Biodiversity Data Journal : Taxonomic Paper
Print
Taxonomic Paper
Megafauna of the UKSRL exploration contract area and eastern Clarion-Clipperton Zone in the Pacific Ocean: Echinodermata
expand article infoDiva J Amon, Amanda F Ziegler, Antonina Kremenetskaia§, Christopher L Mah|, Rich Mooi, Tim O'Hara#, David L Pawson|, Michel Roux¤, Craig R Smith
‡ University of Hawaii at Manoa, Honolulu, United States of America
§ P.P. Shirshov Institute of Oceanology, Moscow, Russia
| Smithsonian Institution National Museum of Natural History, Washington, United States of America
¶ California Academy of Sciences, San Francisco, United States of America
# Museum Victoria, Melbourne, Australia
¤ Museum National d’Histoire Naturelle, Paris, France
Open Access

Abstract

Background

There is growing interest in mining polymetallic nodules from the abyssal Clarion-Clipperton Zone (CCZ) in the tropical Pacific Ocean. Despite being the focus of environmental studies for decades, the benthic megafauna of the CCZ remain poorly known. In order to predict and manage the environmental impacts of mining in the CCZ, baseline knowledge of the megafauna is essential. The ABYSSLINE Project has conducted benthic biological baseline surveys in the UK Seabed Resources Ltd polymetallic-nodule exploration contract area (UK-1). Prior to these research cruises in 2013 and 2015, no biological studies had been done in this area of the eastern CCZ.

New information

Using a Remotely Operated Vehicle and Autonomous Underwater Vehicle, the megafauna within the UKSRL exploration contract area (UK-1) and at a site ~250 km east of the UK-1 area were surveyed, allowing us to make the first estimates of megafaunal morphospecies richness from the imagery collected. Here, we present an atlas of the abyssal echinoderm megafauna observed and collected during the ABYSSLINE cruises to the UK-1 polymetallic-nodule exploration contract area in the CCZ. There appear to be at least 62 distinct morphospecies (13 Asteroidea, 5 Crinoidea, 9 Echinoidea, 29 Holothuroidea and 6 Ophiuroidea) identified mostly by imagery but also using molecular barcoding for a limited number of animals that were collected. This atlas will aid the synthesis of megafaunal presence/absence data collected by contractors, scientists and other stakeholders undertaking work in the CCZ, ultimately helping to decipher the biogeography of the megafauna in this threatened habitat.

Keywords

deep-sea mining, polymetallic nodule, Clarion-Clipperton Zone, megafauna, echinoderm, atlas

Introduction

The Clarion-Clipperton Zone (CCZ) is an abyssal region of the tropical eastern Pacific Ocean where deep-sea mining may take place in the near future (Ramirez-Llodra et al. 2011, Wedding et al. 2013; Fig. 1). High-grade polymetallic nodules, which could provide a commercial source of copper, cobalt, nickel, and manganese (among other metals), are abundant in this six million km2 region that lies in Areas Beyond National Jurisdiction (ABNJ), and thus falls under the legal mandate of the International Seabed Authority (ISA) (Wedding et al. 2013). Thus far, there have been 16 exploration leases (each up to 75,000 km2 in area) granted by the International Seabed Authority in the CCZ, with those for exploitation expected to soon follow (https://www.isa.org.jm/).

Figure 1.

Locations of megafaunal surveys during the ABYSSLINE cruises, AB01 and AB02, in the Clarion-Clipperton Zone. (a) The location of the UKSRL exploration contract area (UK-1) in the eastern Pacific Ocean. (b) The locations of the 30x30-km survey areas, UK-1 Stratum A and UK-1 Stratum B, in relation to the UK-1 exploration contract area and the AB01 ROV dive site, EPIRB, which was approximately 250 km east of the UK-1 contract area. (c) The locations of ROV dives within UK-1 Stratum A, indicated by purple tracklines labelled with the dive number (e.g. RV01). Stations where samples were collected with a Brenke epibenthic sled (EB04) and megacorer (MC10) are also indicated. (d) The locations of AUV dives within UK-1 Stratum B, indicated by purple tracklines labelled with the dive number (e.g. AV01). Stations where samples were collected with a Brenke epibenthic sled (EB02), box corer (BC02, BC19) and megacorer (MC17, MC25) are also indicated, as well as where imagery was collected with a baited camera (CA02). All maps were created by Seafloor Investigations Ltd for the ABYSSLINE Project using ArcGIS software (https://www.arcgis.com/features/).

The ABYSSLINE (ABYSSal BaseLINE) Project was designed to undertake benthic biological baseline studies in accordance with ISA environmental guidelines within the UK Seabed Resources Ltd (UKSRL) exploration contract area (UK-1) (Amon et al. 2016b). The UK-1 exploration contract area is one of the easternmost contract areas in the CCZ and encompasses ~58,000 km2 of seafloor (Fig. 1). The ABYSSLINE Project was led by scientists from the University of Hawai’i at Mānoa (USA), and included scientists from Hawai’i Pacific University (USA), the Natural History Museum, London (UK), the National Oceanography Centre, Southampton (UK), Senckenberg Gesellschaft für Naturforschung (Germany), Uni Research (Norway), and the International Research Institute of Stavanger (Norway). The ABYSSLINE Project aimed to evaluate baseline conditions of community structure and biodiversity for megafauna, macrofauna, meiofauna and microbes within the UK-1 contract area and across the CCZ (Amon et al. 2016a, Amon et al. 2016b, Dahlgren et al. 2016, Glover et al. 2015, Glover et al. 2016, Shulse et al. 2016). No faunal studies had been undertaken in the UK-1 contract area prior to licensing in 2013.

It is expected that nodule mining will drastically alter this unique deep-sea habitat with recovery expected to be slow (Amon et al. 2016b, Vanreusel et al. 2016, Ramirez-Llodra et al. 2011, Oebius et al. 2001) and yet, despite increases in technology and the number of expeditions to the area, very little is known about the ecology and biogeography of the fauna inhabiting the region (Amon et al. 2016a, Amon et al. 2016b, Dahlgren et al. 2016, Foell and Pawson 1986, Bluhm and Gebruk 1999, Glover et al. 2016, Martinez-Arbizu et al. 2013, Pawson 1983, Pawson and Foell 1986, Roux 2004, Roux and Pawson 1999, Shulse et al. 2016, Tilot 2006, Vanreusel et al. 2016). The megafauna constitute an important component of the biodiversity in the abyssal deep sea and play a significant role in deep-sea ecosystem function (Amon et al. 2016a, Amon et al. 2016b, Smith et al. 2008, Vanreusel et al. 2016). It has also been suggested that echinoderms may act as indicators of physical disturbance of the seabed, such as that caused by deep-sea polymetallic-nodule mining (Bluhm and Gebruk 1999). Glover et al. 2016 reported that a search of OBIS listed 698 echinoderm species recorded at abyssal depths worldwide between 3000m and 6000m, but only 50 species within the CCZ. Amon et al. 2016b was able to confirm that there were no echinoderm or megafauna records for the UK-1 exploration contract area in OBIS. This is likely the result of lack of sampling, taxonomy and/or ensuring data are publicly available, especially as an abundant and diverse echinoderm fauna is already known from the tropical Pacific Ocean from photographic and video survey (Amon et al. 2016a, Amon et al. 2016b, Dahlgren et al. 2016, Foell and Pawson 1986, Glover et al. 2016, Martinez-Arbizu et al. 2013, Pawson 1983, Pawson and Foell 1986, Roux and Pawson 1999, Tilot 2006, Vanreusel et al. 2016). In order to predict and manage the environmental impacts of mining in the CCZ and within the UK-1 exploration contract area, baseline knowledge of the megafauna is essential and allows for a complete taxonomic and biogeographic synthesis of the fauna of the CCZ (Wedding et al. 2015).

Here, we present the first section (Echinodermata) of an image atlas of benthic megafauna that inhabit the UK-1 exploration contract area based on ROV and AUV surveys and samples collected during two cruises of the ABYSSLINE project. This section will be supported by the following sections in the near future: 1) Cnidaria, 2) Porifera, and 3) All Other Phyla (Annelida, Arthropoda, Bryozoa, Chordata, Ctenophora, and Mollusca). This atlas was crucial during the ABYSSLINE quantitative megafaunal analyses (Amon et al. 2016b) and we hope that it will facilitate the standardization of the putative morphospecies and be useful to other scientists and stakeholders undertaking research in the CCZ in the future.

Materials and methods

The UKSRL exploration contract area (UK-1) is located in the eastern CCZ in the Pacific Ocean (Fig. 1). There have been two ABYSSLINE research cruises to the UK-1 exploration contract area: the AB01 or MV1313 cruise on the R/V Melville from 3 to 27 October 2013, and the AB02 or TN319 cruise on the R/V Thompson from 12 February to 25 March 2015. The AB01 cruise focused on a 30x30-km stratum (UK-1 Stratum A) centered at 13°49'N, 116°36'W in the northern portion of the UK-1 contract area (Fig. 1). During the AB01 cruise, multibeam bathymetric surveys indicated an abyssal seafloor characterized by ridges and valleys running from NNW to SSE at 3900–4400 m. The commercial Remotely Operated Vehicle (ROV) Remora III, operated by Phoenix International Holdings, performed video surveys and sample collections at four randomly-located sites within UK-1 Stratum A in the UK-1 contract area. Additionally, surveys were done ~250 km to the east of the UK-1 contract area, at a site called ‘EPIRB’ centered at 13°40'N, 114°24'W (Fig. 1). Work at the EPIRB site was dictated by an emergency response to an Emergency Position Indicating Radio Beacon (EPIRB) distress signal and, although unplanned, provided a useful broader context for our study. The ROV was equipped with two manipulators, four ROS QLEDIII lights, one 1Cam Alpha Component high-definition downward-looking “science” video camera (1080p video and 24.1 megapixel stills) and one standard-definition forward-looking “pilot” video camera. During surveys, the vehicle had substantial difficulty maintaining constant altitude, direction and velocity over the seabed, thereby limiting the availability of usable imagery and also the collection of specimens.

The AB02 cruise focused on a 30x30-km stratum (UK-1 Stratum B) centered at 12°28'N, 116°36'W in the central portion of the UK-1 exploration contract area (Fig. 1). During the AB02 cruise, multibeam bathymetric surveys indicated an abyssal seafloor dominated by numerous high-relief volcanic seamounts between 3500-4300 m. The Autonomous Underwater Vehicle (AUV) REMUS 6000, operated by Woods Hole Oceanographic Institution, performed image surveys at five randomly-located sites within UK-1 Stratum B (Fig. 1). The AUV was equipped with four ROS QLEDIII lights, and one Prosilica GT3400 high-definition downward-looking still camera (9 megapixel stills).

Sample collection

The ROV was the primary tool used to collect specimens on the AB01 cruise, however due to significant difficulties, a limited number of megafauna was successfully sampled. As a result, megafauna that were collected serendipitously by the box corer, megacorer, and Brenke epibenthic sled were also included in this study (Amon et al. 2016b). As there was no ROV on AB02, we again relied on samples collected by chance with the box corer, megacorer, and Brenke epibenthic sled. Once the respective sampling equipment was on deck, megafauna were quickly transferred to containers of chilled seawater, photographed, and a tissue subsample taken for DNA analyses. DNA samples were preserved in 80% ethanol and the remainder of the animal was preserved in buffered 4% formalin-seawater solution or 95% ethanol, depending on the taxon. On board, all collected specimens were also imaged, with the resulting images included in this manuscript. After the cruise, morphological samples were sent to taxonomic experts for identification and all specimens sequenced for a range of DNA markers at the Natural History Museum, London, with tissue samples subsequently archived and made openly-available for future taxonomic work (Dahlgren et al. 2016, Glover et al. 2015, Glover et al. 2016). All collected specimens were used for taxonomic identifications including ground-truthing identifications based on images.

Megafaunal image surveys and analyes

All imagery from both “pilot” and “science” cameras on the ROV (covering roughly 8,000 m2) collected during AB01 was used during the creation of this atlas (Amon et al. 2016b). All imagery from the AUV (27,178 images covering roughly 500,000 m2) collected during AB02 was also used, although the majority of these images (>20,000 images) were at too high an altitude (>6 m) for megafauna to be resolved and identified. All video from both cameras on the ROV, as well as from the AUV, were viewed multiple times and frames archived of each identifiable megafaunal morphotype. The ROV imagery from the AB01 cruise was higher resolution than the imagery collected by the AUV during the AB02 cruise.

