Biodiversity Data Journal :
Taxonomic Paper
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Corresponding author: Magdi El-Hawagry (elhawagry@cu.edu.eg), Ahmed Mostafa Soliman (ammsoliman@gmail.com), Hathal Mohammed Al Dhafer (hdhafer@ksu.edu.sa)
Academic editor: Torsten Dikow
Received: 02 Nov 2020 | Accepted: 08 Jan 2021 | Published: 13 Jan 2021
© 2021 Magdi El-Hawagry, Ahmed Soliman, Hathal Al Dhafer
This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
Citation:
El-Hawagry M, Soliman AM, Al Dhafer HM (2021) The family Conopidae (Diptera) in Egypt and Saudi Arabia. Biodiversity Data Journal 9: e60287. https://doi.org/10.3897/BDJ.9.e60287
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The present study is one in a series of planned studies aiming to catalogue the whole order Diptera in both Egypt and Saudi Arabia.
All known Egyptian and Saudi Arabian conopid taxa are systematically catalogued in the present study. Three species are recorded herein for the first time from Saudi Arabia: Conops (Asiconops) elegans Meigen, 1804 and Thecophora atra (Fabricius, 1775) (Al-Baha region, south-western of Saudi Arabia), and Conops (Conops) quadrifasciatus De Geer, 1776 (Tabuk region, north-western of Saudi Arabia). Physocephala variegata (Meigen, 1924) is also recorded for the first time from Gebel Elba, the south-eastern triangle of Egypt. Considering that Gebel Elba in Egypt and Al-Baha in Saudi Arabia are affiliated to the Afrotropical Region, this is the first time Physocephala variegata and Thecophora atra have been recorded from the Afrotropical Region. An updated taxonomy, world and local distributions, dates of collection and some coloured photographs are provided.
thick-headed flies, local distribution, dates of collection, new records
The Conopidae, commonly known as thick-headed flies, are an interesting brachycerous family of usually large-sized wasp-like flies. Many conopids are most frequently found at flowers feeding on nectar using their long proboscis and considered as important pollinators. The majority of conopid larvae are obligatory endoparasitoids of adult Hymenoptera, such as bees and wasps. Adult conopid females aggressively intercept their hymenopterous hosts in the field to deposit their eggs (
Egypt and Saudi Arabia are two neighbouring Middle Eastern countries, separated by the Gulf of Aqaba and the Red Sea (Fig.
Some previous studies have been carried out in Egypt on the taxonomy and faunistics of the family Conopidae, including
The abundance and diversity of conopid species in both Egypt and Saudi Arabia seem to be low (17 species in Egypt and 12 species in Saudi Arabia) comparable with the total number of species known from the Palaearctic Region (173 species) and from the Afrotropical Region (187 species) (Stuke 2017). However, this should be taken with caution, since the family Conopidae seems to lack sampling efforts, especially in Saudi Arabia. This is most likely due to the fact that documentation of biological diversity in Saudi Arabia began in the second half of the 1960s and no systematic studies on the Conopidae have been previously conducted in this country. We think that the faunistic data of these flies in Egypt and Saudi Arabia is still scanty and more efforts would be highly required.
The taxonomy of Conopidae in both Egypt and Saudi Arabia is updated in the present study. Twenty species assigned to five genera and three subfamilies (Conopinae, Myopinae and Zodioninae) are treated (Table
Conopid species treated in the present study as recorded from Egypt and Saudi Arabia (* = recorded, - = not recorded).
