The family Stratiomyidae in Egypt and Saudi Arabia (Diptera: Stratiomyoidea)

Magdi El-Hawagry; Hathal Al Dhafer; Mahmoud Abdel-Dayem; Martin Hauser

doi:10.3897/BDJ.9.e64212

Biodiversity Data Journal : Taxonomic Paper

PDF

Taxonomic Paper

The family Stratiomyidae in Egypt and Saudi Arabia (Diptera: Stratiomyoidea)

Magdi El-Hawagry^‡, Hathal Mohammed Al Dhafer^§, Mahmoud Abdel-Dayem^§, Martin Hauser^|

‡ Entomology Department, Faculty of Science, Cairo University, Giza, Egypt

§ King Saud University, College of Food and Agriculture Sciences, Riyadh, Saudi Arabia

| California Department of Food & Agriculture, Sacramento, United States of America

Corresponding author: Magdi El-Hawagry (elhawagry@cu.edu.eg)

Academic editor: Torsten Dikow

Received: 09 Feb 2021 | Accepted: 18 Mar 2021 | Published: 22 Mar 2021

This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.

Citation: El-Hawagry M, Al Dhafer HM, Abdel-Dayem M, Hauser M (2021) The family Stratiomyidae in Egypt and Saudi Arabia (Diptera: Stratiomyoidea). Biodiversity Data Journal 9: e64212. https://doi.org/10.3897/BDJ.9.e64212

Abstract

Background

This study systematically catalogues all known taxa of the family Stratiomyidae in Egypt and Saudi Arabia. It is one in a series of planned studies aiming to catalogue the whole order in both countries.

New information

Twenty species, belonging to seven genera and three subfamilies (Pachygastrinae, Stratiomyinae and Nemotelinae), are treated. One of these genera, Oplodontha and two species, Oplodontha pulchriceps Loew and Oxycera turcica Üstüner & Hasbenli, are recorded herein for the first time from Saudi Arabia. A lectotype for Nemotelus matrouhensis Mohammad et al., 2009 is designated. An updated classification, synonymies, type localities, world and local distributions, dates of collection and some coloured photographs are provided.

Keywords

soldier-flies, local distribution, dates of collection, new records

Introduction

The Stratiomyidae is a lower brachycerous Dipteran family, including more than 2650 species classified in 375 genera and 12 subfamilies worldwide (Woodley 2001). Flies of the family Stratiomyidae, commonly known as soldier-flies, are small to large, ranging from 2 to 28 mm in length, highly varied in shape and colouration, with wings hyaline to variously patterned or infuscated and some species likely mimic wasps and bees in their flight and body colour (Hauser 2008, Hauser et al. 2017b). These flies can be easily distinguished as adults by the following characters: radial veins grouped together anteriorly, ending before tip of the wing; costal vein (C) usually ending well before wing apex and discal cell (d) short, usually forming distinct short, often squarish cell at middle of the wing (Marshall et al. 2017).

Adult soldier-flies can be found sitting on foliage in sunny locations in damp forest habitats, near water or boggy areas and some flies can be found frequenting flowers, particularly those of the subfamily Stratiomyinae and some of the subfamily Clitellariinae (James 1981, Khaghaninia et al. 2015, Hauser et al. 2017b).

Immature stages of soldier-flies can be found in a variety of habitats. Larvae of the subfamilies Beridinae, Clitellariinae and Sarginae, which have not been represented in Egypt or Saudi Arabia, are usually associated with decaying organic matter, such as leaf litter and rotting fruits. However, larvae of the subfamilies Nemotelinae and Stratiomyinae are aquatic, occurring in slow moving or standing water in rivers, ponds, seepage areas and other transient water sources, with those of the genus Nemotelus Geoffroy, 1762 being remarkably tolerant to high levels of salinity. Larvae of the subfamily Pachygastrinae can be found under the bark of fallen trees (Rozkošný 1983, Hauser 2008, Hauser et al. 2017b).

