gil, 1993

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Notes: 

Ganias 1990

Genus Parellisina Osburn, 1940

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Parellisina curvirostris (Hincks, 1862)

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Notes: 

Harmelin 1969, Hayward 1974, Ganias 1990

Genus Retevirgula Brown, 1948

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Retevirgula akdenizae Chimenz, Nicoletti & Lippi Boncambi, 1997

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Notes: 

Morri et al. 1999

Family Candidae d'Orbigny, 1851

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Genus Caberea Lamouroux, 1816

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Caberea boryi (Audouin, 1826)

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Notes: 

Harmelin 1969, Hayward 1974, Ganias 1990, Morri et al. 1999, Simboura et al. 1995

Genus Cradoscrupocellaria Vieira, Spencer Jones & Winston, 2013

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Cradoscrupocellaria bertholletii (Audouin, 1826)

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Notes: 

Hayward 1974, Ganias 1990

Cradoscrupocellaria macrorhyncha (Gautier, 1962)

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Notes: 

Harmelin 1969, Hayward 1974

Genus Scrupocaberea Vieira, Spencer Jones, Winston, Migotto & Marques, 2014

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Scrupocaberea maderensis (Busk, 1860)

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Notes: 

Castritsi-Catharios et al. 1986a, Castritsi-Catharios et al. 1986b, Castritsi-Catharios and Ganias 1989, Ganias 1990

Genus Scrupocellaria Van Beneden, 1845

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Scrupocellaria aegeensis Harmelin, 1969

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Notes: 

Harmelin 1969, Ganias 1990

Scrupocellaria delilii (Audouin, 1826)

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Notes: 

Harmelin 1969, Ganias 1990, Morri et al. 1999

Scrupocellaria incurvata Waters, 1897

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Notes: 

Harmelin 1969, Ganias 1990

Scrupocellaria muricata (Lamouroux, 1816)

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Notes: 

Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios et al. 1986a

Scrupocellaria scrupea Busk, 1852

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Notes: 

Gautier 1962, Harmelin 1969, Hayward 1974, Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios and Kiortis 1985, Castritsi-Catharios et al. 1986a, Ganias 1990, Simboura et al. 1995, Morri et al. 1999

Scrupocellaria scruposa (Linnaeus, 1758)

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Distribution: 
NIB
Notes: 

Hayward 1974, Ganias 1990, Simboura et al. 1995, Morri et al. 1999

Family Cellariidae Fleming, 1828

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Genus Cellaria Ellis & Solander, 1786

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Cellaria fistulosa (Linnaeus, 1758)

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Notes: 

Ganias 1990

Cellaria normani (Hastings, 1947)

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Notes: 

Harmelin 1969

Cellaria salicornioides Lamouroux, 1816

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Notes: 

Hayward 1974, Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios and Kiortis 1985, Ganias 1990

Family Celleporidae Johnston, 1838

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Genus Buskea Heller, 1867

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Buskea dichotoma (Hincks, 1862)

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Notes: 

Ganias 1990

Buskea nitida Heller, 1867

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Notes: 

Harmelin 1969

Genus Cellepora Linnaeus, 1767

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Cellepora posidoniae (Hayward, 1975)

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Notes: 

Ganias 1990

Cellepora pumicosa (Pallas, 1766)

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Notes: 

Morri et al. 1999

Genus Celleporina Gray, 1848

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Celleporina caminata (Waters, 1879)

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Notes: 

Gautier 1962, Harmelin 1969, Hayward 1974, Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios et al. 1985a, Castritsi-Catharios et al. 1986a, Ganias 1990, Morri et al. 1999

Celleporina hassallii (Johnston, 1847)

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Notes: 

Castritsi-Catharios and Kiortis 1984 (as Cellepora hassalli), Castritsi-Catharios et al. 1985a, Castritsi-Catharios et al. 1985b, Castritsi-Catharios et al. 1986a, Ganias 1990, Simboura et al. 1995, Morri et al. 1999

Celleporina lucida (Hincks, 1880)

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Notes: 

Morri et al. 1999

Celleporina tubulosa (Hincks, 1880)

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Notes: 

Harmelin 1969, Morri et al. 1999

Genus Palmicellaria Alder, 1864

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Palmicellaria elegans Alder, 1864

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Notes: 