Species | Egypt | Saudi Arabia |
Subfamily CONOPINAE | ||
Tribe CONOPINI | ||
Conops (Asiconops) elegans Meigen, 1804 | * | * (new record) |
Conops (Asiconops) flavifrons Meigen, 1804 | * | - |
Conops (Asiconops) nubeculipennis Bezzi, 1901 | * | * |
Conops (Conops) flavicauda (Bigot, 1880) | * | - |
Conops (Conops) quadrifasciatus De Geer, 1776 | - | * (new record) |
Conops (Conops) rufiventris Macquart, 1849 | * | - |
Conops (Conops) tomentosus Kröber, 1916 | - | * |
Tribe PHYSOCEPHALINI | ||
Physocephala antiqua (Wiedemann, 1830) | * | * |
Physocephala chrysorrhoea (Meigen, 1824) | * | * |
Physocephala pusilla (Meigen, 1804) | * | * |
Physocephala variegata (Meigen, 1824) | * | * |
Physocephala vittata (Fabricius, 1794) | * | - |
Subfamily MYOPINAE | ||
Tribe MYOPINI | ||
Myopa dorsalis Fabricius, 1794 | * | - |
Myopa picta Panzer, 1798 | * | - |
Myopa stigma Meigen, 1824 | - | * |
Myopa testacea (Linnaeus, 1767) | * | - |
Tribe THECOPHORINI | ||
Thecophora atra (Fabricius, 1775) | * | * (new record) |
Thecophora fulvipes (Robineau-Desvoidy, 1830) | * | * |
Subfamily ZODIONINAE | ||
Zodion cinereum (Fabricius, 1794) | * | * |
Zodion erythrurum Rondani, 1865 | * | - |
Physocephala variegata (Meigen) is recorded for the first time from Gebel Elba, the south-eastern triangle of Egypt and Thecophora atra (Fabricius) is recorded for the first time from the south-western part of Saudi Arabia. Considering that Gebel Elba in Egypt and the south-western part in Saudi Arabia including Al-Baha Region are affiliated to the Afrotropical Region, this is the first time that these two species from the Afrotropical Region have been recorded.
This study is one in a series of taxonomic studies on different Egyptian and Saudi Arabian dipteran taxa aiming to catalogue the whole order Diptera in the two countries.
Data sources. Data of the present study are obtained from three main sources: 1. Specimens preserved in the Egyptian and Saudi Arabian insect collections and museums, namely: Efflatoun Bey’s collection, Cairo University, Egypt (EFC); Collection of Entomological Society of Egypt (ESEC); King Saud University Museum of Arthropods, Saudi Arabia (KSMA) and Personal collection M. El-Hawagry (MSHC). 2. Material collected by the authors and their co-workers. 3. Previous studies on the Egyptian and Saudi Arabian Conopidae. The majority of specimens preserved in EFC and ESEC were determined by Dr. Otto Kröber. However, all identifications have been rechecked by the first author using Kröber (1927), Stuke and Clements (2008), Azmy (2016) and Azmy et al. (2016). Photos of Thecophora atra have been checked by Dr. Jens-Hermann Stuke (personal communication).
Study area. Ecologists divide Egypt into eight ecological zones: the Coastal Strip (= Mediterranean Coastal Strip), Lower Nile Valley and Delta, Upper Nile Valley, Fayoum, Eastern Desert, Western Desert, Sinai and Gebel Elba (
Classification. The classification and nomenclature in the present study basically follows that used in
World distribution.
Local distribution and dates of collection. Localities within each Egyptian ecological zone or each Saudi Arabian administrative region are arranged alphabetically and written after a colon following each zone or region, followed by the dates of collection between parentheses. Sources for the local distribution of each species in each country are given between square brackets at the end of the section. Coordinates of Egyptian and Saudi Arabian localities of the family Conopidae are listed (Table
A gazetteer of some Egyptian and Saudi Arabian localities of the family Conopidae.