Of the 12 subfamilies, seven are recorded in the Palaearctic Region with about 430 species (Woodley 2001). Of these, only three subfamilies (Pachygastrinae, Stratiomyinae and Nemotelinae) have been represented in Saudi Arabia and/or Egypt by seven genera comprising 20 species. One of these genera, Oplodontha and two species, Oxycera turcica Üstüner & Hasbenli and Oplodontha pulchriceps Loew, are recorded herein for the first time from Saudi Arabia. This is not surprising as no previous faunistic or systematic studies on the Stratiomyidae have been carried out in Saudi Arabia and only two species have been recorded from Saudi Arabia amidst two comprehensive checklists of the Saudi Arabian Diptera: El-Hawagry et al. (2017) [Aspidacantha atra Kertesz] and Abu-Zoherah et al. (1993) [Stratiomys deserticolor Lindner]. We think the overall number of Saudi Arabian taxa treated in the present study is still low and does not represent the real fauna of Stratiomyidae in this country and an extensive faunistic and systematic study on the Saudi Arabian Stratiomyidae is required. On the other hand, some previous studies have been carried out in Egypt to list the Stratiomyidae species (Steyskal and El-Bialy 1967) or to study the fauna and/or taxonomy of the family (Lindner 1930, Badrawy 2006, Mohammad et al. 2009), in addition to some miscellaneous studies which described some new species from Egypt (Olivier 1811, Becker 1902, Wiedemann 1830, Lindner 1925, Lindner 1930, Lindner 1937, Lindner 1974).

Egypt and Saudi Arabia are two neighbouring countries in the Middle East, separated by the Gulf of Aqaba and the Red Sea (Fig. 1). Egypt is a transcontinental country as its majority is located in the north-eastern corner of Africa, while its north-eastern extremity, Sinai Peninsula, is located in the south-western corner of Asia. On the other hand, Saudi Arabia is wholly located in the south-western corner of Asia. Both Egypt and Saudi Arabia are biogeographically located at the junction of the Palaearctic and the Afrotropical Regions. The faunal affiliation of the two countries is mainly Palaearctic. Exceptions are Gebel Elba, the south-eastern triangle of Egypt and the southwestern part of Saudi Arabia, south to the Tropic of Cancer, which are considered as having an Afrotropical faunal affiliation (Wallace 1876, Hölzel 1998, El-Hawagry and Gilbert 2014, El-Hawagry et al. 2017).

Figure 1.

A satellite map of Egypt and Saudi Arabia.

This study is one in a series of taxonomic studies on different Egyptian and Saudi Arabian dipteran taxa aiming to catalogue the whole order Diptera in the two countries (EL-Hawagry 2015, Al Dhafer and El-Hawagry 2016, El-Hawagry 2017, EL-Hawagry 2018, El-Hawagry et al. 2018, El-Hawagry and Al Dhafer 2019, El-Hawagry et al. 2019, El-Hawagry and Gilbert 2019, El-Hawagry et al. 2020b, El-Hawagry et al. 2020a, El-Hawagry et al. 2021, El-Hawagry and El-Azab 2019).

Materials and methods

Data sources. Data of the present study are obtained from three main sources: 1. Specimens preserved in the Egyptian and Saudi Arabian insect collections and museums, namely: Efflatoun Bey’s collection, Cairo University, Egypt (EFC); Collection of Entomological Society of Egypt (ESEC) and King Saud University Museum of Arthropods, Saudi Arabia (KSMA). 2. Previous studies on the Stratiomyidae in Egypt and Saudi Arabia. 3. Specimens collected by the authors and their co-workers, especially from Saudi Arabia using both Malaise traps and aerial nets. A great deal of faunistic and taxonomic information, including type species, type localities, Old World synonymies, world and local distributions and collection dates were obtained from relevant literature as well. These sources are listed in the following subsections.

Study area. Egypt and Saudi Arabia, the study area, are two neighbouring countries in the Middle East, separated by the Red Sea and the Gulf of Aqaba. They constitute a part of the Great Desert Belt, mainly with an arid desert climate characterised by hot summer and a mild winter (Alsharhan et al. 2001, Nasser et al. 2019).

Ecologists divide Egypt into eight ecological zones: the Coastal Strip, Lower Nile Valley & Delta, Upper Nile Valley, Fayoum, Eastern Desert, Western Desert, Sinai and Gebel Elba (Fig. 1, El-Hawagry and Gilbert 2014). These ecological zones are adopted in the present study in the sections of localities and dates of collection. However, Saudi Arabia is not divided into ecological zones, so ecologists usually adopt the administrative divisions instead. These divisions (also called emirates, regions or provinces) are adopted in the present study, namely: Makkah, Riyadh, Eastern Province, Asir, Jazan, Al-Madinah, Al-Qaseem, Tabuk, Hail, Najran, Al-Jawf, Al-Baha and Northern Frontier (El-Hawagry and Al Dhafer 2019).