Harmelin 1969, Ganias 1990

Genus Turbicellepora Ryland, 1963

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Turbicellepora avicularis (Hincks, 1860)

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Notes: 

Ganias 1990

Turbicellepora camera Hayward, 1978

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Notes: 

Hayward 1978, Ganias 1990, Morri et al. 1999

Turbicellepora coronopus (Wood, 1844)

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Notes: 

Hayward 1974, Ganias 1990, Gerovasileiou et al. 2015

Turbicellepora crenulata Hayward, 1978

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Notes: 

Ganias 1990

Turbicellepora magnicostata (Barroso, 1919)

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Notes: 

Ganias 1990

Family Chlidoniidae Busk, 1884

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Genus Chlidonia Lamouroux, 1824

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Chlidonia pyriformis (Bertoloni, 1810)

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Notes: 

Harmelin 1969, Hayward 1974, Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios et al. 1986a, Castritsi-Catharios et al. 1986b, Ganias 1990, Morri et al. 1999

Family Chorizoporidae Vigneaux, 1949

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Genus Chorizopora Hincks, 1879

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Chorizopora brongniartii (Audouin, 1826)

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Notes: 

Gautier 1962, Harmelin 1969, Hayward 1974, Castritsi-Catharios and Marcopoulou-Diacantoni 1983, Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios et al. 1985a, Castritsi-Catharios et al. 1986a, Castritsi-Catharios and Ganias 1989, Ganias 1990, Simboura et al. 1995, Morri et al. 1999, Antoniadou and Chintiroglou 2005

Family Colatooeciidae Winston, 2005

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Genus Trematooecia Osburn, 1940

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Trematooecia ligulata Ayari & Taylor, 2008

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Notes: 

Harmelin 1969 (as Cigclisula turrita)

Family Crepidacanthidae Levinsen, 1909

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Genus Crepidacantha Levinsen, 1909

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Crepidacantha poissonii (Audouin, 1826)

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Distribution: 
NIB
Notes: 

Ganias 1990

Family Cribrilinidae Hincks, 1879

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Genus Collarina Jullien, 1886

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Collarina balzaci (Audouin, 1826)

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Notes: 

Hayward 1974, Castritsi-Catharios and Marcopoulou-Diacantoni 1983, Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios and Kiortis 1985, Castritsi-Catharios et al. 1986a, Castritsi-Catharios and Ganias 1989, Ganias 1990

Genus Figularia Jullien, 1886

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Figularia figularis (Johnston, 1847)

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Notes: 

Harmelin 1969, Hayward 1974, Ganias 1990

Genus Puellina Jullien, 1886

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Puellina (Cribrilaria) innominata (Couch, 1844)

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Notes: 

Harmelin 1969, Hayward 1974, Castritsi-Catharios et al. 1986a, Castritsi-Catharios et al. 1986b, Castritsi-Catharios and Ganias 1989, Ganias 1990

Puellina (Cribrilaria) hincksi (Friedl, 1917)

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Notes: 

Gautier 1962, Morri et al. 1999

Puellina (Cribrilaria) radiata (Moll, 1803)

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Notes: 

Harmelin 1969, Hayward 1974, Castritsi-Catharios and Marcopoulou-Diacantoni 1983, Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios and Kiortis 1985, Castritsi-Catharios et al. 1986a, Castritsi-Catharios and Ganias 1989, Ganias 1990, Simboura et al. 1995, Morri et al. 1999

Puellina (Cribrilaria) venusta (Canu & Bassler, 1925)

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Notes: 

Ganias 1990

Puellina (Glabrilaria) orientalis Harmelin & Aristegui, 1987

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Notes: 

Morri et al. 1999

Puellina (Glabrilaria) pedunculata (Gautier, 1956)

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Notes: 

Harmelin 1969, Hayward 1974

Puellina (Puellina) gattyae (Landsborough, 1852)

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Notes: 

Gautier 1962, Hayward 1974, Castritsi-Catharios and Marcopoulou-Diacantoni 1983, Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios et al. 1986a, Castritsi-Catharios et al. 1986b, Castritsi-Catharios and Ganias 1989, Ganias 1990, Morri et al. 1999

Puellina (Puellina) setosa (Waters, 1899)

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Notes: 