Country | Locality | Governorate | Ecological zone or Region | Latitude (N) / Longitude (E) |
Egypt | Abu-Kir | Alexandria | Coastal Strip |
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Abu-Rawash | Giza | Lower Nile Valley and Delta |
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Abu-Sueir | Ismailia | Eastern Desert |
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Amria | Alexandria | Coastal Strip |
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Ashmoun | Menofiya | Lower Nile Valley and Delta |
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Aswan, south | Aswan | Upper Nile Valley |
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Balteem | Kafr Al-Sheikh | Coastal Strip |
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Barrage | Qalyoubia | Lower Nile Valley and Delta |
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Behig | Alexandria | Coastal Strip |
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Burg | Alexandria | Coastal Strip |
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Burgash | Giza | Lower Nile Valley and Delta |
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Cairo | Cairo | Lower Nile Valley and Delta |
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Cairo-Suez Road | Cairo | Eastern Desert |
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Dahshour | Giza | Lower Nile Valley and Delta |
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Dakhla | New Valley | Western Desert |
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Dekhela | Alexandria | Coastal Strip |
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Edfina | Al-Behaira | Lower Nile Valley and Delta |
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Ein Gedeirat | North Sinai | Sinai |
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Ein Moussa | South Sinai | Sinai |
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El-Arish | North Sinai | Sinai |
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El-Gebel El-Asfar | Qalyoubia | Lower Nile Valley and Delta |
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El-Katta | Giza | Lower Nile Valley and Delta |
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El-Magadlah | Giza | Lower Nile Valley and Delta |
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El-Mallah East | Cairo | Eastern Desert |
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El-Wasfia | Ismailia | Eastern Desert |
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Esna | Qena | Upper Nile Valley |
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Ezbet Nakhl | Qalyoubia | Lower Nile Valley and Delta |
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Fayed | Ismailia | Eastern Desert |
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Fayoum | Fayoum | Fayoum |
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Gebel Elba | Red Sea | Gebel Elba |
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Geneifa | Suez | Eastern Desert |
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Gezira | Cairo | Lower Nile Valley and Delta |
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Girza | Fayoum | Fayoum |
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Giza | Giza | Lower Nile Valley and Delta |
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Halaib | Red Sea | Eastern Desert |
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Hawamdia | Giza | Lower Nile Valley and Delta |
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Helwan | Cairo | Lower Nile Valley and Delta |
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Ismailia | Ismailia | Eastern Desert |
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Kafr Ghatati | Giza | Lower Nile Valley and Delta |
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Kafr Hakim | Giza | Lower Nile Valley and Delta |
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Kerdassa | Giza | Lower Nile Valley and Delta |
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Kom Ombo | Aswan | Upper Nile Valley |
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Kom Osheem | Fayoum | Fayoum |
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Kubba | Cairo | Lower Nile Valley and Delta |
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Maadi | Cairo | Lower Nile Valley and Delta |
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Madinet El-Sadat | Menofiya | Western Desert |
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Madinet Badr | Cairo | Lower Nile Valley and Delta |
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Mansouriah | Giza | Lower Nile Valley and Delta |
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Mariout | Alexandria | Coastal Strip |
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Mazghouna | Giza | Lower Nile Valley and Delta |
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Mersa Matrouh | Matrouh | Coastal Strip |
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Orman | Giza | Lower Nile Valley and Delta |
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Pyramids | Giza | Lower Nile Valley and Delta |
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Rafah | North Sinai | Sinai |
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Salloum | Matrouh | Coastal Strip |
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Saqqara | Giza | Lower Nile Valley and Delta |
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Sennouris | Fayoum | Fayoum |
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Sentris | Menofiya | Lower Nile Valley and Delta |
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Siwa Oasis | Matrouh | Western Desert |
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Suez | Suez | Eastern Desert |
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Tamiya | Fayoum | Fayoum |
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Turah | Cairo | Lower Nile Valley and Delta |
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W. Asal | Red Sea | Eastern Desert |
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W. Dar El-Maskhara | Cairo | Eastern Desert |
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W. Digla | Cairo | Eastern Desert |
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W. El-Arbaein | South Sinai | Sinai |
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W. El-Daiqa | North Sinai | Sinai |
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W. El-Lega | South Sinai | Sinai |
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W. El-Natroun | Al-Behaira | Western Desert |
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W. El-Rahba | South Sinai | Sinai |
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W. Firan | South Sinai | Sinai |
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W. Garawi | Cairo | Eastern Desert |
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W. Gedeirat | North Sinai | Sinai |
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W. Gharagid | Cairo | Eastern Desert |
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W. Hoff | Cairo | Eastern Desert |
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W. Ibib | Red Sea | Gebel Elba |
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W. Isla | South Sinai | Sinai |
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W. Morrah | Cairo | Eastern Desert |
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W. Rishrash | Giza | Eastern Desert |
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W. Um Elek | Cairo | Eastern Desert |
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W. Zohleiga | Cairo | Eastern Desert |
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Saudi Arabia | Abu-Arish | Jazan | Jazan |
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Al-Aqiq | Al-Aqiq | Al-Baha |
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Al-Mekhwa | Al-Mekhwa | Al-Baha |
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Ghabet Shahba | Al-Baha | Al-Baha |
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Jabal Shada al-A’la Nature Reserve | Al-Mekhwa | Al-Baha |
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Jazan | Jazan | Jazan |
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Jeddah | Jeddah | Makka Al-Mukarramah |
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Marabah | Marabah | Asir |
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Najran | Najran | Najran |
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Raydah Nature Reserve | Abha | Asir |
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Tabouk | Tabouk | Tabouk |
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Unayzah | Unayzah | Al-Qassim |
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Abbreviations used:
Conops Linnaeus, 1758: 604. Type species: Conops flavipes Linnaeus, by subsequent designation of
Asiconops Chen, 1939: 171. Type species: Conops aureomaculatus Kröber, by original designation.