Classification. The classification and arrangement of taxa in the present study basically follows that used in Woodley (2001) and subsequent updates by Hauser et al. (2017a).

World distribution. Sources of world distribution of each stratiomyid species are given between square brackets at the end of the list of countries.

Local distribution and dates of collection. Localities within each Egyptian ecological zone or Saudi Arabian administrative region are arranged alphabetically and written after a colon following each zone or region followed by the dates of collection between parentheses; for example, "Coastal Strip: Abu-Kir, Cleopatra, Dekhela (April to September)" and “Tabuk: Tabuk City (May)”. Sources for this distribution are given between square brackets at the end of the section. Coordinates of all Egyptian and Saudi Arabian localities of the family Stratiomyidae are listed (Table 1). Distribution maps of species were made using SimpleMappr (Shorthouse 2010).

Table 1.

Download as

CSV

XLSX

A gazetteer of Egyptian and Saudi Arabian localities of the family Stratiomyidae.

Country	Locality	Governorate	Ecological zone or Region	Latitude (N)	Longitude (E)
Egypt	Abu-Kir	Alexandria	Coastal Strip	31.22429	33.8256
	Abu-Rawash	Giza	Lower Nile Valley & Delta	30.0438	42.4101
	Abu-Zaabal	Al-Qalyubia	Lower Nile Valley & Delta	30.24098	42.4089
	Alexandria	Alexandria	Coastal Strip	31.2129	42.4089
	Behaira	Behaira	Lower Nile Valley & Delta	30.62189	42.4102
	Benha	Al-Qalyubia	Lower Nile Valley & Delta	30.46572	42.4089
	Beni Sweif	Beni Sweif	Lower Nile Valley & Delta	29.07788	42.4089
	Birket Qaroun	Fayoum	Fayoum	29.40879	42.4124
	Cleopatra	Alexandria	Coastal Strip	31.22022	42.4101
	Dakhla Oasis	New Valley	Western Desert	25.5000	41.3115
	Damietta	Damietta	Lower Nile Valley & Delta	31.34595	41.3115
	Dekhela	Alexandria	Coastal Strip	31.12098	41.3101
	Ein Moussa	South Sinai	Sinai	29.8667	41.3101
	El-Alag	Al-Qalyubia	Lower Nile Valley & Delta	30.18009	42.8348
	El-Arish	North Sinai	Sinai	31.1244	39.8749
	El-Baragil	Giza	Lower Nile Valley & Delta	30.07673	41.4107
	El-Ferdan	Ismailia	Eastern Desert	30.65760	36.6297
	El-Gebel El-Asfar	Al-Qalyubia	Lower Nile Valley & Delta	29.1689	34.1412
	El-Kantara	Ismailia	Eastern Desert	30.79392	34.3836
	El-Marg	Al-Qalyubia	Lower Nile Valley & Delta	31.0667	30.5833
	El-Siala	Alexandria	Coastal Strip	31.20849	30.8180