The criteria used for selection of megafaunal morphospecies was that individuals were greater than 2 cm in maximum dimension and that there was sufficient detail to identify them to a putative ‘species-level’ morphotype or morphospecies (Amon et al. 2016b). However, this only applied to imagery from AB01 as the AUV imagery collected during AB02 was of poor resolution resulting in only megafauna above 5 cm in the largest dimension being included in this atlas. Morphospecies that could not be identified to species but appeared morphologically distinct were assigned a unique informal species name (e.g. Echinoidea morphospecies 1). These were identified by taxonomic experts or by using the “Atlas of Abyssal Megafauna Morphotypes of the Clarion-Clipperton Fracture Zone” created for the ISA (http://ccfzatlas.com/) (Martinez-Arbizu et al. 2013), as well as Bluhm and Gebruk 1999, Foell and Pawson 1986, Pawson 1983, Pawson and Foell 1986, Roux 2004, Roux and Pawson 1999, Tilot 2006. Morphospecies from this study that matched morphotypes listed in the “Atlas of Abyssal Megafauna Morphotypes of the Clarion-Clipperton Fracture Zone” have had a section titled "Nomenclature" added to their data, in which the identification from the “Atlas of Abyssal Megafauna Morphotypes of the Clarion-Clipperton Fracture Zone” has been included. This is in an effort to provide coherence between these CCZ atlases. For morphospecies that were morphologically similar to a well-defined species name, we use the open nomenclature expression ”cf.", although a precautionary approach was taken. Specimens in this atlas that were collected have undergone (relevant GenBank numbers are included) or are currently undergoing molecular analyses (Glover et al. 2016; Glover et al., unpublished data).

This process provided an estimate of the number of echinoderm morphospecies in the UK-1 contract area and eastern CCZ, and will aid in delimiting species ranges in the CCZ. However, since the majority of the morphospecies were not collected, it is impossible to confirm species identities in most cases or undertake systematic studies on this fauna (Amon et al. 2016b).

Echinoderms of the UKSRL exploration contract area (UK-1) and the eastern Clarion-Clipperton Zone

Phylum Echinodermata

Material    Download as CSV 

Class Asteroidea de Blainville, 1830

Material    Download as CSV 

Order Brisingida Fisher, 1928

Material    Download as CSV 

Family Freyellidae Downey, 1986

Material    Download as CSV 

Genus Freyella Perrier, 1885

Material    Download as CSV 

cf. Freyella morphospecies

Materials    Download as CSV 
  1. scientificName:
    Freyella sp.
    ; taxonConceptID:
    cf. Freyella morphospecies
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Asteroidea
    ; order:
    Brisingida
    ; family:
    Freyellidae
    ; taxonRank:
    genus
    ; genus:
    Freyella
    ; scientificNameAuthorship:
    Perrier, 1885
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum A
    ; maximumDepthInMeters:
    4020
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.8551
    ; decimalLongitude:
    -116.5477
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-21
    ; eventTime:
    3:09
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 6 (RV06)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    Frequently observed on sponge stalks, rocks and seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Christopher Mah, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
  2. scientificName:
    Freyella sp.
    ; taxonConceptID:
    cf. Freyella morphospecies
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Asteroidea
    ; order:
    Brisingida
    ; family:
    Freyellidae
    ; taxonRank:
    genus
    ; genus:
    Freyella
    ; scientificNameAuthorship:
    Perrier, 1885
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum A
    ; maximumDepthInMeters:
    4050
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.9588
    ; decimalLongitude:
    -116.5605
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-16
    ; eventTime:
    2:20
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 3 (RV03)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    Frequently observed on sponge stalks, rocks and seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Christopher Mah, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
  3. scientificName:
    Freyella sp.
    ; taxonConceptID:
    cf. Freyella morphospecies
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Asteroidea
    ; order:
    Brisingida
    ; family:
    Freyellidae
    ; taxonRank:
    genus
    ; genus:
    Freyella
    ; scientificNameAuthorship:
    Perrier, 1885
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    Eastern Clarion-Clipperton Zone
    ; verbatimLocality:
    Site EPIRB
    ; maximumDepthInMeters:
    3943
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.6794
    ; decimalLongitude:
    -114.4133
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-23
    ; eventTime:
    10:03
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 7 (RV07)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    Frequently observed on sponge stalks, rocks and seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Christopher Mah, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
  4. scientificName:
    Freyella sp.
    ; taxonConceptID:
    cf. Freyella morphospecies
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Asteroidea
    ; order:
    Brisingida
    ; family:
    Freyellidae
    ; taxonRank:
    genus
    ; genus:
    Freyella
    ; scientificNameAuthorship:
    Perrier, 1885
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum B
    ; maximumDepthInMeters:
    4250
    ; locationRemarks:
    RV Thompson Cruise TN319
    ; decimalLatitude:
    12.5011
    ; decimalLongitude:
    -116.6442
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Autonomous Underwater Vehicle
    ; eventDate:
    2015-03-18
    ; eventTime:
    11:08
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 9 (AV09)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    Frequently observed on sponge stalks, rocks and seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; identifiedBy:
    Christopher Mah, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2015
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 2

Figure 2.

cf. Freyella morphospecies observed in the UK-1 exploration contract area and eastern CCZ. Images (a-d) correspond with the relevant data above.

acf. Freyella morphospecies attached to dead sponge stalk. Scale bar is 10 cm. Image attribution: DJ Amon & CR Smith, University of Hawai’i.
bcf. Freyella morphospecies attached to dead sponge stalk. Scale bar is 10 cm. Image attribution: DJ Amon & CR Smith, University of Hawai’i.
ccf. Freyella morphospecies attached to a live sponge. Scale bar is 10 cm. Image attribution: DJ Amon & CR Smith, University of Hawai’i.
dcf. Freyella morphospecies in situ on seafloor. Image attribution: Woods Hole Oceanographic Institution.

Genus Freyastera Sladen, 1889

Material    Download as CSV 

Freyastera cf. benthophila Sladen, 1889

Nomenclature: 

In the “Atlas of Abyssal Megafauna Morphotypes of the Clarion-Clipperton Fracture Zone” created for the ISA (http://ccfzatlas.com/), this species is listed as "Freyastera sp. morphotype".

Materials    Download as CSV 
  1. scientificName:
    Freyastera benthophila
    ; taxonConceptID:
    Freyastera cf. benthophila
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Asteroidea
    ; order:
    Brisingida
    ; family:
    Freyellidae
    ; taxonRank:
    species
    ; genus:
    Freyastera
    ; scientificNameAuthorship:
    Sladen, 1889
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum A
    ; maximumDepthInMeters:
    4064
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.8635
    ; decimalLongitude:
    -116.5486
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-21
    ; eventTime:
    5:07
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 6 (RV06)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    Frequently observed on sponge stalks, rocks and seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Christopher Mah, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
  2. scientificName:
    Freyastera benthophila
    ; taxonConceptID:
    Freyastera cf. benthophila
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Asteroidea
    ; order:
    Brisingida
    ; family:
    Freyellidae
    ; taxonRank:
    species
    ; genus:
    Freyastera
    ; scientificNameAuthorship:
    Sladen, 1889
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum B
    ; maximumDepthInMeters:
    4251
    ; locationRemarks:
    RV Thompson Cruise TN319
    ; decimalLatitude:
    12.4979
    ; decimalLongitude:
    -116.6464
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Autonomous Underwater Vehicle
    ; eventDate:
    2015-03-03
    ; eventTime:
    22:38
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 5 (AV05)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    Frequently observed on sponge stalks, rocks and seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; identifiedBy:
    Christopher Mah, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2015
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
  3. scientificName:
    Freyastera benthophila
    ; taxonConceptID:
    Freyastera cf. benthophila
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Asteroidea
    ; order:
    Brisingida
    ; family:
    Freyellidae
    ; taxonRank:
    species
    ; genus:
    Freyastera
    ; scientificNameAuthorship:
    Sladen, 1889
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum A
    ; maximumDepthInMeters:
    4027
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.8609
    ; decimalLongitude:
    -116.5468
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-21
    ; eventTime:
    1:55
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 6 (RV06)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    Frequently observed on sponge stalks, rocks and seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Christopher Mah, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
  4. scientificName:
    Freyastera benthophila
    ; taxonConceptID:
    Freyastera cf. benthophila
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Asteroidea
    ; order:
    Brisingida
    ; family:
    Freyellidae
    ; taxonRank:
    species
    ; genus:
    Freyastera
    ; scientificNameAuthorship:
    Sladen, 1889
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum A
    ; maximumDepthInMeters:
    4011
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.8622
    ; decimalLongitude:
    -116.5462
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-21
    ; eventTime:
    0:39
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 6 (RV06)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    tissue and DNA voucher stored in 80% non-denatured ethanol aqueous solution and remainder of animal preserved in 4% formaldehyde
    ; behavior:
    Frequently observed on sponge stalks, rocks and seafloor
    ; catalogNumber:
    AB01-RV06-CS-10
    ; recordNumber:
    AB01-RV06-CS-10; NHM413
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; otherCatalogNumbers:
    b7ffe7a2-7be1-4d4fb784-7aaecf0ee743; 5023520
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492 | Glover AG, Wiklund H, Rabone M, Amon DJ, Smith CR, O'Hara T, Mah CL, Dahlgren TG. Abyssal fauna of the UK-1 polymetallic nodule exploration claim, Clarion-Clipperton Zone, central Pacific Ocean: Echinodermata. Biodiversity data journal. 2016(4). doi: 10.3897/BDJ.4.e7251
    ; associatedSequences: ; identifiedBy:
    Christopher Mah, Diva J Amon, Amanda F Ziegler, Adrian Glover, Helena Wiklund, Thomas Dahlgren
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified by morphology and DNA of collected specimen
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    PreservedSpecimen
Notes: 

Fig. 3

Figure 3.

Freyastera cf. benthophila observed in the UK-1 exploration contract area. Images (a-d) correspond with the relevant data above.

aFreyastera cf. benthophila in situ on seafloor. Scale bar is 10 cm. Image attribution: DJ Amon & CR Smith, University of Hawai’i.
bFreyastera cf. benthophila attached to a polymetallic nodule on seafloor. Scale bar is 10 cm. Image attribution: Woods Hole Oceanographic Institution.
cFreyastera cf. benthophila attached to dead stalk regenerating sixth arm. Scale bar is 10 cm. Image attribution: DJ Amon & CR Smith, University of Hawai’i.
dTentacle of Freyastera cf. benthophila only partly recovered. Scale bar is 2 cm. Image attribution: AG Glover, TD Dahlgren & H Wiklund, Natural History Museum, London & Uni Research.

Order Paxillosida Perrier, 1884

Material    Download as CSV 

cf. Paxillosida morphospecies 1

Material    Download as CSV 
  1. scientificName:
    Paxillosida sp.
    ; taxonConceptID:
    cf. Paxillosida morphospecies 1
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Asteroidea
    ; order:
    Paxillosida
    ; taxonRank:
    order
    ; scientificNameAuthorship:
    Perrier, 1884
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum A
    ; maximumDepthInMeters:
    4022
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.8593
    ; decimalLongitude:
    -116.5482
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-21
    ; eventTime:
    4:12
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 6 (RV06)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Christopher Mah, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 4

Figure 4.

cf. Paxillosida morphospecies 1 in situ on seafloor in the UK-1 exploration contract area. Image corresponds with the data above. Scale bar is 10 cm. Image attribution: DJ Amon & CR Smith, University of Hawai’i.

cf. Paxillosida morphospecies 2

Material    Download as CSV 
  1. scientificName:
    Paxillosida sp.
    ; taxonConceptID:
    cf. Paxillosida morphospecies 2
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Asteroidea
    ; order:
    Paxillosida
    ; taxonRank:
    order
    ; scientificNameAuthorship:
    Perrier, 1884
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum B
    ; maximumDepthInMeters:
    4144
    ; locationRemarks:
    RV Thompson Cruise TN319
    ; decimalLatitude:
    12.3606
    ; decimalLongitude:
    -116.5133
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Autonomous Underwater Vehicle
    ; eventDate:
    2015-02-18
    ; eventTime:
    20:09
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 1 (AV01)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; identifiedBy:
    Christopher Mah, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2015
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 5

Figure 5.

cf. Paxillosida morphospecies 2 in situ on seafloor in the UK-1 exploration contract area. Image corresponds with the data above. Scale bar is 10 cm. Image attribution: Woods Hole Oceanographic Institution.

cf. Paxillosida morphospecies 3

Nomenclature: 

In the “Atlas of Abyssal Megafauna Morphotypes of the Clarion-Clipperton Fracture Zone” created for the ISA (http://ccfzatlas.com/), this morphospecies is listed as "Astropectinidae Dytaster morphotype".