Conops elegans Meigen, 1804: 275. Type locality: France (Marseille).
Conops elegans var. fuscipennis Macquart 1849: 472. Type locality: Algeria.
Conops algirus Macquart 1849: 473. Type locality: Algeria (environs du Cercle de Lacalle).
Conopaejus aegyptiacus Rondani, 1850: 167. Type locality: Egypt.
Conops fuscanipennis Bigot, 1887: 32. Type locality: Italy (Sicily).
Conops elegans var. minutus Kröber, 1916: 48. Type locality: Not given (Cyprus, Limassol in
AF: Democratic Republic of the Congo, Ethiopia, Gabon, Guinea, Guinea-Bissau, Malawi, Mozambique, Namibia, Nigeria, Saudi Arabia [as “South-western part”] (first record), South Africa, Tanzania, Yemen. PA: Algeria, Bulgaria, Cyprus, Egypt, France, Israel, Italy, Morocco, Romania, Spain, Syria, Tunisia, Turkey.
Local distribution and dates of collection (Fig.
Conops flavifrons Meigen, 1804: 281. Type locality: France (Lyon).
Physocephala pugioniformis Becker, 1913: 611. Type locality: Iran.
Conops minor Becker, 1922: 201. Type locality: Russia, Italy, Hungary & Greece.
Conops kroeberi var. immaculata Paramonov, 1927: 4. Type locality: Armenia.
Conops kroeberi Paramonov, 1927: 4. Type locality: Armenia.
This species was listed by
Conops nubeculipennis Bezzi, 1901: 21. Type locality: Eritrea.
Conops atrogonius Séguy 1930: 134. Type locality: Morocco.
AF: Cameroon, Ethiopia, Guinea-Bissau, Mozambique, Nigeria, Saudi Arabia [as “South-western part”] (
Local distribution and dates of collection (Fig.
This species was listed by
Sphixosoma flavicauda Bigot, 1880: 149. Type locality: Iran (Northern Iran and Caucasus).
Conops euzonatus Bigot, 1887: 33. Type locality: Caucasus Mountains.
Conops superbus Röder, 1889: 6. Type locality: Lebanon (Beirut) [as “Beyrut, Syria (Asia minor)”].
Conops cypris Janssens, 1955: 2. Type locality: Cyprus (Akrotiri Bay, Yermdoyia & Erimi).
AF: Uganda. PA: Afghanistan, Armenia, Cyprus, Egypt, Greece, Iran, Israel, Lebanon, Syria, Turkey.
Local distribution and dates of collection (Fig.
Conops 4-fasciata De Geer, 1776: 261. Type locality: Presumably Sweden.
PA: Andorra, Austria, Belarus, Belgium, Bosnia and Herzegovina, Bulgaria, Croatia, Czech Republic, Denmark, Finland, France, Germany, Great Britain, Hungary, Iran, Ireland, Italy, Kazakhstan, Latvia, Lithuania, Netherlands, Norway, Poland, Romania, Russia, Saudi Arabia (first record), Slovakia, Spain, Sweden, Switzerland, Turkey, Ukraine.
Local distribution and dates of collection (Fig.
This species is recorded herein for the first time from Saudi Arabia.
Conops rufiventris Macquart, 1849: 474. Type locality: Algeria (Constantine).
Algeria, Egypt, Tunisia.
Local distribution and dates of collection (Fig.
Conops tomentosus Kröber, 1916: 45. Type locality: Turkmenistan.