	El-Tour	South Sinai	Sinai	28.24024	29.1667
	Ezbet El-Nakhl	Al-Qalyubia	Lower Nile Valley & Delta	31.1111	32.6500
	Fayed	Ismailia	Eastern Desert	30.32382	29.8805
	Fayoum	Fayoum	Fayoum	29.32061	32.3008
	Gebel Elba	Red Sea	Gebel Elba	22.2008	30.2167
	Gezeira	Cairo	Lower Nile Valley & Delta	30.04596	34.3836
	Girza	Fayoum	Fayoum	29.49968	34.1412
	Giza	Giza	Lower Nile Valley & Delta	30.01350	31.3333
	Giza-Fayoum Road	Giza	Western Desert	29.85176	31.18121
	Helwan	Cairo	Lower Nile Valley & Delta	29.8500	31.22435
	Hurghada	Red Sea	Eastern Desert	27.2337	30.5833
	Ismailia	Ismailia	Eastern Desert	30.59428	30.8180
	Kerdassa	Giza	Lower Nile Valley & Delta	30.0297	30.48755
	Kharga Oasis	New Valley	Western Desert	25.2500	31.6317
	Khosous	Cairo	Lower Nile Valley & Delta	30.15957	31.1117
	Kosseimah	North Sinai	Sinai	30.90307	30.9898
	Maadi	Cairo	Lower Nile Valley & Delta	29.95772	31.3125
	Mariout	Alexandria	Coastal Strip	31.0172	31.3531
	Max	Alexandria	Coastal Strip	31.1636	30.2167
	Mersa Matrouh	Matrouh	Coastal Strip	29.5696	30.6468
	Moharram Bey	Alexandria	Coastal Strip	31.17796	31.10713
	Moweileh	South Sinai	Sinai	30.3924	30.5833
	Nazla	Fayoum	Fayoum	29.29792	30.5833
	Nubar Bey	Alexandria	Coastal Strip	31.18079	31.2505
	Nuzha (Alex.)	Alexandria	Coastal Strip	31.2001	31.3333
	Quisna	Menofiya	Lower Nile Valley & Delta	30.53514	31.3333
	Ramleh	Alexandria	Coastal Strip	31.2279	30.8180
	Sandoub	Dakahlyia	Lower Nile Valley & Delta	31.00782	30.2167
	Sherbin	Dakahlyia	Lower Nile Valley & Delta	31.19461	31.3333
	Shubra	Al-Qalyoubia	Lower Nile Valley & Delta	30.1012	30.5833
	Siwa Oasis	Matrouh	Western Desert	29.20427	26.4194
	Suez	Suez	Eastern Desert	29.95278	32.3008
	Tanta	Gharbia	Lower Nile Valley & Delta	30.75725	32.3008
	Wadi El-Lega	South Sinai	Sinai	28.5469	32.3008
	Wadi El-Natroun	Behaira	Western Desert	30.3814	29.8157
	Wadi Hebran	South Sinai	Sinai	28.40225	31.3521
	Wadi Hoff	Cairo	Eastern Desert	29.8821	32.3008
	Wadi Watir	South Sinai	Sinai	29.02147	31.3110
	Zaranik	North Sinai	Sinai	31.10345	30.3441
Saudi Arabia	Al-Mekhwa	Al-Mekhwa	Al-Baha	19.798133	30.3441
	Dawmat Al-Jandal	Dawmat Al-Gandal	Al-Jwaf	29.809552	30.8180
	Hassan Ameen farm	Tabouk	Tabouk	28.36661	26.4194
	Jabal Shada al-A’la Nature Reserve	Al-Mekhwa	Al-Baha	19.8429	29.7600
	Jazan	Jazan	Jazan	16.9595	29.7600
	Raydah Nature Reserve	Abha	Asir	18.20525	30.8180