Material    Download as CSV 
  1. scientificName:
    Paxillosida sp.
    ; taxonConceptID:
    cf. Paxillosida morphospecies 3
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Asteroidea
    ; order:
    Paxillosida
    ; taxonRank:
    order
    ; scientificNameAuthorship:
    Perrier, 1884
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum B
    ; maximumDepthInMeters:
    4159
    ; locationRemarks:
    RV Thompson Cruise TN319
    ; decimalLatitude:
    12.3655
    ; decimalLongitude:
    -116.5184
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Autonomous Underwater Vehicle
    ; eventDate:
    2015-02-18
    ; eventTime:
    20:56
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 1 (AV01)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; identifiedBy:
    Christopher Mah, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2015
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 6

Figure 6.

cf. Paxillosida morphospecies 3 observed in situ on sponge on seafloor in the UK-1 exploration contract area. Image corresponds with the data above. Scale bar is 10 cm. Image attribution: Woods Hole Oceanographic Institution.

Family Porcellanasteridae Sladen, 1883

Material    Download as CSV 

cf. Porcellanasteridae morphospecies

Materials    Download as CSV 
  1. scientificName:
    Porcellanasteridae sp.
    ; taxonConceptID:
    cf. Porcellanasteridae morphospecies
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Asteroidea
    ; order:
    Paxillosida
    ; family:
    Porcellanasteridae
    ; taxonRank:
    family
    ; scientificNameAuthorship:
    Sladen, 1883
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum A
    ; maximumDepthInMeters:
    4029
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.8632
    ; decimalLongitude:
    -116.5464
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-21
    ; eventTime:
    9:11
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 6 (RV06)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Christopher Mah, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
  2. scientificName:
    Porcellanasteridae sp.
    ; taxonConceptID:
    cf. Porcellanasteridae morphospecies
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Asteroidea
    ; order:
    Paxillosida
    ; family:
    Porcellanasteridae
    ; taxonRank:
    family
    ; scientificNameAuthorship:
    Sladen, 1883
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    Eastern Clarion-Clipperton Zone
    ; verbatimLocality:
    Site EPIRB
    ; maximumDepthInMeters:
    3950
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.6798
    ; decimalLongitude:
    -114.4144
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-23
    ; eventTime:
    13:27
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 7 (RV07)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Christopher Mah, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 7

Figure 7.

cf. Porcellanasteridae morphospecies observed in the UK-1 exploration contract area and eastern CCZ. Images (a-b) correspond with the data above.

acf. Porcellanasteridae morphospecies in situ on seafloor. Image attribution: DJ Amon & CR Smith, University of Hawai’i.
bcf. Porcellanasteridae morphospecies in situ on seafloor. Scale bar is 10 cm. Image attribution: DJ Amon & CR Smith, University of Hawai’i.

Genus Porcellanaster Wyville Thomson, 1877

Material    Download as CSV 

cf. Porcellanaster morphospecies

Materials    Download as CSV 
  1. scientificName:
    Porcellanaster sp.
    ; taxonConceptID:
    cf. Porcellanaster morphospecies
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Asteroidea
    ; order:
    Paxillosida
    ; family:
    Porcellanasteridae
    ; taxonRank:
    genus
    ; genus:
    Porcellanaster
    ; scientificNameAuthorship:
    Wyville Thomson, 1877
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum A
    ; maximumDepthInMeters:
    4110
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.8501
    ; decimalLongitude:
    -116.6456
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-10
    ; eventTime:
    16:24
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 1 (RV01)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Christopher Mah, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
  2. scientificName:
    Porcellanaster sp.
    ; taxonConceptID:
    cf. Porcellanaster morphospecies
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Asteroidea
    ; order:
    Paxillosida
    ; family:
    Porcellanasteridae
    ; taxonRank:
    genus
    ; genus:
    Porcellanaster
    ; scientificNameAuthorship:
    Wyville Thomson, 1877
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    Eastern Clarion-Clipperton Zone
    ; verbatimLocality:
    Site EPIRB
    ; maximumDepthInMeters:
    3947
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.6794
    ; decimalLongitude:
    -114.4137
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-23
    ; eventTime:
    9:58
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 7 (RV07)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Christopher Mah, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 8

Figure 8.

cf. Porcellanaster morphospecies observed in the UK-1 exploration contract area and eastern CCZ. Images (a-b) correspond with the data above.

acf. Porcellanaster morphospecies in situ on seafloor. Image attribution: DJ Amon & CR Smith, University of Hawai’i.
bcf. Porcellanaster morphospecies in situ on seafloor. Image attribution: DJ Amon & CR Smith, University of Hawai’i.

Porcellanaster ceruleus Wyville Thomson, 1877

Material    Download as CSV 
  1. scientificName:
    Porcellanaster ceruleus
    ; taxonConceptID:
    Porcellanaster ceruleus
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Asteroidea
    ; order:
    Paxillosida
    ; family:
    Porcellanasteridae
    ; taxonRank:
    species
    ; genus:
    Porcellanaster
    ; scientificNameAuthorship:
    Wyville Thomson, 1877
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum A
    ; maximumDepthInMeters:
    4076
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.7558
    ; decimalLongitude:
    -116.4867
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    500
    ; samplingProtocol:
    Brenke Epibenthic Sled
    ; eventDate:
    2013-10-17
    ; eventTime:
    1:50
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Brenke Epibenthic Sled (EB04)
    ; individualCount:
    2
    ; lifeStage:
    Adult
    ; preparations:
    tissue and DNA voucher stored in 80% non-denatured ethanol aqueous solution and remainder of animal preserved in 4% formaldehyde
    ; behavior:
    On seafloor
    ; catalogNumber:
    AB01-EB04-NHM253
    ; recordNumber:
    AB01-EB04-NHM253
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; otherCatalogNumbers:
    95d0bd7f-0df9-47e4-8003-cd12007d54b4
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492 | Glover AG, Wiklund H, Rabone M, Amon DJ, Smith CR, O'Hara T, Mah CL, Dahlgren TG. Abyssal fauna of the UK-1 polymetallic nodule exploration claim, Clarion-Clipperton Zone, central Pacific Ocean: Echinodermata. Biodiversity data journal. 2016(4). doi: 10.3897/BDJ.4.e7251
    ; associatedSequences: ; identifiedBy:
    Christopher Mah, Diva J Amon, Amanda F Ziegler, Adrian Glover, Helena Wiklund, Thomas Dahlgren
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified by morphology and DNA of collected specimen
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    NHMUK
    ; collectionCode:
    ZOO
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    PreservedSpecimen
Notes: 

Fig. 9

Figure 9.

Porcellanaster ceruleus after collection from the UK-1 exploration contract area. Aboral view on left and oral view on right. Image corresponds with the data above. Image attribution: AG Glover, TD Dahlgren & H Wiklund, Natural History Museum, London & Uni Research.

Genus Styracaster Sladen, 1883

Material    Download as CSV 

Styracaster paucispinus Ludwig, 1907

Material    Download as CSV 
  1. scientificName:
    Syracaster paucispinus
    ; taxonConceptID:
    Syracaster paucispinus
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Asteroidea
    ; order:
    Paxillosida
    ; family:
    Porcellanasteridae
    ; taxonRank:
    species
    ; genus:
    Styracaster
    ; scientificNameAuthorship:
    Ludwig, 1907
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum B
    ; maximumDepthInMeters:
    4237
    ; locationRemarks:
    RV Thompson Cruise TN319
    ; decimalLatitude:
    12.5212
    ; decimalLongitude:
    -116.698
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    50
    ; samplingProtocol:
    Megacorer
    ; eventDate:
    2015-03-09
    ; eventTime:
    10:56
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Megacorer 17 (MC17)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    tissue and DNA voucher stored in 80% non-denatured ethanol aqueous solution and remainder of animal preserved in 4% formaldehyde
    ; behavior:
    On seafloor
    ; catalogNumber:
    AB02-MC17-CS-31
    ; recordNumber:
    AB02-MC17-CS-31; NHM1608
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492 | Glover AG, Wiklund H, Rabone M, Amon DJ, Smith CR, O'Hara T, Mah CL, Dahlgren TG. Abyssal fauna of the UK-1 polymetallic nodule exploration claim, Clarion-Clipperton Zone, central Pacific Ocean: Echinodermata. Biodiversity data journal. 2016(4). doi: 10.3897/BDJ.4.e7251
    ; identifiedBy:
    Christopher Mah, Diva J Amon, Amanda F Ziegler, Adrian Glover, Helena Wiklund, Thomas Dahlgren
    ; dateIdentified:
    2015
    ; identificationRemarks:
    Identified by morphology and DNA of collected specimen
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    PreservedSpecimen
Notes: 

Fig. 10

Figure 10.

Styracaster paucispinus after collection from the UK-1 exploration contract area. Images (a) and (b) correspond with the data in (a) above.

aAboral view of Styracaster paucispinus after collection. The pink dots on the specimen are parasites. Scale bar is 1 cm. Image attribution: DJ Amon and CR Smith.
bOral view of Styracaster paucispinus after collection. Scale bar is 1 cm. Image attribution: DJ Amon and CR Smith.

Order Velatida Perrier, 1884

Material    Download as CSV 

Family Pterasteridae Perrier, 1875

Material    Download as CSV 

cf. Pterasteridae morphospecies 1

Nomenclature: 

In the “Atlas of Abyssal Megafauna Morphotypes of the Clarion-Clipperton Fracture Zone” created for the ISA (http://ccfzatlas.com/), this morphospecies is listed as "Pteraster morphotype".

Materials    Download as CSV 
  1. scientificName:
    Pterasteridae sp.
    ; taxonConceptID:
    cf. Pterasteridae morphospecies 1
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Asteroidea
    ; order:
    Velatida
    ; family:
    Pterasteridae
    ; taxonRank:
    family
    ; scientificNameAuthorship:
    Perrier, 1875
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum A
    ; maximumDepthInMeters:
    4059
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.9669
    ; decimalLongitude:
    -116.55796
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-16
    ; eventTime:
    0:06
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 3 (RV03)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Christopher Mah, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
  2. scientificName:
    Pterasteridae sp.
    ; taxonConceptID:
    cf. Pterasteridae morphospecies 1
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Asteroidea
    ; order:
    Velatida
    ; family:
    Pterasteridae
    ; taxonRank:
    family
    ; scientificNameAuthorship:
    Perrier, 1875
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum B
    ; maximumDepthInMeters:
    4226
    ; locationRemarks:
    RV Thompson Cruise TN319
    ; decimalLatitude:
    12.5783
    ; decimalLongitude:
    -116.6872
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Autonomous Underwater Vehicle
    ; eventDate:
    2015-03-09
    ; eventTime:
    3:12
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 6 (AV06)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Christopher Mah, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2015
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 11

Figure 11.

cf. Pterasteridae morphospecies 1 observed in the UK-1 exploration contract area. Images correspond with the data above.

acf. Pterasteridae morphospecies 1 in situ on seafloor. Scale bar is 10 cm. Image attribution: DJ Amon & CR Smith, University of Hawai’i.
bcf. Pterasteridae morphospecies 1 in situ on seafloor. Scale bar is 10 cm. Image attribution: Woods Hole Oceanographic Institution.

cf. Pterasteridae morphospecies 2

Material    Download as CSV 
  1. scientificName:
    Pterasteridae sp.
    ; taxonConceptID:
    cf. Pterasteridae morphospecies 2
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Asteroidea
    ; order:
    Velatida
    ; family:
    Pterasteridae
    ; taxonRank:
    family
    ; scientificNameAuthorship:
    Perrier, 1875
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum A
    ; maximumDepthInMeters:
    4107
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.85096
    ; decimalLongitude:
    -116.6453
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-10
    ; eventTime:
    16:48
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 1 (RV01)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Christopher Mah, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 12

Figure 12.

cf. Pterasteridae morphospecies 2 observed in the UK-1 exploration contract area. Image corresponds with the data above. Image attribution: DJ Amon & CR Smith, University of Hawai’i.