AF: Saudi Arabia [as “South-western part”]. PA: Iraq, Tajikistan, Turkey, Turkmenistan.
Local distribution and dates of collection (Fig.
Physocephala Schiner, 1861: 138. Type species: Conops rufipes Fabricius, by original designation.
Conops antiqua Wiedemann, 1830: 239. Type locality: Egypt.
Conops arabica Macquart, 1851: 134. Type locality: Saudi Arabia (Jeddah).
Physocephala furax Becker, 1913: 612. Type locality: Iran.
Physocephala syriaca Kröber, 1915: 72. Type locality: Egypt (Cairo) & Palestinian West Bank (Jericho).
Physocephala obscurifacies Kröber, 1924: 67. Type locality: Egypt (Wadi Hoff).
AF: Egypt [as “Gebel Elba”], Saudi Arabia [as “South-western part”], United Arab Emirates, Yemen. PA: Afghanistan, Algeria, China, Cyprus, Egypt, Greece, Iran, Iraq, Israel, Italy, Jordan, Kazakhstan, Kyrgyzstan, Mongolia, Palestinian Territories, Russia, Saudi Arabia (Jeddah), Syria, Tunisia, Turkey, Turkmenistan.
Local distribution and dates of collection (Fig.
Conops chrysorrhoea Meigen, 1824: 128. Type locality: Austria.
Conops pallasi Meigen, 1824: 128. Type locality: Russia.
Conops serpylleti Zeller, 1842: 837. Type locality: Germany.
Physocephala zarudnyi Becker, 1913: 614. Type locality: Iran.
Physocephala truncata var. pseudomaculigera Kröber, 1915: 72. Type locality: Egypt & Tunisia.
AF: Saudi Arabia [as “South-western part”]. PA: Algeria, Armenia, Austria, Belgium, Bulgaria, China, Croatia, Czech, Egypt, France, Germany, Greece, Hungary, Iran, Italy, Kyrgyzstan, Lithuania, Mongolia, Morocco, Netherlands, Poland, Romania, Russia, Serbia, Slovakia, Spain, Switzerland, Turkey, Turkmenistan, Ukraine.
Local distribution and dates of collection (Fig.
Conops pusilla Meigen, 1824: 280. Type locality: Germany (Frankreich).
Conops lacera Meigen, 1824: 130. Type locality: Austria.
Conops pumila Macquart, 1835: 26. Type locality: France.
Conops persica Becker, 1913: 609. Type locality: Iran.
Conops punctithorax Becker, 1913: 611. Type locality: Iran.
PA: Afghanistan, China, Egypt, Europe (widespread), Kazakhstan, Mongolia, Morocco, Saudi Arabia, Syria, Tunisia, Turkmenistan.
Local distribution and dates of collection (Fig.
Conops variegata Meigen, 1824: 132. Type locality: Austria & France.
AF: Egypt [as “Gebel Elba”]. PA: Austria, Bulgaria, Egypt, France, Greece, Hungary, Iran, Kazakhstan, Kyrgyzstan, Mongolia, Romania, Russia, Saudi Arabia, Serbia, Turkey, Turkmenistan, Ukraine.
Local distribution and dates of collection (Fig.
Conops vittata Fabricius, 1794: 392. Type locality: Germany (Kiel).
Conops dorsalis Wiedemann in Meigen, 1824: 133. Type locality: Austria.
Conops solaeformis Gimmerthal, 1842: 672. Type locality: Latvia.
Conops semiatrata Costa, 1844: 89. Type locality: Italy.
Conops fraternus Loew, 1847: 18. Type locality: Turkey, Greece, Croatia & Italy.
Conops truncata Loew, 1847: 21. Type locality: Italy (Sicily).
Physocephala detecta Becker, 1913: 615. Type locality: Iran.
Physocephala vittata var. abdominalis Kröber, 1915: 57. Type locality: Turkey, Syria & Cyprus.
Physocephala vittata var. semirufa Kröber, 1915: 58. Type locality: Israel.
Physocephala truncata var. maculigera Kröber, 1915: 71. Type locality: Egypt, Tunisia, Algeria & Syria.
Physocephala vittata var. immaculata Kröber, 1939: 365. Type locality: Cyprus.