Harmelin 1969, Ganias 1990, Morri et al. 1999

Family Cryptosulidae Vigneaux, 1949

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Genus Cryptosula Canu & Bassler, 1925

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Cryptosula pallasiana (Moll, 1803)

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Notes: 

Hayward 1974, Castritsi-Catharios and Marcopoulou-Diacantoni 1983, Castritsi-Catharios et al. 1985a, Castritsi-Catharios et al. 1986a, Castritsi-Catharios et al. 1986b, Ganias 1990, Antoniadou and Chintiroglou 2005

Family Electridae d'Orbigny, 1851

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Genus Conopeum Gray, 1848

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Conopeum seurati (Canu, 1928)

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Notes: 

Ganias 1990

Genus Electra Lamouroux, 1816

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Electra pilosa (Linnaeus, 1767)

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Notes: 

Castritsi-Catharios et al. 1986a, Castritsi-Catharios and Ganias 1989, Ganias 1990, Antoniadou and Chintiroglou 2005

Electra posidoniae Gautier, 1954

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Notes: 

Hayward 1974, Castritsi-Catharios and Marcopoulou-Diacantoni 1983, Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios and Kiortis 1985, Castritsi-Catharios et al. 1985a, Castritsi-Catharios et al. 1986a, Castritsi-Catharios and Ganias 1989, Ganias 1990

Family Epistomiidae Gregory, 1893

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Genus Synnotum Pieper, 1881

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Synnotum aegyptiacum (Audouin, 1826)

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Notes: 

Harmelin 1969, Hayward 1974, Ganias 1990

Family Escharinidae Tilbrook, 2006

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Genus Escharina Milne Edwards, 1836

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Escharina dutertrei subsp. protecta Zabala, Maluquer & Harmelin, 1993

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Notes: 

Harmelin 1969, Hayward 1974, Morri et al. 1999

Escharina johnstoni (Quelch, 1884)

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Notes: 

Ganias 1990

Escharina vulgaris (Moll, 1803)

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Notes: 

Harmelin 1969, Hayward 1974, Castritsi-Catharios et al. 1986a (as Schizoporella vulgaris), Castritsi-Catharios et al. 1986b (as S. vulgaris), Ganias 1990, ​Morri et al. 1999

Genus Herentia Gray, 1848

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Herentia hyndmanni (Johnston, 1847)

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Notes: 

Harmelin 1969, Morri et al. 1999

Genus Hippomenella Canu & Bassler, 1917

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Hippomenella mucronelliformis (Waters, 1899)

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Notes: 

Harmelin 1969, Hayward 1974, Ganias 1990

Genus Therenia David & Pouyet, 1978

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Therenia rosei Berning, Tilbrook & Rosso, 2008

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Notes: 

Hayward 1974 (as Escharina porosa), Ganias 1990 (as E. porosa), Berning et al. 2008

Family Exechonellidae Harmer, 1957

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Genus Anarthropora Smitt, 1868

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Anarthropora monodon (Busk, 1860)

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Notes: 

Harmelin 1969, Hayward 1974, Ganias 1990, Morri et al. 1999

Genus Exechonella Duvergier, 1924

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Exechonella antillea (Osburn, 1927)

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Distribution: 
NIB
Notes: 

Hayward 1974

Family Exochellidae Bassler, 1935

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Genus Escharoides Milne Edwards, 1836

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Escharoides coccinea (Abildgaard, 1806)

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Notes: 

Harmelin 1969, Hayward 1974, Castritsi-Catharios and Marcopoulou-Diacantoni 1983, Castritsi-Catharios and Kiortis 1984 (as Cellepora coccinea), Castritsi-Catharios et al. 1986a, Castritsi-Catharios and Ganias 1989, Ganias 1990, Morri et al. 1999​, Antoniadou and Chintiroglou 2005

Family Flustridae Fleming, 1828

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Genus Chartella Gray, 1848

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Chartella papyrea (Pallas, 1766)

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Notes: 

Hayward 1974

Genus Gregarinidra Barroso, 1949

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Gregarinidra gregaria (Heller, 1867)

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Notes: 

Harmelin 1969, Hayward 1974, Castritsi-Catharios and Kiortis 1984

Genus Hincksina Norman, 1903

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Hincksina flustroides (Hincks, 1877)

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Notes: 