Genus Hymenaster Thomson, 1873

Material    Download as CSV 

cf. Hymenaster morphospecies 1

Material    Download as CSV 
  1. scientificName:
    Hymenaster sp.
    ; taxonConceptID:
    cf. Hymenaster morphospecies 1
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Asteroidea
    ; order:
    Velatida
    ; family:
    Pterasteridae
    ; taxonRank:
    genus
    ; genus:
    Hymenaster
    ; scientificNameAuthorship:
    Thomson, 1873
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum A
    ; maximumDepthInMeters:
    4027
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.86097
    ; decimalLongitude:
    -116.5468
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-21
    ; eventTime:
    1:54
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 6 (RV06)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Christopher Mah, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 13

Figure 13.

cf. Hymenaster morphospecies 1 in situ on seafloor in the UK-1 exploration contract area. Image corresponds with the data above. Scale bar is 10 cm. Image attribution: DJ Amon & CR Smith, University of Hawai’i.

cf. Hymenaster morphospecies 2

Material    Download as CSV 
  1. scientificName:
    Hymenaster sp.
    ; taxonConceptID:
    cf. Hymenaster morphospecies 2
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Asteroidea
    ; order:
    Velatida
    ; family:
    Pterasteridae
    ; taxonRank:
    genus
    ; genus:
    Hymenaster
    ; scientificNameAuthorship:
    Thomson, 1873
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum A
    ; maximumDepthInMeters:
    4022
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.8577
    ; decimalLongitude:
    -116.5479
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-21
    ; eventTime:
    3:46
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 6 (RV06)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Christopher Mah, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 14

Figure 14.

cf. Hymenaster morphospecies 2 observed in the UK-1 exploration contract area. Both image (a) and image (b) correspond with the data in (a) above.

aSide view of cf. Hymenaster morphospecies 2 in situ on seafloor. Scale bar is 10 cm. Image attribution: DJ Amon & CR Smith, University of Hawai’i.
bAboral view of cf. Hymenaster morphospecies 2 in situ on seafloor. Scale bar is 10 cm. Image attribution: DJ Amon & CR Smith, University of Hawai’i.

Class Crinoidea

Material    Download as CSV 

Order Comatulida Sieverts-Doreck, 1953

Material    Download as CSV 

cf. Comatulida morphospecies 1

Nomenclature: 

In the “Atlas of Abyssal Megafauna Morphotypes of the Clarion-Clipperton Fracture Zone” created for the ISA (http://ccfzatlas.com/), this morphospecies is listed as "Featherstar sp. 1".

Materials    Download as CSV 
  1. scientificName:
    Comatulida sp.
    ; taxonConceptID:
    cf. Comatulida morphospecies 1
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Crinoidea
    ; order:
    Comatulida
    ; taxonRank:
    order
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    Eastern Clarion-Clipperton Zone
    ; verbatimLocality:
    Site EPIRB
    ; maximumDepthInMeters:
    3914
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.6793
    ; decimalLongitude:
    -114.4072
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-23
    ; eventTime:
    11:52
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 7 (RV07)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On nodule
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Michel Roux, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
  2. scientificName:
    Comatulida sp.
    ; taxonConceptID:
    cf. Comatulida morphospecies 1
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Crinoidea
    ; order:
    Comatulida
    ; taxonRank:
    order
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum A
    ; maximumDepthInMeters:
    4025
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.8639
    ; decimalLongitude:
    -116.5487
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-21
    ; eventTime:
    5:14
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 6 (RV06)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On nodule
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Michel Roux, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 15

Figure 15.

cf. Comatulida morphospecies 1 observed in the UK-1 exploration contract area and eastern CCZ. Images (a-b) correspond with the data above.

acf. Comatulida morphospecies 1 in situ on seafloor. Scale bar is 10 cm. Image attribution: DJ Amon & CR Smith, University of Hawai’i.
bcf. Comatulida morphospecies 1 in situ on seafloor. Scale bar is 10 cm. Image attribution: DJ Amon & CR Smith, University of Hawai’i.

cf. Comatulida morphospecies 2

Nomenclature: 

In the “Atlas of Abyssal Megafauna Morphotypes of the Clarion-Clipperton Fracture Zone” created for the ISA (http://ccfzatlas.com/), this morphospecies is listed as "Pentametrocrinus sp. 1".

Material    Download as CSV 
  1. scientificName:
    Comatulida sp.
    ; taxonConceptID:
    cf. Comatulida morphospecies 2
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Crinoidea
    ; order:
    Comatulida
    ; taxonRank:
    order
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum A
    ; maximumDepthInMeters:
    4110
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.8498
    ; decimalLongitude:
    -116.6458
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-10
    ; eventTime:
    13:48
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 1 (RV01)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On nodule
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Michel Roux, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 16

Figure 16.

cf. Comatulida morphospecies 2 observed in the UK-1 exploration contract area. Image (a-b) correspond with the data in (a) above.

acf. Comatulida morphospecies 2 in situ on seafloor. Image attribution: DJ Amon & CR Smith, University of Hawai’i.
bcf. Comatulida morphospecies 2 in situ on seafloor. Image attribution: DJ Amon & CR Smith, University of Hawai’i.

Family Bathycrinidae Bather, 1899

Material    Download as CSV 

Genus Bathycrinus Thomson, 1872

Material    Download as CSV 

Bathycrinus cf. equatorialis A.H. Clark, 1908

Nomenclature: 

In the “Atlas of Abyssal Megafauna Morphotypes of the Clarion-Clipperton Fracture Zone” created for the ISA (http://ccfzatlas.com/), this morphospecies is listed as "Bourgueticrinina sp. 1".

Materials    Download as CSV 
  1. scientificName:
    Bathycrinus equatorialis
    ; taxonConceptID:
    Bathycrinus cf. equatorialis
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Crinoidea
    ; order:
    Comatulida
    ; family:
    Bathycrinidae
    ; taxonRank:
    species
    ; genus:
    Bathycrinus
    ; scientificNameAuthorship:
    AH Clark, 1908
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum A
    ; maximumDepthInMeters:
    4071
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.7605
    ; decimalLongitude:
    -116.468
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-18
    ; eventTime:
    2:14
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 5 (RV05)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On nodule
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492.
    ; identifiedBy:
    Michel Roux, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
  2. scientificName:
    Bathycrinus equatorialis
    ; taxonConceptID:
    Bathycrinus cf. equatorialis
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Crinoidea
    ; order:
    Comatulida
    ; family:
    Bathycrinidae
    ; taxonRank:
    species
    ; genus:
    Bathycrinus
    ; scientificNameAuthorship:
    AH Clark, 1908
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    Eastern Clarion-Clipperton Zone
    ; verbatimLocality:
    Site EPIRB
    ; maximumDepthInMeters:
    3909
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.6785
    ; decimalLongitude:
    -114.406
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-23
    ; eventTime:
    11:21
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 7 (RV07)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On nodule
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492.
    ; identifiedBy:
    Michel Roux, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 17

Figure 17.

Bathycrinus cf. equatorialis observed in the UK-1 exploration contract area and eastern CCZ. Images (a-b) correspond with the data above.

aBathycrinus cf. equatorialis in situ on seafloor. Image attribution: DJ Amon & CR Smith, University of Hawai’i.
bBathycrinus cf. equatorialis in situ on seafloor. Scale bar is 10 cm. Image attribution: DJ Amon & CR Smith, University of Hawai’i.

Order Hyocrinida Rasmussen, 1978

Material    Download as CSV 

Family Hyocrinidae Carpenter, 1884

Material    Download as CSV 

cf. Hyocrinidae morphospecies

Nomenclature: 

In the “Atlas of Abyssal Megafauna Morphotypes of the Clarion-Clipperton Fracture Zone” created for the ISA (http://ccfzatlas.com/), this morphospecies is listed as "Bourgueticrinina sp. 1".

Material    Download as CSV 
  1. scientificName:
    Hyocrinidae sp.
    ; taxonConceptID:
    cf. Hyocrinidae morphospecies
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Crinoidea
    ; order:
    Hyocrinida
    ; family:
    Hyocrinidae
    ; taxonRank:
    family
    ; scientificNameAuthorship:
    Carpenter, 1884
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum A
    ; maximumDepthInMeters:
    3919
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.6787
    ; decimalLongitude:
    -114.4072
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-23
    ; eventTime:
    11:09
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 7 (RV07)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On nodule
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Michel Roux, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 18

Figure 18.

cf. Hyocrinidae morphospecies in situ on seafloor in the eastern CCZ. Image corresponds with the data above. Scale bar is 10 cm. Image attribution: DJ Amon & CR Smith, University of Hawai’i.

Genus Hyocrinus Thomson, 1876

Material    Download as CSV 

Hyocrinus cf. foelli Roux & Pawson, 1999

Nomenclature: 

In the “Atlas of Abyssal Megafauna Morphotypes of the Clarion-Clipperton Fracture Zone” created for the ISA (http://ccfzatlas.com/), this morphospecies is listed as "Hyocrinidae sp. 1".

Materials    Download as CSV 
  1. scientificName:
    Hyocrinus foelli
    ; taxonConceptID:
    Hyocrinus cf. foelli
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Crinoidea
    ; order:
    Hyocrinida
    ; family:
    Hyocrinidae
    ; taxonRank:
    species
    ; genus:
    Hyocrinus
    ; scientificNameAuthorship:
    Roux & Pawson, 1999
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    Eastern Clarion-Clipperton Zone
    ; verbatimLocality:
    Site EPIRB
    ; maximumDepthInMeters:
    3944
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.6794
    ; decimalLongitude:
    -114.41297
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-23
    ; eventTime:
    10:06
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 7 (RV07)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On nodule
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016, 6. doi: 10.1038/srep30492.
    ; identifiedBy:
    Michel Roux, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationReferences:
    Roux M & Pawson DL. Two New Pacific Ocean Species of Hyocrinid Crinoids (Echinodermata),with Comments on Presumed Giant-Dwarf Gradients Related to Seamountsand Abyssal Plains. Pacific Science. 1999, 53(3), 289-298; Roux M (2004). New hyocrinid crinoids (Echinodermata) from submersible investigations in the Pacific Ocean. Pacific Science, 58:597-613.
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
  2. scientificName:
    Hyocrinus foelli
    ; taxonConceptID:
    Hyocrinus cf. foelli
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Crinoidea
    ; order:
    Hyocrinida
    ; family:
    Hyocrinidae
    ; taxonRank:
    species
    ; genus:
    Hyocrinus
    ; scientificNameAuthorship:
    Roux & Pawson, 1999
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    Eastern Clarion-Clipperton Zone
    ; verbatimLocality:
    Site EPIRB
    ; maximumDepthInMeters:
    3919
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.6792
    ; decimalLongitude:
    -114.4079
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-23
    ; eventTime:
    11:59
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 7 (RV07)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On nodule
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016, 6. doi: 10.1038/srep30492
    ; identifiedBy:
    Michel Roux, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationReferences:
    Roux M & Pawson DL. Two New Pacific Ocean Species of Hyocrinid Crinoids (Echinodermata),with Comments on Presumed Giant-Dwarf Gradients Related to Seamountsand Abyssal Plains. Pacific Science. 1999, 53(3), 289-298; Roux M (2004). New hyocrinid crinoids (Echinodermata) from submersible investigations in the Pacific Ocean. Pacific Science, 58:597-613.
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 19

Figure 19.

Hyocrinus cf. foelli observed in the eastern CCZ. Images (a-b) correspond with the data above.

aHyocrinus cf. foelli in situ on seafloor. Scale bar is 10 cm. Image attribution: DJ Amon & CR Smith, University of Hawai’i.
bHyocrinus cf. foelli in situ on seafloor. Scale bar is 10 cm. Image attribution: DJ Amon & CR Smith, University of Hawai’i.

