This species is widely distributed in Europe, occurs also in the Caucasus, the Transcaucasus, Turkey, Kazakhstan and Turkmenistan. In Russia, it is known from the steppe zone from Kursk Prov. to West Siberia.

In Russia, the species occurs in rodent burrows (mostly of marmots and sousliks). In some localities, beetles can be found in large numbers near burrow entrances in May-June (Isajev 1995). However, in the UK, this species was collected under vegetable debris and sheep, cow and horse dung (Jessop 1986). It was also collected from rabbit latrines (Jason Mate, pers. comm.) Probably, in west Europe, in the areas without Marmotini rodents, this species develop in burrows of other rodents or rabbits, but its life cycle needs further research.

The species is distributed in central, south and eastern Europe, the Transcaucasus, Asia Minor, Kazakhstan and Turkmenistan. In Russia, it is known from Ulyanovsk and Novosibirsk Provinces, Volgograd, Rostov, Orenburg Provinces and Krasnoyarsk Terr.

The species occurs in open biotopes, in marmot holes and in the dung of domestic animals.

The species occurs in central and eastern Europe, north and eastern Kazakhstan, Kyrgyzstan. In Russia, it is known from the European part and West Siberia (up to Krasnoyarsk in the north-east).

The beetles and larvae feed on horse dung.

The species is distributed in the steppe zone of central and eastern Europe and Kazakhstan.

A nidicolous steppe species occurring in gopher and marmot holes.

The distribution range of this species includes almost the whole of Europe (except for the extreme north), the Transcaucasus, Asia Minor, west Kazakhstan and Tajikistan. In Russia, the species is known from Leningrad, Nizhny Novgorod, Voronezh, Saratov, Volgograd and Astrakhan Provinces.

A coprophagous species occurring in the dung of cows, horses and wild ungulates. The species occurs in different biotopes: in eastern Europe, in forests; Isajev (1995) recorded it from steppes; we collected it on sandy soils on the Akhtuba riverside in Dosang environs (Astrakhan Prov.).

The distribution range of this species includes the whole of Europe (except for the extreme north), North Africa, the Transcaucasus, Asia Minor, Kazakhstan, Middle Asia, Mongolia. It was imported to North America. In Russia, it occurs throughout the country in the European part, reaching Baikal Region in the east.

The species occurs in cow and horse dung, in marmot holes and also is attracted to light. In Russia, the beetles are active from March to October. It is a common, locally abundant species.

The species occurs in Europe (except for the extreme north), west, middle and Central Asia; it was imported to North America. In Russia, it is distributed throughout the country.

This eurybiont species is abundant throughout its range occurring mostly in open biotopes, in the dung of domestic and wild animals.

The species is widely distributed in Europe, North Africa, Kazakhstan, Middle and Central Asia, imported to North America and Australia. In Russia, it occurs throughout the country up to Eastern Siberia in the east.

In Russia, imagoes are active from April to October occurring in open biotopes, pastures, in cow, sheep and horse dung.

The species is distributed throughout Europe (except for the extreme north), west Asia, Kazakhstan and Middle Asia; imported to North America. In Russia, it occurs from the western border up to Transbaikal Region.

The species is common in all the parts of its range. It occurs in open biotopes in the dung of different animals, in carrion and in marmot holes.

The distribution range of this species includes the whole of Europe, North Africa, the Caucasus, the Transcaucasus, Kazakhstan, Middle Asia, Afghanistan, Mongolia, north and south-western China, the Korean Peninsula and Japan; it was imported to North America. In Russia, it is distributed throughout the country.

The species mostly occurs in cow and horse dung, in open biotopes.

The species occurs throughout Europe, in Morocco, Egypt, the Caucasus, the Transcaucasus, Asia Minor, Syria, Kazakhstan, middle and Central Asia. In Russia, it occurs from the western borders up to southern Yakutia.

The species is common in the major part of its range and prefers open biotopes. The beetles feed on horse, cow and donkey dung; they are attracted to light and occur from April to September.

The species occurs in Europe, North Africa, the Transcaucasus, Asia Minor, Kazakhstan, the mountains of Middle Asia; it was imported to North America. In Russia, it is distributed throughout the European part and occurs in south of West Siberia.

A coprophagous species feeding on cow, horse, donkey and sheep dung.