Hayward 1974, Ganias 1990, Morri et al. 1999

Family Haplopomidae Gordon in De Blauwe, 2009

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Genus Haplopoma Levinsen, 1909

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Haplopoma impressum (Audouin, 1826)

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Notes: 

Harmelin 1969, Hayward 1974, Ganias 1990, Morri et al. 1999

Family Hippaliosinidae Winston, 2005

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Genus Hippaliosina Canu, 1919

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Hippaliosina depressa (Busk, 1854)

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Notes: 

Busk 1854, Harmelin 1969, Hayward 1974, Castritsi-Catharios and Marcopoulou-Diacantoni 1983, Castritsi-Catharios et al. 1985a, Castritsi-Catharios et al. 1985b, Castritsi-Catharios et al. 1986a, Castritsi-Catharios and Ganias 1989, Ganias 1990, Morri et al. 1999, Gerovasileiou et al. 2013, Gerovasileiou et al. 2015

Family Hippopodinidae Levinsen, 1909

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Genus Hippopodina Levinsen, 1909

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Hippopodina ambita (Hayward, 1974)

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Notes: 

Harmelin 1969 (as Cosciniopsis sp.), Hayward 1974, Ganias 1990

Hippopodina feegeensis (Busk, 1884)

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Distribution: 
NIB
Notes: 

Morri et al. 1999

Family Hippoporidridae Vigneaux, 1949

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Genus Hagiosynodos Bishop & Hayward, 1989

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Hagiosynodos latus (Busk, 1856)

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Notes: 

Hayward 1974, Ganias 1990, Morri et al. 1999

Genus Hippoporidra Canu & Bassler, 1927

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Hippoporidra picardi Gautier, 1962

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Notes: 

Gautier 1962, Morri et al. 1999

Family Hippothoidae Busk, 1859

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Genus Hippothoa Lamouroux, 1821

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Hippothoa flagellum Manzoni, 1870

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Notes: 

Ganias 1990

Family Lacernidae Jullien, 1888

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Genus Arthropoma Levinsen, 1909

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Arthropoma cecilii (Audouin, 1826)

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Notes: 

Hayward 1974, Ganias 1990

Family Lanceoporidae Harmer, 1957

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Genus Calyptotheca Harmer, 1957

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Calyptotheca rugosa Hayward, 1974

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Notes: 

Hayward 1974, Ganias 1990

Family Margarettidae Harmer, 1957

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Genus Margaretta Gray, 1843

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Margaretta cereoides (Ellis & Solander, 1786)

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Notes: 

Harmelin 1969, Hayward 1974, Castritsi-Catharios et al. 1985a, Ganias 1990

Family Microporellidae Hincks, 1879

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Genus Diporula Hincks, 1879

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Diporula verrucosa (Peach, 1868)

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Notes: 

Harmelin 1969, Ganias 1990

Genus Fenestrulina Jullien, 1888

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Fenestrulina malusii (Audouin, 1826)

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Notes: 

Harmelin 1969, Hayward 1974, Castritsi-Catharios et al. 1986b, Castritsi-Catharios and Ganias 1989, Ganias 1990, Morri et al. 1999

Genus Microporella Hincks, 1877

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Microporella coronata (Audouin, 1826)

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Distribution: 
NIB
Notes: 

Harmelin et al. (2011) suggested that specimens from the Aegean Sea reported as Microporella orientalis could belong to M. coronata (followed by Rosso and Di Martino 2016). Recorded by Hayward 1974, Morri et al. 1999.

Microporella ciliata (Pallas, 1766)

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Notes: 

This taxon corresponds to a complex of species (Rosso et al. 2010, Rosso and Di Martino 2016). Recorded by Harmelin 1969, Castritsi-Catharios and Marcopoulou-Diacantoni 1983, Castritsi-Catharios et al. 1986a, Ganias 1990, Antoniadou and Chintiroglou 2005.