Class Echinoidea Leske, 1778

Material    Download as CSV 

cf. Echinoidea morphospecies 1

Materials    Download as CSV 
  1. scientificName:
    Echinoidea sp.
    ; taxonConceptID:
    cf. Echinoidea morphospecies 1
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Echinoidea
    ; taxonRank:
    class
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum B
    ; maximumDepthInMeters:
    4163
    ; locationRemarks:
    RV Thompson Cruise TN319
    ; decimalLatitude:
    12.369004
    ; decimalLongitude:
    -116.5195
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Autonomous Underwater Vehicle
    ; eventDate:
    2015-02-18
    ; eventTime:
    16:29
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 1 (AV01)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; identifiedBy:
    Richard Mooi, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2015
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
  2. scientificName:
    Echinoidea sp.
    ; taxonConceptID:
    cf. Echinoidea morphospecies 1
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Echinoidea
    ; taxonRank:
    class
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum B
    ; maximumDepthInMeters:
    4253
    ; locationRemarks:
    RV Thompson Cruise TN319
    ; decimalLatitude:
    12.4963
    ; decimalLongitude:
    -116.64899
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Autonomous Underwater Vehicle
    ; eventDate:
    2015-03-04
    ; eventTime:
    0:04
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 5 (AV05)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; identifiedBy:
    Richard Mooi, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2015
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 20

Figure 20.

cf. Echinoidea morphospecies 1 observed in the UK-1 exploration contract area. Images (a-b) correspond with the data above.

acf. Echinoidea morphospecies 1 in situ on seafloor. Scale bar is 10 cm. Image attribution: Woods Hole Oceanographic Institution.
bcf. Echinoidea morphospecies 1 in situ on seafloor. Scale bar is 10 cm. Image attribution: Woods Hole Oceanographic Institution.

cf. Echinoidea morphospecies 2

Material    Download as CSV 
  1. scientificName:
    Echinoidea sp.
    ; taxonConceptID:
    cf. Echinoidea morphospecies 2
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Echinoidea
    ; taxonRank:
    class
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum B
    ; maximumDepthInMeters:
    4251
    ; locationRemarks:
    RV Thompson Cruise TN319
    ; decimalLatitude:
    12.5014
    ; decimalLongitude:
    -116.64696
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Autonomous Underwater Vehicle
    ; eventDate:
    2015-03-03
    ; eventTime:
    21:44
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 5 (AV05)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; identifiedBy:
    Richard Mooi, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2015
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 21

Figure 21.

cf. Echinoidea morphospecies 2 in situ on seafloor in the UK-1 exploration contract area. Image corresponds with the data above. Scale bar is 10 cm. Image attribution: Woods Hole Oceanographic Institution.

Order Aspidodiadematoida Kroh & Smith, 2010

Material    Download as CSV 

Family Aspidodiadematidae Duncan 1889

Material    Download as CSV 

cf. Aspidodiadematidae morphospecies

Materials    Download as CSV 
  1. scientificName:
    Aspidodiadematidae sp.
    ; taxonConceptID:
    cf. Aspidodiadematidae morphospecies
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Echinoidea
    ; order:
    Aspidodiadematoida
    ; family:
    Aspidodiadematidae
    ; taxonRank:
    family
    ; scientificNameAuthorship:
    Duncan 1889
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum B
    ; maximumDepthInMeters:
    4255
    ; locationRemarks:
    RV Thompson Cruise TN319
    ; decimalLatitude:
    12.5025
    ; decimalLongitude:
    -116.6489
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Autonomous Underwater Vehicle
    ; eventDate:
    2015-03-18
    ; eventTime:
    8:35
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 9 (AV09)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; identifiedBy:
    Richard Mooi, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2015
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
  2. scientificName:
    Aspidodiadematidae sp.
    ; taxonConceptID:
    cf. Aspidodiadematidae morphospecies
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Echinoidea
    ; order:
    Aspidodiadematoida
    ; family:
    Aspidodiadematidae
    ; taxonRank:
    family
    ; scientificNameAuthorship:
    Duncan 1889
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum A
    ; maximumDepthInMeters:
    4108
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.8502
    ; decimalLongitude:
    -116.6457
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-10
    ; eventTime:
    12:05
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 1 (RV01)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Richard Mooi, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 22

Figure 22.

cf. Aspidodiadematidae morphospecies observed in the UK-1 exploration contract. Images (a-b) correspond with the data above.

acf. Aspidodiadematidae morphospecies in situ on seafloor. Scale bar is 10 cm. Image attribution: Woods Hole Oceanographic Institution.
bcf. Aspidodiadematidae morphospecies in situ on seafloor. Image attribution: DJ Amon & CR Smith, University of Hawai’i.

Order Holasteroida Durham & Melville, 1957

Material    Download as CSV 

cf. Holasteroida morphospecies 1

Material    Download as CSV 
  1. scientificName:
    Holasteroida sp.
    ; taxonConceptID:
    cf. Holasteroida morphospecies 1
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Echinoidea
    ; order:
    Holasteroida
    ; taxonRank:
    order
    ; scientificNameAuthorship:
    Durham & Melville, 1957
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum B
    ; maximumDepthInMeters:
    4224
    ; locationRemarks:
    RV Thompson Cruise TN319
    ; decimalLatitude:
    12.5797
    ; decimalLongitude:
    -116.7271
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Autonomous Underwater Vehicle
    ; eventDate:
    2015-03-09
    ; eventTime:
    7:56
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 6 (AV06)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; identifiedBy:
    Richard Mooi, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2015
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 23

Figure 23.

cf. Holasteroida morphospecies 1 in situ on seafloor in the UK-1 exploration contract area. Image corresponds with the data above. Scale bar is 10 cm. Image attribution: Woods Hole Oceanographic Institution.

cf. Holasteroida morphospecies 2

Materials    Download as CSV 
  1. scientificName:
    Holasteroida sp.
    ; taxonConceptID:
    cf. Holasteroida morphospecies 2
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Echinoidea
    ; order:
    Holasteroida
    ; taxonRank:
    order
    ; scientificNameAuthorship:
    Durham & Melville, 1957
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum B
    ; maximumDepthInMeters:
    4253
    ; locationRemarks:
    RV Thompson Cruise TN319
    ; decimalLatitude:
    12.4965
    ; decimalLongitude:
    -116.64997
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Autonomous Underwater Vehicle
    ; eventDate:
    2015-03-04
    ; eventTime:
    6:59
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 5 (AV05)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; identifiedBy:
    Richard Mooi, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2015
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
  2. scientificName:
    Holasteroida sp.
    ; taxonConceptID:
    cf. Holasteroida morphospecies 2
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Echinoidea
    ; order:
    Holasteroida
    ; taxonRank:
    order
    ; scientificNameAuthorship:
    Durham & Melville, 1957
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum B
    ; maximumDepthInMeters:
    4227
    ; locationRemarks:
    RV Thompson Cruise TN319
    ; decimalLatitude:
    12.5796
    ; decimalLongitude:
    -116.7233
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Autonomous Underwater Vehicle
    ; eventDate:
    2015-03-09
    ; eventTime:
    11:03
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 6 (AV06)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; identifiedBy:
    Richard Mooi, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2015
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 24

Figure 24.

cf. Holasteroida morphospecies 2 observed in the UK-1 exploration contract area. Images (a-b) correspond with the data above.

acf. Holasteroida morphospecies 2 in situ on seafloor. Scale bar is 10 cm. Image attribution: Woods Hole Oceanographic Institution.
bcf. Holasteroida morphospecies 2 in situ on seafloor. Scale bar is 10 cm. Image attribution: Woods Hole Oceanographic Institution.

Family Pourtalesiidae A. Agassiz, 1881

Material    Download as CSV 

Genus Cystocrepis Mortensen, 1907

Material    Download as CSV 

cf. Cystocrepis morphospecies

Material    Download as CSV 
  1. scientificName:
    Cystocrepis sp.
    ; taxonConceptID:
    cf. Cystocrepis morphospecies
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Echinoidea
    ; order:
    Holasteroida
    ; family:
    Pourtalesiidae
    ; taxonRank:
    genus
    ; genus:
    Cystocrepis
    ; scientificNameAuthorship:
    Mortensen, 1907
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum A
    ; maximumDepthInMeters:
    4022
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.85801
    ; decimalLongitude:
    -116.5473
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-21
    ; eventTime:
    2:33
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 6 (RV06)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Richard Mooi, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 25

Figure 25.

cf. Cystocrepis morphospecies in situ on seafloor in the UK-1 exploration contract area. Image corresponds with the data above. Scale bar is 10 cm. Image attribution: DJ Amon & CR Smith, University of Hawai’i.

Genus Echinocrepis A. Agassiz, 1879

Material    Download as CSV 

cf. Echinocrepis morphospecies

Materials    Download as CSV 
  1. scientificName:
    Echinocrepis sp.
    ; taxonConceptID:
    cf. Echinocrepis morphospecies
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Echinoidea
    ; order:
    Holasteroida
    ; family:
    Pourtalesiidae
    ; taxonRank:
    genus
    ; genus:
    Echinocrepis
    ; scientificNameAuthorship:
    A Agassiz, 1879
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum A
    ; maximumDepthInMeters:
    4026
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.8601
    ; decimalLongitude:
    -116.5484
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-21
    ; eventTime:
    4:22
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 6 (RV06)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Richard Mooi, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
  2. scientificName:
    Echinocrepis sp.
    ; taxonConceptID:
    cf. Echinocrepis morphospecies
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Echinoidea
    ; order:
    Holasteroida
    ; family:
    Pourtalesiidae
    ; taxonRank:
    genus
    ; genus:
    Echinocrepis
    ; scientificNameAuthorship:
    A. Agassiz, 1879
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum B
    ; maximumDepthInMeters:
    4200
    ; locationRemarks:
    RV Thompson Cruise TN319
    ; decimalLatitude:
    12.56795
    ; decimalLongitude:
    -116.7361
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Autonomous Underwater Vehicle
    ; eventDate:
    2015-03-09
    ; eventTime:
    10:36
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 6 (AV06)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; identifiedBy:
    Richard Mooi, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2015
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
  3. scientificName:
    Echinocrepis sp.
    ; taxonConceptID:
    cf. Echinocrepis morphospecies
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Echinoidea
    ; order:
    Holasteroida
    ; family:
    Pourtalesiidae
    ; taxonRank:
    genus
    ; genus:
    Echinocrepis
    ; scientificNameAuthorship:
    A Agassiz, 1879
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum B
    ; maximumDepthInMeters:
    4213
    ; locationRemarks:
    RV Thompson Cruise TN319
    ; decimalLatitude:
    12.5707
    ; decimalLongitude:
    -116.7072
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Autonomous Underwater Vehicle
    ; eventDate:
    2015-03-09
    ; eventTime:
    15:06
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 6 (AV06)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; identifiedBy:
    Richard Mooi, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2015
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 26

Figure 26.

Echinocrepis morphospecies observed in the UK-1 exploration contract area. Images (a-c) correspond with the data above.

acf. Echinocrepis morphospecies in situ on seafloor. Scale bar is 10 cm. Image attribution: DJ Amon & CR Smith, University of Hawai’i.
bcf. Echinocrepis morphospecies in situ on seafloor. Scale bar is 10 cm. Image attribution: Woods Hole Oceanographic Institution.
ccf. Echinocrepis morphospecies in situ on seafloor. Scale bar is 10 cm. Image attribution: Woods Hole Oceanographic Institution.

Family Urechinidae Duncan, 1889

Material    Download as CSV 

cf. Urechinidae morphospecies

Material    Download as CSV 
  1. scientificName:
    Urechinidae sp.
    ; taxonConceptID:
    cf. Urechinidae morphospecies
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Echinoidea
    ; order:
    Holasteroida
    ; family:
    Urechinidae
    ; taxonRank:
    family
    ; scientificNameAuthorship:
    Duncan, 1889
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum A
    ; maximumDepthInMeters:
    4020
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.8579
    ; decimalLongitude:
    -116.54799
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-21
    ; eventTime:
    3:50
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 6 (RV06)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Richard Mooi, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 27

Figure 27.

cf. Urechinidae morphospecies in situ on seafloor in the UK-1 exploration contract area. Image corresponds with the data above. Scale bar is 10 cm. Image attribution: DJ Amon & CR Smith, University of Hawai’i.