Abbreviations used:

AF, Afrotropical
CSCA, California State Collection of Arthropods, Sacramento, California, USA
EFC, Efflatoun Bey’s collection, Department of Entomology, Faculty of Science, Cairo University, Egypt
ESEC, Collection of Entomological Society of Egypt, Cairo, Egypt
KSMA, King Saud University Museum of Arthropods, Riyadh, Saudi Arabia
MCCB, Museum of Community College, Al-Baha University, Saudi Arabia
OR, Oriental
PA, Palearctic

The Catalogue

Family Stratiomyidae

Subfamily Stratiomyiinae

Genus Aspidacantha Kertsz, 1916

GBIF https://www.gbif.org/species/1578168

Nomenclature:

Aspidacantha Kertész, 1916: 154. Type species: Aspidacantha atra Kertész, by original designation.

Aspidacantha atra Kertsz, 1916

GBIF https://www.gbif.org/species/1578171

Nomenclature:

Aspidacantha atra Kertész, 1916: 155. Type locality: Eritrea (Assab) [Assab belongs now to Eritrea not to Ethiopia as written in world catalogues].

Materials Download as CSV

scientificName:
Aspidacantha atra
; taxonID:
https://www.gbif.org/species/1578171
; country:
Egypt
; locality:
Hurghada
; decimalLatitude:
27.23370
; decimalLongitude:
33.8256
; samplingProtocol:
Light trap
; eventDate:
9/11-22/1994
; sex:
1 female
; lifeStage:
Adult
; recordedBy:
H. Pohl
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
CSCA
scientificName:
Aspidacantha atra
; taxonID:
https://www.gbif.org/species/1578171
; country:
Saudi Arabia
; locality:
Raydah
; decimalLatitude:
18.20525
; decimalLongitude:
42.4101
; samplingProtocol:
Sweeping
; eventDate:
06-07-2014
; lifeStage:
Adult
; recordedBy:
Aldhafer H.M. et al.
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
KSMA
scientificName:
Aspidacantha atra
; taxonID:
https://www.gbif.org/species/1578171
; country:
Saudi Arabia
; locality:
Raydah
; decimalLatitude:
18.201583
; decimalLongitude:
42.4089
; samplingProtocol:
Light trap
; eventDate:
02-21-2014
; lifeStage:
Adult
; recordedBy:
Aldhafer H.M. et al.
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
KSMA
scientificName:
Aspidacantha atra
; taxonID:
https://www.gbif.org/species/1578171
; country:
Saudi Arabia
; locality:
Raydah
; decimalLatitude:
18.201583
; decimalLongitude:
42.4089
; samplingProtocol:
Light trap
; eventDate:
07-31-2015
; lifeStage:
Adult
; recordedBy:
Aldhafer H.M. et al.
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
KSMA
scientificName:
Aspidacantha atra
; taxonID:
https://www.gbif.org/species/1578171
; country:
Saudi Arabia
; locality:
Raydah
; decimalLatitude:
18.20525
; decimalLongitude:
42.4102
; samplingProtocol:
Sweeping
; eventDate:
09-05-2015
; lifeStage:
Adult
; recordedBy:
Aldhafer H.M. et al.
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
KSMA
scientificName:
Aspidacantha atra
; taxonID:
https://www.gbif.org/species/1578171
; country:
Saudi Arabia
; locality:
Raydah
; decimalLatitude:
18.201583
; decimalLongitude:
42.4089
; samplingProtocol:
Light trap
; eventDate:
09-05-2015
; lifeStage:
Adult
; recordedBy:
Aldhafer H.M. et al.
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
KSMA
scientificName:
Aspidacantha atra
; taxonID:
https://www.gbif.org/species/1578171
; country:
Saudi Arabia
; locality:
Raydah
; decimalLatitude:
18.201583
; decimalLongitude:
42.4089
; samplingProtocol:
Malaise trap
; eventDate:
09-05-2015
; lifeStage:
Adult
; recordedBy:
Aldhafer H.M. et al.
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
KSMA
scientificName:
Aspidacantha atra
; taxonID:
https://www.gbif.org/species/1578171
; country:
Saudi Arabia
; locality:
Raydah
; decimalLatitude:
18.204417
; decimalLongitude:
42.4124
; samplingProtocol:
Malaise trap
; eventDate:
02-20-2014
; lifeStage:
Adult
; recordedBy:
Aldhafer H.M. et al.
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
KSMA
scientificName:
Aspidacantha atra
; taxonID:
https://www.gbif.org/species/1578171
; country:
Saudi Arabia
; locality:
Raydah
; decimalLatitude:
18.20525
; decimalLongitude:
42.4101
; samplingProtocol:
Malaise trap
; eventDate:
06-07-2014
; lifeStage:
Adult
; recordedBy:
Aldhafer H.M. et al.
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
KSMA

Distribution:

AF: Democratic Republic of Congo, Eritrea, Ethiopia, Saudi Arabia [as “south western part”], Tanzania, Uganda, United Arab Emirates, Zimbabwe. PA: Egypt, Israel, Russia, Turkmenistan. [Sources: Lindner (1936), Woodley (2001), Hauser (2008) and El-Hawagry et al. (2017)]

Local distribution and dates of collection (Fig. 2): EGYPT: Eastern Desert: Hurghada (September). Western Desert: Siwa Oasis (August). [Sources: Lindner (1930) and museum material]. SAUDI ARABIA: Asir: Raydah Nature Reserve (February to December). [Sources: El-Hawagry et al. (2017) and collected material]

Figure 2.

Distribution map of Aspidacantha atra Kertész and Sternobrithes sp.

Genus Sternobrithes Loew, 1857

GBIF https://www.gbif.org/species/1576881

Nomenclature:

Sternobrithes Loew, 1857: 264. Type species Sternobrithes tumidus Loew, by monotypy.

Sternobrithes sp.