Microporella marsupiata (Busk, 1860)

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Notes: 

Hayward 1974

Family Microporidae Gray, 1848

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Genus Calpensia Jullien, 1888

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Calpensia nobilis (Esper, 1796)

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Notes: 

Busk 1854, Hayward 1974, Castritsi-Catharios and Marcopoulou-Diacantoni 1983, Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios and Kiortis 1985, Castritsi-Catharios and Ganias 1989, Ganias 1990, Morri et al. 1999

Genus Coronellina Prenant & Bobin, 1966

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Coronellina fagei (Gautier, 1962)

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Notes: 

Harmelin 1969, Hayward 1974

Genus Micropora Gray, 1848

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Micropora coriacea (Johnston, 1847)

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Notes: 

Hayward 1974, Ganias 1990

Genus Mollia Lamouroux, 1821

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Mollia multijuncta (Waters, 1879)

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Notes: 

Hayward 1974, Morri et al. 1999

Mollia patellaria (Moll, 1803)

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Notes: 

Gautier 1962, Harmelin 1969, Hayward 1974, Ganias 1990, Morri et al. 1999

Family Monoporellidae Hincks, 1882

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Genus Monoporella Hincks, 1881

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Monoporella bouchardii (Audouin, 1826)

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Notes: 

Mediterranean specimens of Monoporella nodulifera and M. fimbriata carinifera actually belong to this species, as suggested by Harmelin (2014a) and followed by Rosso and Di Martino (2016). Recorded by Harmelin 1969, Hayward 1974, Ganias 1990, Morri et al. 1999.

Family Myriaporidae Gray, 1841

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Genus Myriapora de Blainville, 1830

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Myriapora truncata (Pallas, 1766)

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Notes: 

Harmelin 1969, Hayward 1974, Morri et al. 1999, Gerovasileiou et al. 2015

Family Onychocellidae Jullien, 1882

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Genus Onychocella Jullien, 1882

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Onychocella angulosa (Reuss, 1847)

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Notes: 

Hayward 1974, Ganias 1990

Onychocella marioni (Jullien, 1881)

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Notes: 

Harmelin 1969, Morri et al. 1999, Gerovasileiou et al. 2015

Onychocella vibraculifera Neviani, 1895

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Notes: 

Hayward 1974, Castritsi-Catharios and Marcopoulou-Diacantoni 1983, Ganias 1990

Genus Smittipora Jullien, 1882

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Smittipora disjuncta (Canu & Bassler, 1930)

Nomenclature: 

Smittipora is a synonym of Rectonychocella (see Rosso and Di Martino 2016​).

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Notes: 

Harmelin 1969, Hayward 1974, Ganias 1990

Family Phidoloporidae Gabb & Horn, 1862

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Genus Dentiporella Barroso, 1926

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Dentiporella sardonica (Waters, 1879)

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Notes: 

Gautier 1962, Harmelin 1969

Genus Plesiocleidochasma Soule, Soule & Chaney, 1991

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Plesiocleidochasma mediterraneum Chimenz-Gusso & Soule, 2003

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Notes: 

Harmelin 1969 (as Cleidochasma sp.), Hayward 1974 (as Cleidochasma porcellanum)

Genus Reteporella Busk, 1884

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Reteporella complanata (Waters, 1894)

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Notes: 

Ganias 1990

Reteporella couchii (Hincks, 1878)

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Notes: 

Harmelin 1969, Ganias 1990

Reteporella grimaldii (Jullien & Calvet, 1903)

Nomenclature: 

Synonym valid for R. septentrionalis (see Rosso et al. 2010​).

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Notes: 

Hayward 1974, Ganias 1990, Morri et al. 1999, Antoniadou and Chintiroglou 2005, Chimenz Gusso et al. 2014

Reteporella mediterranea (Hass, 1948)

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Notes: 

Hayward 1974, Ganias 1990

Genus Reteporellina Harmer, 1933

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Reteporellina delicatula Hayward, 1974

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Notes: 

Hayward 1974, Harmelin 1976, Ganias 1990

Genus Rhynchozoon Hincks, 1895

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Rhynchozoon bispinosum (Johnston, 1847)

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Notes: 

Hayward 1974, Ganias 1990

Rhynchozoon neapolitanum Gautier, 1962

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Notes: 

Harmelin 1969, Hayward 1974, Ganias 1990, Morri et al. 1999, Gerovasileiou et al. 2015

Rhynchozoon pseudodigitatum Zabala & Maluquer, 1988

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Notes: 

Morri et al. 1999

Rhynchozoon sp. 1 sensu Hayward, 1974

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Notes: 

Hayward 1974

Genus Schizoretepora Gregory, 1893

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Schizoretepora imperati (Busk, 1884)