AF: Ethiopia, Kenya, Yemen. PA: Afghanistan, Algeria, Armenia, Austria, Belarus, Belgium, Bulgaria, China, Croatia, Cyprus, Czech Republic, Egypt, France, Germany, Hungary, Iran, Israel, Italy, Lithuania, Malta, Mongolia, Morocco, Netherlands, Norway, Palestinian Territories, Poland, Portugal, Romania, Russia, Serbia, Slovakia, Spain, Sweden, Switzerland, Syria, Tunisia, Turkey, Turkmenistan, Ukraine, Uzbekistan.
Local distribution and dates of collection (Fig.
Myopa Fabricius, 1775: 798. Type species: Conops buccata Linnaeus, by subsequent designation of
Myopa dorsalis Fabricius, 1794: 397. Type locality: Germany.
Myopa ferruginea Panzer, 1794: 24. Type locality: Germany.
Myopa grandis Meigen, 1804: 284. Type locality: Germany.
Myopa dorsalis var. nigrifacies Becker, 1922: 289. Type locality: Not given.
OR: India. PA: Albania, Algeria, Armenia, Austria, Belarus, Belgium, Bulgaria, China, Croatia, Czech Republic, Denmark, Egypt, Finland, France, Germany, Greece, Hungary, Iran, Italy, Kazakhstan, Luxembourg, Morocco, Netherlands, Norway, Poland, Romania, Russia, Slovakia, Spain, Sweden, Switzerland, Syria, Tunisia, Turkey, Turkmenistan, Ukraine.
Local distribution and dates of collection (Fig.
Myopa picta Panzer, 1798: 22. Type locality: Austria.
Myopa varia Wiedemann, 1830: 242. Type locality: Egypt.
Myopa meridionalis Macquart, 1835: 34. Type locality: Egypt.
AF: Ethiopia. OR: India, Pakistan. PA: Afghanistan, Austria, Belarus, Belgium, Bosnia and Herzegovina, Bulgaria, China, Croatia, Czech Republic, Egypt, France, Germany, Greece, Iran, Italy, Kyrgyzstan, Morocco, Poland, Romania, Russia, Slovakia, Spain, Switzerland, Tunisia, Turkey, Turkmenistan, Ukraine.
Local distribution and dates of collection (Fig.
Myopa stigma Meigen, 1824: 148. Type locality: Austria.
Myopa arabica Macquart 1851: 138. Type locality: Saudi Arabia (Jeddah).
AF: Ethiopia. PA: Afghanistan, Algeria, Austria, Belarus, Bulgaria, Cyprus, Czech Republic, France, Germany, Greece, Hungary, Italy, Kazakhstan, Morocco, Poland, Portugal, Romania, Russia, Saudi Arabia (Jeddah), Slovakia, Spain, Switzerland, Tunisia, Turkey, Ukraine.
Local distribution and dates of collection (Fig.
Conops testacea Linnaeus, 1767: 1006. Type locality: "Europa australis Ascanius".
Myopa longirostris Robineau-Desvoidy, 1830: 243. Type locality: Not given.
Myopa pictipennis Robineau-Desvoidy, 1830: 243. Type locality: Not given.
Myopa umbripennis Robineau-Desvoidy, 1830: 243. Type locality: Not given.
Myopa testacea var. japonica Kröber, 1916: 89. Type locality: Japan.
OR: India. PA: Afghanistan, Austria, Belarus, Belgium, Bulgaria, China, Croatia, Czech Republic, Denmark, Egypt, Finland, France, Germany, Great Britain, Greece, Hungary, Ireland, Italy, Japan, Kazakhstan, Kyrgyzstan, Lithuania, Luxembourg, Malta, Mongolia, Netherlands, Norway, Poland, Portugal, Romania, Russia, Slovakia, Spain, Sweden, Switzerland, Tunisia, Turkey, Turkmenistan, Ukraine.
Local distribution and dates of collection (Fig.
Thecophora Rondani, 1845: 15. Type species: Myopa atra Fabricius, by monotypy.
Myopa atra Fabricius, 1775: 799. Type locality: Denmark (Copenhagen).
Myopa annulata Fabricius, 1794: 399. Type locality: Italy.