GBIF https://www.gbif.org/species/1576881

Materials Download as CSV

scientificName:
Sternobrithes sp.
; taxonID:
https://www.gbif.org/species/1576881
; country:
Saudi Arabia
; locality:
Shada
; decimalLatitude:
19.8429
; decimalLongitude:
41.3115
; samplingProtocol:
Malaise trap
; eventDate:
11-15-2015
; lifeStage:
Adult
; recordedBy:
Aldhafer H.M. et al.
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
KSMA
scientificName:
Sternobrithes sp.
; taxonID:
https://www.gbif.org/species/1576881
; country:
Saudi Arabia
; locality:
Shada
; decimalLatitude:
19.8429
; decimalLongitude:
41.3115
; samplingProtocol:
Malaise trap
; eventDate:
05-05-2015
; lifeStage:
Adult
; recordedBy:
Aldhafer H.M. et al.
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
KSMA
scientificName:
Sternobrithes sp.
; taxonID:
https://www.gbif.org/species/1576881
; country:
Saudi Arabia
; locality:
Shada
; decimalLatitude:
19.8388
; decimalLongitude:
41.3101
; samplingProtocol:
Malaise trap
; eventDate:
11-15-2015
; lifeStage:
Adult
; recordedBy:
Aldhafer H.M. et al.
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
KSMA
scientificName:
Sternobrithes sp.
; taxonID:
https://www.gbif.org/species/1576881
; country:
Saudi Arabia
; locality:
Shada
; decimalLatitude:
19.8388
; decimalLongitude:
41.3101
; samplingProtocol:
Light trap
; eventDate:
09-02-2015
; lifeStage:
Adult
; recordedBy:
Aldhafer H.M. et al.
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
KSMA

Distribution:

The genus Sternobrithes is widely distributed all over the African continent (Botswana, Burundi, Cameroon, Cape Verde Islands, Democratic Republic of the Congo, Ethiopia, Ghana, Guinea, Ivory Coast, Kenya, Liberia, Malawi, Mozambique, Namibia, Nigeria, Senegal, Sierra Leone, Somalia, South Africa, Tanzania, Togo, Uganda and Zimbabwe. [Sources: Woodley (2001)]

Distribution of Sternobrithes sp.: AF: Kenya, Saudi Arabia [as “south western part”], Yemen.

Local distribution and dates of collection (Fig. 2): SAUDI ARABIA: Al-Baha: Jabal Shada al-A’la Nature Reserve (May to November). Asir: Raydah Nature Reserve (June). [Sources: El-Hawagry et al. (2017) and collected material]

Notes:

El-Hawagry et al. (2017) reported this species as Sternobrithes sp. and further research indicated that this is an undescribed species (Fig. 3). As we do not have a male specimen available, we are not yet describing this species here. According the generic key in Hauser et al. (2017b), the difference between Argyrobrithes and Sternobrithes is the colour of the last antennal segment, but further studies indicate that Argyrobrithes is characterised by the dichoptic male eyes and a very long last antennal segment. The two genera are widely distributed through Africa and the Oriental Region and a more thorough revision is needed to clarify the limits of the genera and which species should be included in each genus. Material of this undescribed species are known from Yemen and Kenya.

Figure 3.

Sternobrithes sp.: A. female habitus, lateral; B. same, dorsal.

Subfamily Stratiomyinae

Tribe Oxycerini

Genus Oxycera Meigen, 1803

GBIF https://www.gbif.org/species/1577931

Nomenclature:

Oxycera Meigen, 1803: 265. Type species: Musca hypoleon Linnaeus [= Musca trilineata Linnaeus], by designation of Latreille (1810).

Oxycera orientalis (Lindner, 1974)

GBIF https://www.gbif.org/species/5078177

Nomenclature:

Heraclina orientalis Lindner, 1974: 103. Type locality: Egypt (Sinai Mountains: Wadi Watir) [Wadi Watir is located in Sinai, Egypt, not in Israel as given in world catalogues].

Material Download as CSV

scientificName:
Oxycera orientalis
; taxonID:
https://www.gbif.org/species/5078177
; country:
Egypt
; locality:
Wadi Watir
; decimalLatitude:
29.02147
; decimalLongitude:
34.6731
; samplingProtocol:
Not given
; eventDate:
09-06-1941
; sex:
1 male
; lifeStage:
Adult
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC

Distribution:

PA: Egypt. [Sources: Furth (1983), Woodley (2001) and Badrawy (2006)]

Local distribution and dates of collection (Fig. 4): EGYPT: Sinai: Wadi El-Lega, Wadi Watir (Ain Fortaga) (September to November). [Sources: original description (Lindner 1974) and museum material]

Figure 4.