Order Spatangoida L. Agassiz, 1840

Material    Download as CSV 

Family Schizasteridae Lambert, 1905

Material    Download as CSV 

Genus Aceste Thomson, 1877

Material    Download as CSV 

Aceste cf. ovata A. Agassiz, 1840

Materials    Download as CSV 
  1. scientificName:
    Aceste ovata
    ; taxonConceptID:
    Aceste cf. ovata
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Echinoidea
    ; order:
    Spatangoida
    ; family:
    Schizasteridae
    ; taxonRank:
    species
    ; genus:
    Aceste
    ; scientificNameAuthorship:
    A Agassiz & HL Clark, 1907
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum A
    ; maximumDepthInMeters:
    4028
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.8619
    ; decimalLongitude:
    -116.5484
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-21
    ; eventTime:
    4:47
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 6 (RV06)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Richard Mooi, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
  2. scientificName:
    Aceste ovata
    ; taxonConceptID:
    Aceste ovata
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Echinoidea
    ; order:
    Spatangoida
    ; family:
    Schizasteridae
    ; taxonRank:
    species
    ; genus:
    Aceste
    ; scientificNameAuthorship:
    A Agassiz & HL Clark, 1907
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum B
    ; maximumDepthInMeters:
    4647
    ; locationRemarks:
    RV Thompson Cruise TN319
    ; decimalLatitude:
    12.531
    ; decimalLongitude:
    -116.6233
    ; geodeticDatum:
    WGS85
    ; coordinateUncertaintyInMeters:
    500
    ; samplingProtocol:
    Brenke Epibenthic Sled
    ; eventDate:
    2015-02-20
    ; eventTime:
    22:09
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Brenke Epibenthic Sled (EB02)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    tissue and DNA voucher stored in 80% non-denatured ethanol aqueous solution and remainder of animal preserved in 4% formaldehyde
    ; behavior:
    On seafloor
    ; catalogNumber:
    AB02-EB02-CS-18
    ; recordNumber:
    AB02-EB02-CS-18; NHM1372
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Richard Mooi, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2015
    ; identificationRemarks:
    Identified by morphology and DNA of collected specimen
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    PreservedSpecimen
Notes: 

Fig. 28

Figure 28.

Aceste cf. ovata observed in the UK-1 exploration contract area. Image (a) corresponds with the data in (a) and images (b) and (c) correspond with the data in (b) above.

aAceste ovata in situ on seafloor. Image attribution: DJ Amon and CR Smith.
bAboral view of Aceste ovata after collection. Scale bar is 1 cm. Image attribution: DJ Amon and CR Smith.
cOral view of Aceste ovata after collection. Scale bar is 1 cm. Image attribution: DJ Amon and CR Smith.

Class Holothuroidea

Material    Download as CSV 

Order Aspidochirotida Grube, 1840

Material    Download as CSV 

Family Mesothuriidae Smirnov, 2012

Material    Download as CSV 

Genus Mesothuria Ludwig, 1894

Material    Download as CSV 

cf. Mesothuria morphospecies

Nomenclature: 

In the “Atlas of Abyssal Megafauna Morphotypes of the Clarion-Clipperton Fracture Zone” created for the ISA (http://ccfzatlas.com/), this morphospecies is listed as "Mesothuria morphotype".

Materials    Download as CSV 
  1. scientificName:
    Mesothuria sp.
    ; taxonConceptID:
    cf. Mesothuria morphospecies
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Holothuroidea
    ; order:
    Aspidochirotida
    ; family:
    Mesothuriidae
    ; taxonRank:
    genus
    ; genus:
    Mesothuria
    ; scientificNameAuthorship:
    Ludwig, 1894
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    Eastern Clarion-Clipperton Zone
    ; verbatimLocality:
    Site EPIRB
    ; maximumDepthInMeters:
    3918
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.6793
    ; decimalLongitude:
    -114.4074
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-23
    ; eventTime:
    11:55
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 7 (RV07)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Antonina Kremenetskaia, David L Pawson, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
  2. scientificName:
    Mesothuria sp.
    ; taxonConceptID:
    cf. Mesothuria morphospecies
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Holothuroidea
    ; order:
    Aspidochirotida
    ; family:
    Mesothuriidae
    ; taxonRank:
    genus
    ; genus:
    Mesothuria
    ; scientificNameAuthorship:
    Ludwig, 1894
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum B
    ; maximumDepthInMeters:
    4221
    ; locationRemarks:
    RV Thompson Cruise TN319
    ; decimalLatitude:
    12.5767
    ; decimalLongitude:
    -116.6767
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Autonomous Underwater Vehicle
    ; eventDate:
    2015-03-09
    ; eventTime:
    4:12
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 6 (AV06)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; identifiedBy:
    Antonina Kremenetskaia, David L Pawson, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2015
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 29

Figure 29.

cf. Mesothuria morphospecies observed in the UK-1 exploration contract area and eastern CCZ. Images (a-b) correspond with the data above.

acf. Mesothuria morphospecies in situ on seafloor. Scale bar is 10 cm. Image attribution: DJ Amon & CR Smith, University of Hawai’i.
bcf. Mesothuria morphospecies in situ on seafloor. Scale bar is 10 cm. Image attribution: Woods Hole Oceanographic Institution.

Family Synallactidae Ludwig, 1894

Material    Download as CSV 

cf. Synallactidae morphospecies 1

Materials    Download as CSV 
  1. scientificName:
    Synallactidae sp.
    ; taxonConceptID:
    cf. Synallactidae morphospecies 1
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Holothuroidea
    ; order:
    Aspidochirotida
    ; family:
    Synallactidae
    ; taxonRank:
    family
    ; scientificNameAuthorship:
    Ludwig, 1894
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    Eastern Clarion-Clipperton Zone
    ; verbatimLocality:
    Site EPIRB
    ; maximumDepthInMeters:
    3922
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.6793
    ; decimalLongitude:
    -114.4096
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-23
    ; eventTime:
    12:17
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 7 (RV07)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Antonina Kremenetskaia, David L Pawson, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
  2. scientificName:
    Synallactidae sp.
    ; taxonConceptID:
    cf. Synallactidae morphospecies 1
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Holothuroidea
    ; order:
    Aspidochirotida
    ; family:
    Synallactidae
    ; taxonRank:
    family
    ; scientificNameAuthorship:
    Ludwig, 1894
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum B
    ; maximumDepthInMeters:
    4219
    ; locationRemarks:
    RV Thompson Cruise TN319
    ; decimalLatitude:
    12.57901
    ; decimalLongitude:
    -116.6978
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Autonomous Underwater Vehicle
    ; eventDate:
    2015-03-09
    ; eventTime:
    15:36
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 6 (AV06)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; identifiedBy:
    Antonina Kremenetskaia, David L Pawson, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2015
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
  3. scientificName:
    Synallactidae sp.
    ; taxonConceptID:
    cf. Synallactidae morphospecies 1
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Holothuroidea
    ; order:
    Aspidochirotida
    ; family:
    Synallactidae
    ; taxonRank:
    family
    ; scientificNameAuthorship:
    Ludwig, 1894
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum B
    ; maximumDepthInMeters:
    4225
    ; locationRemarks:
    RV Thompson Cruise TN319
    ; decimalLatitude:
    12.5797
    ; decimalLongitude:
    -116.7276
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Autonomous Underwater Vehicle
    ; eventDate:
    2015-03-09
    ; eventTime:
    7:55
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 6 (AV06)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; identifiedBy:
    Antonina Kremenetskaia, David L Pawson, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2015
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 30

Figure 30.

cf. Synallactidae morphospecies 1 observed in the UK-1 exploration contract area and eastern CCZ. Images (a-c) correspond with the data above.

acf. Synallactidae morphospecies 1 in situ on seafloor. Scale bar is 10 cm. Image attribution: DJ Amon & CR Smith, University of Hawai’i.
bcf. Synallactidae morphospecies 1 in situ on seafloor. Scale bar is 10 cm. Image attribution: Woods Hole Oceanographic Institution.
ccf. Synallactidae morphospecies 1 in situ on seafloor. Scale bar is 10 cm. Image attribution: Woods Hole Oceanographic Institution.

cf. Synallactidae morphospecies 2

Material    Download as CSV 
  1. scientificName:
    Synallactidae sp.
    ; taxonConceptID:
    cf. Synallactidae morphospecies 2
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Holothuroidea
    ; order:
    Aspidochirotida
    ; family:
    Synallactidae
    ; taxonRank:
    family
    ; scientificNameAuthorship:
    Ludwig, 1894
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum B
    ; maximumDepthInMeters:
    4216
    ; locationRemarks:
    RV Thompson Cruise TN319
    ; decimalLatitude:
    12.5894
    ; decimalLongitude:
    -116.7152
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Autonomous Underwater Vehicle
    ; eventDate:
    2015-03-09
    ; eventTime:
    9:47
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 6 (AV06)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; identifiedBy:
    Antonina Kremenetskaia, David L Pawson, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2015
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 31

Figure 31.

cf. Synallactidae morphospecies 2 in situ on seafloor in the UK-1 exploration contract area. Image corresponds with the data above. Scale bar is 10 cm. Image attribution: Woods Hole Oceanographic Institution.

Genus Molpadiodemas Heding, 1935

Material    Download as CSV 

cf. Molpadiodemas morphospecies

Nomenclature: 

In the “Atlas of Abyssal Megafauna Morphotypes of the Clarion-Clipperton Fracture Zone” created for the ISA (http://ccfzatlas.com/), this morphospecies is listed as "Molpadiodemas​ morphotype".

Materials    Download as CSV 
  1. scientificName:
    Molpadiodemas sp.
    ; taxonConceptID:
    cf. Molpadiodemas morphospecies
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Holothuroidea
    ; order:
    Aspidochirotida
    ; family:
    Synallactidae
    ; taxonRank:
    genus
    ; genus:
    Molpadiodemas
    ; scientificNameAuthorship:
    Heding, 1935
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum A
    ; maximumDepthInMeters:
    4053
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.9608
    ; decimalLongitude:
    -116.5548
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-16
    ; eventTime:
    3:43
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 3 (RV03)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Antonina Kremenetskaia, David L Pawson, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
  2. scientificName:
    Molpadiodemas sp.
    ; taxonConceptID:
    cf. Molpadiodemas morphospecies
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Holothuroidea
    ; order:
    Aspidochirotida
    ; family:
    Synallactidae
    ; taxonRank:
    genus
    ; genus:
    Molpadiodemas
    ; scientificNameAuthorship:
    Heding, 1935
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum A
    ; maximumDepthInMeters:
    4055
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.9611
    ; decimalLongitude:
    -116.5541
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-16
    ; eventTime:
    3:52
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 3 (RV03)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Antonina Kremenetskaia, David L Pawson, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
  3. scientificName:
    Molpadiodemas sp.
    ; taxonConceptID:
    cf. Molpadiodemas morphospecies
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Holothuroidea
    ; order:
    Aspidochirotida
    ; family:
    Synallactidae
    ; taxonRank:
    genus
    ; genus:
    Molpadiodemas
    ; scientificNameAuthorship:
    Heding, 1935
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum B
    ; maximumDepthInMeters:
    4228
    ; locationRemarks:
    RV Thompson Cruise TN319
    ; decimalLatitude:
    12.5801
    ; decimalLongitude:
    -116.7228
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Autonomous Underwater Vehicle
    ; eventDate:
    2015-03-09
    ; eventTime:
    11:04
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 6 (AV06)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; identifiedBy:
    Antonina Kremenetskaia, David L Pawson, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2015
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
  4. scientificName:
    Molpadiodemas sp.
    ; taxonConceptID:
    cf. Molpadiodemas morphospecies
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Holothuroidea
    ; order:
    Aspidochirotida
    ; family:
    Synallactidae
    ; taxonRank:
    genus
    ; genus:
    Molpadiodemas
    ; scientificNameAuthorship:
    Heding, 1935
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum B
    ; maximumDepthInMeters:
    4213
    ; locationRemarks:
    RV Thompson Cruise TN319
    ; decimalLatitude:
    12.5708
    ; decimalLongitude:
    -116.7066
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Autonomous Underwater Vehicle
    ; eventDate:
    2015-03-09
    ; eventTime:
    15:07
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 6 (AV06)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; identifiedBy:
    Antonina Kremenetskaia, David L Pawson, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2015
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 32

Figure 32.

cf. Molpadiodemas morphospecies observed in the UK-1 exploration contract area. Images (a-d) correspond with the data above.

acf. Molpadiodemas morphospecies in situ on seafloor. Scale bar is 10 cm. Image attribution: DJ Amon & CR Smith, University of Hawai’i.
bcf. Molpadiodemas morphospecies in situ on seafloor. Scale bar is 10 cm. Image attribution: DJ Amon & CR Smith, University of Hawai’i.
ccf. Molpadiodemas morphospecies in situ on seafloor. Scale bar is 10 cm. Image attribution: Woods Hole Oceanographic Institution.
dcf. Molpadiodemas morphospecies in situ on seafloor. Scale bar is 10 cm. Image attribution: Woods Hole Oceanographic Institution.