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Notes: 

Chimenz Gusso et al. 2014

Schizoretepora serratimargo (Hincks, 1886)

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Notes: 

Gautier 1962, Harmelin 1969, Ganias 1990

Schizoretepora solanderia (Risso, 1826)

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Notes: 

Harmelin 1969, Ganias 1990

Genus Schizotheca Hincks, 1877

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Schizotheca fissa (Busk, 1856)

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Notes: 

Harmelin 1969, Hayward 1974, Ganias 1990

Genus Stephanollona Duvergier, 1921

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Stephanollona armata (Hincks, 1861)

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Notes: 

Harmelin 1969, Hayward 1974, Castritsi-Catharios and Marcopoulou-Diacantoni 1983, Castritsi-Catharios et al. 1985a, Castritsi-Catharios et al. 1985b, Ganias 1990

Family Phoceanidae Vigneaux, 1949

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Genus Phoceana Jullien in Jullien & Calvet, 1903

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Phoceana tubulifera (Heller, 1867)

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Notes: 

Harmelin 1969, Hayward 1974, Ganias 1990

Family Romancheinidae Jullien, 1888

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Genus Escharella Gray, 1848

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Escharella rylandi Geraci, 1974

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Notes: 

Ganias 1990

Genus Hemicyclopora Norman, 1894

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Hemicyclopora multispinata (Busk, 1861)

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Notes: 

Ganias 1990, Morri et al. 1999

Genus Hippopleurifera Canu & Bassler, 1925

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Hippopleurifera pulchra (Manzoni, 1870)

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Notes: 

Harmelin 1969

Family Savignyellidae Levinsen, 1909

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Genus Savignyella Levinsen, 1909

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Savignyella lafontii (Audouin, 1826)

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Notes: 

Gautier 1962, Hayward 1974, Morri et al. 1999

Family Schizoporellidae Jullien, 1883

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Genus Schizobrachiella Canu & Bassler, 1920

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Schizobrachiella sanguinea (Norman, 1868)

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Notes: 

Gautier 1962, Harmelin 1969, Ganias 1990, Morri et al. 1999

Genus Schizoporella Hincks, 1877

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Schizoporella dunkeri (Reuss, 1848)

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Notes: 

Harmelin 1969, Hayward 1974, Castritsi-Catharios and Marcopoulou-Diacantoni 1983, Castritsi-Catharios et al. 1986a, Morri et al. 1999, Ganias 1990

Schizoporella errata (Waters, 1878)

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Notes: 

Hayward 1974, Castritsi-Catharios et al. 1985a, Ganias 1990

Schizoporella magnifica Hincks, 1886

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Notes: 

Harmelin 1969, Hayward 1974, Ganias 1990

Schizoporella unicornis (Johnston in Wood, 1844)

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Notes: 

Castritsi-Catharios et al. 1986a, Ganias 1990, Morri et al. 1999, Antoniadou and Chintiroglou 2005

Family Setosellidae Levinsen, 1909

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Genus Setosella Hincks, 1877

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Setosella vulnerata (Busk, 1860)

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Notes: 

Hayward 1974, Ganias 1990

Family Smittinidae Levinsen, 1909

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Genus Parasmittina Osburn, 1952

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Parasmittina raigii (Audouin, 1826)

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Distribution: 
NIB
Notes: 

Ganias 1990

Parasmittina rouvillei (Calvet, 1902)

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Notes: 

Harmelin 1969, Hayward 1974, Ganias 1990

Genus Prenantia Gautier, 1962

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Prenantia cheilostoma (Manzoni, 1869)

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Notes: 

Hayward 1974, Ganias 1990

Prenantia inerma (Calvet, 1906)

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Notes: 

Harmelin 1969

Genus Smittina Norman, 1903

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Smittina cervicornis (Pallas, 1766)

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Notes: 

Gautier 1962 (as Porella cervicornis), Ganias 1990 (as P. cervicornis), Morri et al. 1999, Chimenz Gusso et al. 2014

Smittina colleti (Jullien & Calvet, 1903)

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Notes: 

Harmelin 1969, Ganias 1990

Smittina landsborovii (Johnston, 1847)

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Notes: 

Ganias 1990, Antoniadou and Chintiroglou 2005

Smittina remotorostrata (Canu & Bassler, 1928)