Myopa maculata Meigen, 1804: 287. Type locality: France.
Myopa micans Meigen, 1804: 288. Type locality: France.
Myopa femorata Fabricius, 1805: 181. Type locality: Germany.
Occemya fulvifrons Robineau-Desvoidy, 1853: 134. Type locality: France.
Occemya meigeni Robineau-Desvoidy, 1853: 135. Type locality: France.
Occemya grisea Robineau-Desvoidy, 1853: 137. Type locality: France.
Occemya lamarckii Robineau-Desvoidy, 1853: 140. Type locality: France.
Occemya guerini Robineau-Desvoidy, 1853: 141. Type locality: France.
Occemya bigoti Robineau-Desvoidy, 1853: 142. Type locality: France.
Occemya brunipes Robineau-Desvoidy, 1853: 143. Type locality: France.
Occemya lucasi Robineau-Desvoidy, 1853: 144. Type locality: France.
AF: Saudi Arabia [as “South-western part”] (first record). OR: India, Sri Lanka. PA: Afghanistan, Albania, Algeria, Austria, Belgium, Bosnia and Herzegovina, Bulgaria, China, Croatia, Cyprus, Czech Republic, Denmark, Egypt, Finland, France, Germany, Great Britain, Greece, Hungary, Iran, Ireland, Italy, Japan, Kyrgyzstan, Lithuania, Luxembourg, Macedonia, Malta, Mongolia, Montenegro, Morocco, Netherlands, North Korea, Norway, Poland, Portugal, Romania, Russia, Serbia, Slovakia, Spain, Sweden, Switzerland, Tunisia, Turkey, Turkmenistan, Ukraine.
Local distribution and dates of collection (Fig.
This species (Fig.
Myopa fulvipes Robineau-Desvoidy, 1830: 246. Type locality: France (Paris).
Myopa sundewalli Zetterstedt, 1844: 942. Type locality: Sweden.
AF: Saudi Arabia [as “South-western part”] (El-Hawagry et al. 2017
Local distribution and dates of collection (Fig.
Zodion Latreille, 1797: 162. Type species: Myopa cinerea Fabricius, by Subsequent monotypy of
Myopa cinerea Fabricius, 1794: 399. Type locality: Italy.
Myopa notata Meigen, 1804: 288. Type locality: France.
Myopa tibialis Fabricius, 1805: 182. Type locality: Europe.
Zodion conopsoides Latreille, 1809: 337. Type locality: France.
Zodion pedicillatum Robineau-Desvoidy, 1830: 252. Type locality: Not given.
Zodion fuliginosum Robineau-Desvoidy, 1853: 156. Type locality: France.
Zodion fulvipes Robineau-Desvoidy, 1853: 157. Type locality: France.
Zodion fulvicorne Robineau-Desvoidy, 1853: 158. Type locality: France.
Zodion cinereum var. rubescens Szilády, 1925: 223. Type locality: Hungary.
Zodion cinereum var. kerteszi Szilády, 1926: 589. Type locality: Hungary.
AF: Central African Republic, Saudi Arabia [as “South-western part”] (
Local distribution and dates of collection (Fig.
Zodion erythrurum Rondani, 1865: 146. Type locality: Italy (Etruria).
Zodion pulchrum Loew, 1868: 384. Type locality: Italy.
Zodion vittipes Strobl, 1906: 331. Type locality: Spain.
PA: Algeria, Bulgaria, Egypt, France, Hungary, Israel, Italy, Kazakhstan, Kyrgyzstan, Morocco, Poland, Romania, Russia, Serbia, Spain, Tunisia, Turkey, Turkmenistan, Ukraine.
Local distribution and dates of collection (Fig.
The authors would like to extend their sincere appreciation to the Deanship of Scientific Research at King Saud University for funding this work [Research group No: RG-1437-009].
We are grateful to Dr. Jens-Hermann Stuke, Roter Weg 22, D-26789 Leer, Germany and Dr. David Clements, 7 Vista Rise, Llandaff, Cardiff, CF5 2SD, U.K. for providing critical assistance throughout this study and for supplying us with some relevant papers. We are also grateful to the research team in KSMA for collecting, mounting and photographing specimens.