Genus Paroriza Herouard, 1902

Material    Download as CSV 

cf. Paroriza morphospecies

Nomenclature: 

In the “Atlas of Abyssal Megafauna Morphotypes of the Clarion-Clipperton Fracture Zone” created for the ISA (http://ccfzatlas.com/), this morphospecies is listed as "Paroriza morphotype".

Material    Download as CSV 
  1. scientificName:
    Paroriza sp.
    ; taxonConceptID:
    cf. Paroriza morphospecies
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Holothuroidea
    ; order:
    Aspidochirotida
    ; family:
    Synallactidae
    ; taxonRank:
    genus
    ; genus:
    Paroriza
    ; scientificNameAuthorship:
    Herouard, 1902
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum B
    ; maximumDepthInMeters:
    4217
    ; locationRemarks:
    RV Thompson Cruise TN319
    ; decimalLatitude:
    12.57799
    ; decimalLongitude:
    -116.7028
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Autonomous Underwater Vehicle
    ; eventDate:
    2015-03-09
    ; eventTime:
    15:24
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 6 (AV06)
    ; individualCount:
    2
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; identifiedBy:
    Antonina Kremenetskaia, David L Pawson, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2015
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 33

Figure 33.

Two cf. Paroriza morphospecies specimens (likely male and female) in situ on seafloor in the UK-1 exploration contract area. Image corresponds with the data above. Scale bar is 10 cm. Image attribution: Woods Hole Oceanographic Institution.

Genus Pseudostichopus Théel, 1886

Material    Download as CSV 

cf. Pseudostichopus morphospecies

Nomenclature: 

In the “Atlas of Abyssal Megafauna Morphotypes of the Clarion-Clipperton Fracture Zone” created for the ISA (http://ccfzatlas.com/), this morphospecies is listed as "Pseudostichopus morphotype".

Materials    Download as CSV 
  1. scientificName:
    Pseudostichopus sp.
    ; taxonConceptID:
    cf. Pseudostichopus morphospecies
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Holothuroidea
    ; order:
    Aspidochirotida
    ; family:
    Synallactidae
    ; taxonRank:
    genus
    ; genus:
    Pseudostichopus
    ; scientificNameAuthorship:
    Théel, 1886
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum A
    ; maximumDepthInMeters:
    4032
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.8628
    ; decimalLongitude:
    -116.5485
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-21
    ; eventTime:
    4:59
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 6 (RV06)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Antonina Kremenetskaia, David L Pawson, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
  2. scientificName:
    Pseudostichopus sp.
    ; taxonConceptID:
    cf. Pseudostichopus morphospecies
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Holothuroidea
    ; order:
    Aspidochirotida
    ; family:
    Synallactidae
    ; taxonRank:
    genus
    ; genus:
    Pseudostichopus
    ; scientificNameAuthorship:
    Théel, 1886
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum A
    ; maximumDepthInMeters:
    4028
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.8616
    ; decimalLongitude:
    -116.5483
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-21
    ; eventTime:
    4:43
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 6 (RV06)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Antonina Kremenetskaia, David L Pawson, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 34

Figure 34.

cf. Pseudostichopus morphospecies observed in the UK-1 exploration contract area. Both images (a) and (b) correspond with the data in (a) and image (c) corresponds with the data in (b) above.

acf. Pseudostichopus morphospecies in situ on seafloor. Scale bar is 10 cm. Image attribution: DJ Amon & CR Smith, University of Hawai’i.
bcf. Pseudostichopus morphospecies in situ on seafloor. Scale bar is 10 cm. Image attribution: DJ Amon & CR Smith, University of Hawai’i.
ccf. Pseudostichopus morphospecies in situ on seafloor. Scale bar is 10 cm. Image attribution: DJ Amon & CR Smith, University of Hawai’i.

Genus Synallactes Ludwig, 1894

Material    Download as CSV 

cf. Synallactes morphospecies 1

Nomenclature: 

In the “Atlas of Abyssal Megafauna Morphotypes of the Clarion-Clipperton Fracture Zone” created for the ISA (http://ccfzatlas.com/), this morphospecies is listed as "Synallactes morphotype "white"".

Materials    Download as CSV 
  1. scientificName:
    Synallactes sp.
    ; taxonConceptID:
    cf. Synallactes morphospecies 1
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Holothuroidea
    ; order:
    Aspidochirotida
    ; family:
    Synallactidae
    ; taxonRank:
    genus
    ; genus:
    Synallactes
    ; scientificNameAuthorship:
    Ludwig, 1894
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum A
    ; maximumDepthInMeters:
    4021
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.8578
    ; decimalLongitude:
    -116.5481
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-21
    ; eventTime:
    3:48
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 6 (RV06)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Antonina Kremenetskaia, David L Pawson, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
  2. scientificName:
    Synallactes sp.
    ; taxonConceptID:
    cf. Synallactes morphospecies 1
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Holothuroidea
    ; order:
    Aspidochirotida
    ; family:
    Synallactidae
    ; taxonRank:
    genus
    ; genus:
    Synallactes
    ; scientificNameAuthorship:
    Ludwig, 1894
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum A
    ; maximumDepthInMeters:
    4079
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.8615
    ; decimalLongitude:
    -116.5483
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-21
    ; eventTime:
    4:40
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 6 (RV06)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Antonina Kremenetskaia, David L Pawson, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
  3. scientificName:
    Synallactes sp.
    ; taxonConceptID:
    cf. Synallactes morphospecies 1
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Holothuroidea
    ; order:
    Aspidochirotida
    ; family:
    Synallactidae
    ; taxonRank:
    genus
    ; genus:
    Synallactes
    ; scientificNameAuthorship:
    Ludwig, 1894
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    Eastern Clarion-Clipperton Zone
    ; verbatimLocality:
    Site EPIRB
    ; maximumDepthInMeters:
    3949
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.6794
    ; decimalLongitude:
    -114.4138
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-23
    ; eventTime:
    9:57
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 7 (RV07)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Antonina Kremenetskaia, David L Pawson, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 35

Figure 35.

cf. Synallactes morphospecies 1 observed in the UK-1 exploration contract area and eastern CCZ. Images (a-c) correspond with the data above.

acf. Synallactes morphospecies 1 in situ on seafloor. Scale bar is 10 cm. Image attribution: DJ Amon & CR Smith, University of Hawai’i.
bcf. Synallactes morphospecies 1 in situ on seafloor. Scale bar is 10 cm. Image attribution: DJ Amon & CR Smith, University of Hawai’i.
ccf. Synallactes morphospecies 1 in situ on seafloor. Image attribution: DJ Amon & CR Smith, University of Hawai’i.

cf. Synallactes morphospecies 2

Nomenclature: 

In the “Atlas of Abyssal Megafauna Morphotypes of the Clarion-Clipperton Fracture Zone” created for the ISA (http://ccfzatlas.com/), this morphospecies is listed as "Synallactes morphotype "pink"".

Materials    Download as CSV 
  1. scientificName:
    Synallactes sp.
    ; taxonConceptID:
    cf. Synallactes morphospecies 2
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Holothuroidea
    ; order:
    Aspidochirotida
    ; family:
    Synallactidae
    ; taxonRank:
    genus
    ; genus:
    Synallactes
    ; scientificNameAuthorship:
    Ludwig, 1894
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum B
    ; maximumDepthInMeters:
    4254
    ; locationRemarks:
    RV Thompson Cruise TN319
    ; decimalLatitude:
    12.4955
    ; decimalLongitude:
    -116.6505
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Autonomous Underwater Vehicle
    ; eventDate:
    2015-03-18
    ; eventTime:
    8:46
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 9 (AV09)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; identifiedBy:
    Antonina Kremenetskaia, David L Pawson, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2015
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
  2. scientificName:
    Synallactes sp.
    ; taxonConceptID:
    cf. Synallactes morphospecies 2
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Holothuroidea
    ; order:
    Aspidochirotida
    ; family:
    Synallactidae
    ; taxonRank:
    genus
    ; genus:
    Synallactes
    ; scientificNameAuthorship:
    Ludwig, 1894
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum B
    ; maximumDepthInMeters:
    4224
    ; locationRemarks:
    RV Thompson Cruise TN319
    ; decimalLatitude:
    12.4947
    ; decimalLongitude:
    -116.6308
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Autonomous Underwater Vehicle
    ; eventDate:
    2015-03-18
    ; eventTime:
    9:48
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 9 (AV09)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; identifiedBy:
    Antonina Kremenetskaia, David L Pawson, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2015
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 36

Figure 36.

cf. Synallactes morphospecies 2 observed in the UK-1 exploration contract area. Images (a-b) correspond with the data above.

acf. Synallactes morphospecies 2 in situ on seafloor. Image attribution: Woods Hole Oceanographic Institution.
bcf. Synallactes morphospecies 2 in situ on seafloor. Image attribution: Woods Hole Oceanographic Institution.

cf. Synallactes morphospecies 3

Nomenclature: 

In the “Atlas of Abyssal Megafauna Morphotypes of the Clarion-Clipperton Fracture Zone” created for the ISA (http://ccfzatlas.com/), this morphospecies is listed as "Synallactidae morphotype".

Materials    Download as CSV 
  1. scientificName:
    Synallactes sp.
    ; taxonConceptID:
    cf. Synallactes morphospecies 3
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Holothuroidea
    ; order:
    Aspidochirotida
    ; family:
    Synallactidae
    ; taxonRank:
    genus
    ; genus:
    Synallactes
    ; scientificNameAuthorship:
    Ludwig, 1894
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum B
    ; maximumDepthInMeters:
    4249
    ; locationRemarks:
    RV Thompson Cruise TN319
    ; decimalLatitude:
    12.4991
    ; decimalLongitude:
    -116.6409
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Autonomous Underwater Vehicle
    ; eventDate:
    2015-03-03
    ; eventTime:
    21:52
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 5 (AV05)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; identifiedBy:
    Antonina Kremenetskaia, David L Pawson, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2015
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
  2. scientificName:
    Synallactes sp.
    ; taxonConceptID:
    cf. Synallactes morphospecies 3
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Holothuroidea
    ; order:
    Aspidochirotida
    ; family:
    Synallactidae
    ; taxonRank:
    genus
    ; genus:
    Synallactes
    ; scientificNameAuthorship:
    Ludwig, 1894
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum B
    ; maximumDepthInMeters:
    4253
    ; locationRemarks:
    RV Thompson Cruise TN319
    ; decimalLatitude:
    12.4936
    ; decimalLongitude:
    -116.6506
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Autonomous Underwater Vehicle
    ; eventDate:
    2015-03-03
    ; eventTime:
    23:47
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 5 (RV05)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; identifiedBy:
    Antonina Kremenetskaia, David L Pawson, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2015
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 37

Figure 37.

cf. Synallactes morphospecies 3 observed in the UK-1 exploration contract area. Images (a-b) correspond with the data above.

acf. Synallactes morphospecies 3 in situ on seafloor. Scale bar is 10 cm. Image attribution: Woods Hole Oceanographic Institution.
bcf. Synallactes morphospecies 3 in situ on seafloor. Scale bar is 10 cm. Image attribution: Woods Hole Oceanographic Institution.

Order Elasipodida Théel, 1882

Material    Download as CSV 

Family Deimatidae Théel, 1882

Material    Download as CSV 

cf. Deimatidae morphospecies 1

Nomenclature: 

In the “Atlas of Abyssal Megafauna Morphotypes of the Clarion-Clipperton Fracture Zone” created for the ISA (http://ccfzatlas.com/), this morphospecies is listed as "Deimatidae gen. sp. morphotype".