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Notes: 

Hayward 1974

Genus Smittoidea Osburn, 1952

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Smittoidea marmorea (Hincks, 1877)

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Notes: 

Hayward 1974, Ganias 1990

Smittoidea ophidiana (Waters, 1879)

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Notes: 

Harmelin 1969, Hayward 1974

Smittoidea reticulata (MacGillivray, 1842)

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Notes: 

Gautier 1962, Morri et al. 1999, Antoniadou and Chintiroglou 2005

Family Tessaradomidae Jullien, 1903

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Genus Tessaradoma Norman, 1869

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Tessaradoma boreale (Busk, 1860)

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Notes: 

Ganias 1990

Family Trypostegidae Gordon, Tilbrook & Winston, 2005

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Genus Trypostega Levinsen, 1909

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Trypostega venusta (Norman, 1864)

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Notes: 

Ganias 1990

Family Umbonulidae Canu, 1904

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Genus Umbonula Hincks, 1880

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Umbonula ovicellata Hastings, 1944

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Notes: 

Hayward 1974, Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios et al. 1986a, Ganias 1990, Morri et al. 1999

Family Watersiporidae Vigneaux, 1949

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Genus Watersipora Neviani, 1896

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Watersipora complanata (Norman, 1864)

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Notes: 

Ganias 1990

Watersipora cucullata (Busk, 1854)

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Notes: 

Busk 1854, Hayward 1974, Castritsi-Catharios et al. 1986b, Castritsi-Catharios and Ganias 1989, Ganias 1990. Records of Watersipora subovoidea (d'Orbigny, 1852) probably belong to W. cucullata.

Order Ctenostomatida Busk, 1852

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Family Alcyonidiidae Johnston, 1838

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Genus Alcyonidium Lamouroux, 1813

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Alcyonidium cellarioides (Calvet, 1900)

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Notes: 

Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios and Kiortis 1985

Alcyonidium duplex Prouho, 1892

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Notes: 

Ganias 1990

Alcyonidium polyoum (Hassall, 1841)

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Notes: 

Ganias 1990

Family Buskiidae Hincks, 1880

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Genus Buskia Alder, 1857

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Buskia nitens Alder, 1857

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Notes: 

Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios and Kiortis 1985, Ganias 1990

Family Farrellidae d'Hondt, 1983

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Genus Farrella Ehrenberg, 1838

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Farrella repens (Farre, 1837)

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Notes: 

Castritsi-Catharios et al. 1986a, Castritsi-Catharios et al. 1986b

Family Mimosellidae Hincks, 1877

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Genus Bantariella Jebram, 1973

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Bantariella verticillata (Heller, 1867)

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Notes: 

Castritsi-Catharios and Marcopoulou-Diacantoni 1983, Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios and Kiortis 1985, Castritsi-Catharios et al. 1985a, Castritsi-Catharios et al. 1986a, Castritsi-Catharios et al. 1986b, Castritsi-Catharios and Ganias 1989, Ganias 1990, Morri et al. 1999

Genus Mimosella Hincks, 1851

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Mimosella gracilis Hincks, 1851

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Notes: 

Harmelin 1969, Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios and Kiortis 1985, Castritsi-Catharios et al. 1985a, Castritsi-Catharios et al. 1986a, Castritsi-Catharios and Ganias 1989, Ganias 1990, Morri et al. 1999

Family Nolellidae Harmer, 1915

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Genus Nolella Gosse, 1851

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Nolella dilatata (Hincks, 1860)

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Notes: 

Harmelin 1969, Ganias 1990

Nolella stipata Gosse, 1855

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Notes: 

Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios et al. 1986a, Castritsi-Catharios and Ganias 1989, Ganias 1990, Morri et al. 1999

Family Pherusellidae Osburn & Soule, 1953

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Genus Pherusella Soule, 1951

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Pherusella tubulosa (Ellis & Solander, 1786)

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Notes: 

Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios and Kiortis 1985, Castritsi-Catharios et al. 1986a, Ganias 1990, Morri et al. 1999

Family Vesiculariidae Hincks, 1880

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Genus Amathia Lamouroux, 1812

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Amathia gracilis (Leidy, 1855)

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Notes: 

Ganias 1990, Morri et al. 1999

Amathia gracillima (Hincks, 1877)