The species occurs in central, south and eastern Europe, North Africa, west Asia, Kazakhstan and Middle Asia. In Russia, it is distributed from the western border to Transbaikal Region, mostly in the steppe zone and the forest-steppe subzone of the deciduous forest zone.

The species occurs in open biotopes, in the dung of domestic animal and in marmot holes. On the territory of Russia, the beetles are active from March to October. The species is common throughout its range.

The species occurs throughout Europe, in North Africa (Morocco, Algeria), the Caucasus, the Transcaucasus, Asia Minor, Syria, Lebanon, Israel, Kazakhstan, middle Asia and Mongolia; it was imported to USA and Canada. In Russia, it is distributed from the western border to Yakutia.

A coprophagous species feeding on dung of cows, horses and wild ungulates. In Russia, this species is common; in southern regions, it occurs from March to December.

The species is distributed in Europe, except for the extreme north, North Africa (Morocco, Tunisia), the Trancaucasus, west Asia, Kazakhstan and Middle Asia. In Russia, it is common in the European part, reaching the Transbaikal Region to the east.

The species occurs in the dung of domestic animals and in marmot holes.

The distribution range of this species includes almost the whole of Europe (except for the extreme north), the Caucasus, the Transcaucasus, Kazakhstan, Middle Asia, Asia Minor, Mongolia, north China, Korean Peninsula and Japan. In Russia, it is distributed throughout the country.

It is a common, locally abundant species feeding on cow, horse and sheep dung.

The species is distributed in Europe, North Africa, west Asia, Kazakhstan and Middle Asia. In Russia, it mostly occurs in the Vorga Region and the Ciscaucasus.

A coprophagous species occurring in the dung of different animals. It prefers arid biotopes with sandy and sandy clay soils.

The species is widely distributed in central and south Europe; it is also known from North Africa, the Transcaucasus, Asia Minor, Syria and Turkmenistan. In Russia, it is known from Samara and Astrakhan Provinces.

The species occurs in cow dung.

The species occurs in central andeEastern Europe, the Transcaucasus, Middle Asia. In Russia, it is mainly distributed in northern Caucasus and the Volga Region.

The species occurs in cow and horse dung, mostly in open biotopes.

The species occurs in Europe (up to Finland in the north), North Africa (Morocco), the Caucasus, the Transcaucasus, Asia Minor, Kazakhstan, Middle Asia, Afghanistan, Mongolia, China, North Korea. It was imported to Canada and the USA. In Russia, the species is mostly distributed in the forest-steppe subzone of the deciduous forest zone from the western border to the Amur Region.

A coprophagous species occurring in ungulate dung.

The distribution range of this species includes central and southern Europe, North Africa, the Transcaucasus, Asia Minor, Kazakhstan and Middle Asia. In Russia, it is mostly distributed in the steppe zone and forest-steppe subzone of the zone of deciduous forest from the western border to South Siberia.

The species occurs in diverse biotopes, in cow and horse dung and in riverside debris.

The species is distributed in southern part of western and central Palaearctic.

The species mostly occurs in steppe biotopes. A generalist coprophage, feeding on ungulate dung, faeces of rodents and birds.

The spcies is widely distributed in the Palaearctic Region, introduced in North America. In Russia, it occurs throughout the European part, in West and Central Siberia.

A generalist coprophagous species feeding mostly on cattle dung. It was also registered feeding on carrion and rotten food.

A western Palaearctic species. In Russia, it is common in the central and southern European part, reaching southern Urals Mountains in the east.

A generalist coprophagous species feeding mostly on cattle dung.

Southern Ukraine, southern European Russia up to Volga Basin in the east, north-western Kazakhstan.

A nidicolous species inhabiting burrows of steppe rodents. It was also registered feeding on sheep and cattle dung.

The species is distributed in southern and middle Europe, northern Africa, Asia Minor and Iranian Plateau.

A common, sometimes abundant generalist coprophage feeding mostly on cattle, horse and sheep dung.

The species is widely distributed in the southern Palaearctic.

The species mostly occurs in steppe and semi-desert biotopes. A generalist coprophage, feeding mostly on ungulate dung.

The species is distributed in southern Europe, Asia Minor and western Central Asia.

A generalist coprophagous species feeding mostly on cattle and horse dung.