Materials    Download as CSV 
  1. scientificName:
    Deimatidae sp.
    ; taxonConceptID:
    cf. Deimatidae morphospecies 1
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Holothuroidea
    ; order:
    Elasipodida
    ; family:
    Deimatidae
    ; taxonRank:
    family
    ; scientificNameAuthorship:
    Théel, 1882
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum B
    ; maximumDepthInMeters:
    4162
    ; locationRemarks:
    RV Thompson Cruise TN319
    ; decimalLatitude:
    12.3703
    ; decimalLongitude:
    -116.5194
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Autonomous Underwater Vehicle
    ; eventDate:
    2015-02-18
    ; eventTime:
    17:39
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 1 (AV01)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; identifiedBy:
    Antonina Kremenetskaia, David L Pawson, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2015
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
  2. scientificName:
    Deimatidae sp.
    ; taxonConceptID:
    cf. Deimatidae morphospecies
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Holothuroidea
    ; order:
    Elasipodida
    ; family:
    Deimatidae
    ; taxonRank:
    family
    ; scientificNameAuthorship:
    Théel, 1882
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum B
    ; maximumDepthInMeters:
    4254
    ; locationRemarks:
    RV Thompson Cruise TN319
    ; decimalLatitude:
    12.4939
    ; decimalLongitude:
    -116.6504
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Autonomous Underwater Vehicle
    ; eventDate:
    2015-03-03
    ; eventTime:
    23:47
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 5 (AV05)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; identifiedBy:
    Antonina Kremenetskaia, David L Pawson, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2015
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 38

Figure 38.

cf. Deimatidae morphospecies 1 observed in the UK-1 exploration contract area. Images (a-b) correspond with the data above.

acf. Deimatidae morphospecies 1 in situ on seafloor. Scale bar is 10 cm. Image attribution: Woods Hole Oceanographic Institution.
bcf. Deimatidae morphospecies 1 in situ on seafloor. Scale bar is 5 cm. Image attribution: Woods Hole Oceanographic Institution.

cf. Deimatidae morphospecies 2

Nomenclature: 

In the “Atlas of Abyssal Megafauna Morphotypes of the Clarion-Clipperton Fracture Zone” created for the ISA (http://ccfzatlas.com/), this morphospecies is listed as "Orphnurgus morphotype".

Material    Download as CSV 
  1. scientificName:
    Deimatidae sp.
    ; taxonConceptID:
    cf. Deimatidae morphospecies 2
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Holothuroidea
    ; order:
    Elasipodida
    ; family:
    Deimatidae
    ; taxonRank:
    family
    ; scientificNameAuthorship:
    Théel, 1882
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum B
    ; maximumDepthInMeters:
    4256
    ; locationRemarks:
    RV Thompson Cruise TN319
    ; decimalLatitude:
    12.49096
    ; decimalLongitude:
    -116.6511
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Autonomous Underwater Vehicle
    ; eventDate:
    2015-03-03
    ; eventTime:
    22:23
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 5 (AV05)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; identifiedBy:
    Antonina Kremenetskaia, David L Pawson, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2015
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 39

Figure 39.

cf. Deimatidae morphospecies 2 observed in the UK-1 exploration contract area. Image corresponds with the data above. Scale bar is 10 cm. Image attribution: Woods Hole Oceanographic Institution.

Genus Deima Théel, 1879

Material    Download as CSV 

Deima cf. validum Théel, 1879

Nomenclature: 

In the “Atlas of Abyssal Megafauna Morphotypes of the Clarion-Clipperton Fracture Zone” created for the ISA (http://ccfzatlas.com/), this morphospecies is listed as "Deima morphotype".

Materials    Download as CSV 
  1. scientificName:
    Deima validum
    ; taxonConceptID:
    Deima cf. validum
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Holothuroidea
    ; order:
    Elasipodida
    ; family:
    Deimatidae
    ; taxonRank:
    species
    ; genus:
    Deima
    ; scientificNameAuthorship:
    Théel, 1879
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum B
    ; maximumDepthInMeters:
    4254
    ; locationRemarks:
    RV Thompson Cruise TN319
    ; decimalLatitude:
    12.4977
    ; decimalLongitude:
    -116.6525
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Autonomous Underwater Vehicle
    ; eventDate:
    2015-03-04
    ; eventTime:
    4:01
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 5 (AV05)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; identifiedBy:
    Antonina Kremenetskaia, David L Pawson, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2015
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
  2. scientificName:
    Deima validum
    ; taxonConceptID:
    Deima cf. validum
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Holothuroidea
    ; order:
    Elasipodida
    ; family:
    Deimatidae
    ; taxonRank:
    species
    ; genus:
    Deima
    ; scientificNameAuthorship:
    Théel, 1879
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum B
    ; maximumDepthInMeters:
    4226
    ; locationRemarks:
    RV Thompson Cruise TN319
    ; decimalLatitude:
    12.5814
    ; decimalLongitude:
    -116.7244
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Autonomous Underwater Vehicle
    ; eventDate:
    2015-03-09
    ; eventTime:
    8:42
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 6 (AV06)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; identifiedBy:
    Antonina Kremenetskaia, David L Pawson, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2015
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 40

Figure 40.

Deima cf. validum observed in the UK-1 exploration contract area. Images (a-b) correspond with the data above.

aDeima cf. validum in situ on seafloor. Scale bar is 10 cm. Image attribution: Woods Hole Oceanographic Institution.
bDeima cf. validum in situ on seafloor. Scale bar is 10 cm. Image attribution: Woods Hole Oceanographic Institution.

Genus Oneirophanta Théel, 1879

Material    Download as CSV 

cf. Oneirophanta morphospecies

Nomenclature: 

In the “Atlas of Abyssal Megafauna Morphotypes of the Clarion-Clipperton Fracture Zone” created for the ISA (http://ccfzatlas.com/), this morphospecies is listed as "Oneirophanta setigera Ludwig, 1893".

Material    Download as CSV 
  1. scientificName:
    Oneirophanta sp.
    ; taxonConceptID:
    cf. Oneirophanta morphospecies
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Holothuroidea
    ; order:
    Elasipodida
    ; family:
    Deimatidae
    ; taxonRank:
    genus
    ; genus:
    Oneirophanta
    ; scientificNameAuthorship:
    Théel, 1879
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum A
    ; maximumDepthInMeters:
    4033
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.8627
    ; decimalLongitude:
    -116.5484
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-21
    ; eventTime:
    4:57
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 6 (RV06)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Antonina Kremenetskaia, David L Pawson, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 41

Figure 41.

cf. Oneirophanta morphospecies observed in the UK-1 exploration contract area. Both images (a) and (b) correspond with the data in (a) above.

acf. Oneirophanta morphospecies in situ on seafloor. Scale bar is 10 cm. Image attribution: DJ Amon & CR Smith, University of Hawai’i.
bcf. Oneirophanta morphospecies in situ on seafloor. Scale bar is 10 cm. Image attribution: DJ Amon & CR Smith, University of Hawai’i.

Family Elpidiidae Théel, 1882

Material    Download as CSV 

cf. Elpidiidae morphospecies 1

Material    Download as CSV 
  1. scientificName:
    Elpidiidae sp.
    ; taxonConceptID:
    cf. Elpidiidae morphospecies 1
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Holothuroidea
    ; order:
    Elasipodida
    ; family:
    Elpidiidae
    ; taxonRank:
    family
    ; scientificNameAuthorship:
    Théel, 1882
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum A
    ; maximumDepthInMeters:
    4020
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.8554
    ; decimalLongitude:
    -116.5477
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-21
    ; eventTime:
    3:05
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 6 (RV06)
    ; individualCount:
    1
    ; lifeStage:
    Young specimen
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Antonina Kremenetskaia, David L Pawson, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 42

Figure 42.

cf. Elpidiidae morphospecies 1 observed in the UK-1 exploration contract area. Image corresponds with the data above. Scale bar is 10 cm. Image attribution: DJ Amon & CR Smith, University of Hawai’i.

cf. Elpidiidae morphospecies 2

Nomenclature: 

In the “Atlas of Abyssal Megafauna Morphotypes of the Clarion-Clipperton Fracture Zone” created for the ISA (http://ccfzatlas.com/), this morphospecies is listed as "Elpidiidae morphotype "double velum"".

Material    Download as CSV 
  1. scientificName:
    Elpidiidae sp.
    ; taxonConceptID:
    cf. Elpidiidae morphospecies 2
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Holothuroidea
    ; order:
    Elasipodida
    ; family:
    Elpidiidae
    ; taxonRank:
    family
    ; scientificNameAuthorship:
    Théel, 1882
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    UK Seabed Resources Ltd exploration contract area (UK-1)
    ; verbatimLocality:
    UK-1 Stratum B
    ; maximumDepthInMeters:
    4254
    ; locationRemarks:
    RV Thompson Cruise TN319
    ; decimalLatitude:
    12.4988
    ; decimalLongitude:
    -116.6496
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Autonomous Underwater Vehicle
    ; eventDate:
    2015-03-18
    ; eventTime:
    8:40
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 9 (AV09)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; identifiedBy:
    Antonina Kremenetskaia, David L Pawson, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2015
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 43

Figure 43.

cf. Elpidiidae morphospecies 2 in situ on seafloor in the UK-1 exploration contract area. Image corresponds with the data above. Scale bar is 10 cm. Image attribution: Woods Hole Oceanographic Institution.

Genus Amperima Pawson, 1965

Material    Download as CSV 

cf. Amperima morphospecies

Nomenclature: 

In the “Atlas of Abyssal Megafauna Morphotypes of the Clarion-Clipperton Fracture Zone” created for the ISA (http://ccfzatlas.com/), this morphospecies is listed as "Amperima morphotype".

Material    Download as CSV 
  1. scientificName:
    Amperima sp.
    ; taxonConceptID:
    cf. Amperima morphospecies
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ; class:
    Holothuroidea
    ; order:
    Elasipodida
    ; family:
    Elpidiidae
    ; taxonRank:
    genus
    ; genus:
    Amperima
    ; scientificNameAuthorship:
    Pawson, 1965
    ; waterBody:
    Pacific Ocean
    ; stateProvince:
    Clarion-Clipperton Zone
    ; locality:
    Eastern Clarion-Clipperton Zone
    ; verbatimLocality:
    Site EPIRB
    ; maximumDepthInMeters:
    3938
    ; locationRemarks:
    RV Melville Cruise MV1313
    ; decimalLatitude:
    13.6794
    ; decimalLongitude:
    -114.4113
    ; geodeticDatum:
    WGS84
    ; coordinateUncertaintyInMeters:
    25
    ; samplingProtocol:
    Remotely Operated Vehicle
    ; eventDate:
    2013-10-23
    ; eventTime:
    12:36
    ; habitat:
    Abyssal polymetallic-nodule field
    ; fieldNumber:
    Dive 7 (RV07)
    ; individualCount:
    1
    ; lifeStage:
    Adult
    ; preparations:
    Imaged only
    ; behavior:
    On seafloor
    ; recordedBy:
    Diva J Amon, Amanda F Ziegler
    ; occurrenceStatus:
    present
    ; associatedReferences:
    Amon DJ, Ziegler AF, Dahlgren TG, Glover AG, Goineau A, Gooday AJ, Wiklund H, Smith CR. Insights into the abundance and diversity of abyssal megafauna in a polymetallic-nodule region in the eastern Clarion-Clipperton Zone. Scientific Reports. 2016;6. doi: 10.1038/srep30492
    ; identifiedBy:
    Antonina Kremenetskaia, David L Pawson, Diva J Amon, Amanda F Ziegler
    ; dateIdentified:
    2014
    ; identificationRemarks:
    Identified only from imagery
    ; identificationQualifier:
    cf.
    ; language:
    en
    ; institutionCode:
    UHM
    ; datasetName:
    ABYSSLINE
    ; basisOfRecord:
    HumanObservation
Notes: 

Fig. 44

Figure 44.

cf. Amperima morphospecies observed in the eastern Clarion-Clipperton Zone. Both images (a) and (b) correspond with the data in (a) above.

acf. Amperima morphospecies in situ on seafloor. Scale bar is 10 cm. Image attribution: DJ Amon & CR Smith, University of Hawai’i.
bcf. Amperima morphospecies in situ on seafloor. Scale bar is 10 cm. Image attribution: DJ Amon & CR Smith, University of Hawai’i.

Genus Peniagone Théel, 1882

Material    Download as CSV 

cf. Peniagone morphospecies 1

Nomenclature: 

In the “Atlas of Abyssal Megafauna Morphotypes of the Clarion-Clipperton Fracture Zone” created for the ISA (http://ccfzatlas.com/), this morphospecies is listed as "Peniagone morphotype “pink, large velum"".

Material    Download as CSV 
  1. scientificName:
    Peniagone sp.
    ; taxonConceptID:
    cf. Peniagone morphospecies 1
    ; kingdom:
    Animalia
    ; phylum:
    Echinodermata
    ;