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Distribution: 
NIB
Notes: 

Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios and Kiortis 1985, Ganias 1990

Amathia imbricata (Adams, 1798)

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Notes: 

Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios and Kiortis 1985, Castritsi-Catharios et al. 1986a, Castritsi-Catharios et al. 1986b, Castritsi-Catharios and Ganias 1989, Ganias 1990

Amathia lendigera (Linnaeus, 1758)

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Notes: 

Castritsi-Catharios and Ganias 1989, Ganias 1990, Morri et al. 1999

Amathia pruvoti Calvet, 1911

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Notes: 

Castritsi-Catharios and Marcopoulou-Diacantoni 1983, Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios and Kiortis 1985, Castritsi-Catharios et al. 1986a, Castritsi-Catharios and Ganias 1989, Ganias 1990, Morri et al. 1999

Amathia pustulosa (Ellis & Solander, 1786)

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Notes: 

Castritsi-Catharios et al. 1986a, Castritsi-Catharios et al. 1986b

Amathia semiconvoluta Lamouroux, 1824

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Notes: 

Castritsi-Catharios and Ganias 1989, Ganias 1990

Amathia verticillata (Delle Chiaje, 1822)

Material    Download as CSV 
Distribution: 
NIB
Notes: 

Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios and Kiortis 1985, Castritsi-Catharios et al. 1986a, Castritsi-Catharios et al. 1986b, Castritsi-Catharios and Ganias 1989, Ganias 1990, Conides et al. 1999

Family Victorellidae Hincks, 1880

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Genus Victorella Saville-Kent, 1870

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Victorella pavida Saville-Kent, 1870

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Notes: 

Castritsi-Catharios et al. 1986a, Castritsi-Catharios et al. 1986b

Family Walkeriidae Hincks, 1880

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Genus Walkeria Fleming, 1823

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Walkeria tuberosa Heller, 1867

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Notes: 

Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios et al. 1985a, Castritsi-Catharios et al. 1986a, Castritsi-Catharios and Ganias 1989, Ganias 1990, Morri et al. 1999

Walkeria uva (Linnaeus, 1758)

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Notes: 

Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios et al. 1986a, Castritsi-Catharios and Ganias 1989, Ganias 1990

Class Stenolaemata Borg, 1926

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Order Cyclostomatida Busk, 1852

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Family Annectocymidae Hayward & Ryland, 1985

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Genus Annectocyma Hayward & Ryland, 1985

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Annectocyma arcuata (Harmelin, 1976)

Material    Download as CSV 
Notes: 

Ganias 1990

Annectocyma major (Johnston, 1847)

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Notes: 

Castritsi-Catharios et al. 1985a (as Stomatopora major), Castritsi-Catharios et al. 1986a, Castritsi-Catharios et al. 1986b, Castritsi-Catharios and Ganias 1989 (as S. major), Ganias 1990, Simboura et al. 1995, Morri et al. 1999

Annectocyma tubulosa (Busk, 1875)

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Notes: 

Ganias 1990

Genus Entalophoroecia Harmelin, 1976

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Entalophoroecia deflexa (Couch, 1844)

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Castritsi-Catharios et al. 1986a, Castritsi-Catharios et al. 1986b, Castritsi-Catharios and Ganias 1989, Ganias 1990

Entalophoroecia gracilis Harmelin, 1976

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Ganias 1990

Entalophoroecia robusta Harmelin, 1976

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Ganias 1990

Family Crisiidae Johnston, 1838

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Genus Crisia Lamouroux, 1812

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Crisia denticulata (Lamarck, 1816)

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Ganias 1990

Crisia eburnea (Linnaeus, 1758)

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Castritsi-Catharios and Kiortis 1984, Castritsi-Catharios et al. 1986a, Castritsi-Catharios et al. 1986b, Castritsi-Catharios and Ganias 1989, Ganias 1990

Crisia elongata Milne Edwards, 1838

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Castritsi-Catharios et al. 1986a, Castritsi-Catharios et al. 1986b

Crisia fistulosa (Heller, 1867)

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Castritsi-Catharios and Marcopoulou-Diacantoni 1983, Castritsi-Catharios et al. 1986a, Ganias 1990

Crisia pyrula Harmelin, 1990

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Harmelin 1969 (as Crisia sp. II), Ganias 1990