Distribution map of Oxycera orientalis (Lindner) and Oxycera turcica Üstüner & Hasbenli.

Oxycera turcica Üstüner & Hasbenli, 2005

GBIF https://www.gbif.org/species/5078139

Nomenclature:

Oxycera turcica Üstüner & Hasbenli, 2005: 163. Type locality: Turkey (Sivas: Sarkisla, Karacaören Village).

Material Download as CSV

scientificName:
Oxycera turcica
; taxonID:
https://www.gbif.org/species/5078139
; country:
Saudi Arabia
; locality:
Al-Mekhwa
; decimalLatitude:
19.798133
; decimalLongitude:
41.4107
; samplingProtocol:
Malaise trap
; eventDate:
03-15-2012
; lifeStage:
Adult
; recordedBy:
El-Hawagry
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
MCCB

Distribution:

PA: Saudi Arabia, Turkey. [Sources: Woodley (2011) and collected material]

Local distribution and dates of collection (Fig. 4): SAUDI ARABIA: Al-Baha: Al-Mekhwa (March). [Source: collected material]

Notes:

This species is recorded herein for the first time from Saudi Arabia.

Tribe Stratiomyini

Genus Odontomyia Meigen, 1803

GBIF https://www.gbif.org/species/1580809

Nomenclature:

Odontomyia Meigen, 1803: 265. Type species: Musca hydroleon Linnaeus, by designation of Westwood (1840).

Odontomyia angulata (Panzer, 1798)

GBIF https://www.gbif.org/species/1581104

Nomenclature:

Stratiomys angulata Panzer, 1798: 19. Type locality: Germany (Nürnberg).

Stratiomys vulpina Panzer, 1798: 24. Type locality: Germany (Nürnberg).

Stratiomys hydropota Meigen, 1822: 147. Type locality: Europe.

Odontomyia latifaciata Macquart, 1834: 248. Type locality: France.

Stratiomys brevicornis Loew, 1840: 25. Type locality: Poland (Poznañ).

Stratiomys brevicornis Loew, 1840: 557. Type locality: Poland (Poznañ). Preoccupied, primary homonym of Stratiomys brevicornis Loew, 1840.

Stratiomys ruficornis Zetterstedt, 1842: 139. Type locality: Sweden (Gotland: Ejsta). Preoccupied, primary homonym of Stratiomyia ruficornis Macquart, 1838.

Odontomyia hydrophila Loew, 1846: 486. Type locality: Turkey (Makri) and Italy (Sicily: Siracusa).

Eulalia latifasciata Kertész, 1908: 74. Unjustified emendation of Odontomyia latifaciata Macquart.

Distribution:

PA: Afghanistan, Albania, Algeria, Austria, Belgium, Bulgaria, China, Czech Republic, Denmark, Egypt, England, Estonia, Finland, France, Germany, Greece, Hungary, Iran, Israel, Italy including Sardinia, Kazakhstan, Morocco, Netherlands, Poland, Romania, Russia, Slovakia, Spain, Sweden, Switzerland, Turkey, UAE, former Yugoslavia. [Sources: Woodley (2001), Mason et al. (2009), Hauser (2014) and Yimlahi et al. (2017)]

Local distribution and dates of collection: Unknown.

Notes:

This species was listed by Steyskal and El-Bialy (1967), Woodley (2001) and Badrawy (2006) as recorded from Egypt, but no specimens have been collected or preserved in Egyptian museums to confirm these records.

Odontomyia megacephala Olivier, 1811

GBIF https://www.gbif.org/species/7863197

Nomenclature:

Odontomyia megacephala Olivier, 1811: 432. Type locality: Egypt (borders of the Nile [as “bords du Nil”]).

Materials Download as CSV

scientificName:
Odontomyia megacephala
; taxonID:
https://www.gbif.org/species/7863197
; country:
Egypt
; locality:
Moweileh
; decimalLatitude:
30.3924
; decimalLongitude:
34.1412
; samplingProtocol:
Not given
; eventDate:
Sept.1924
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
ESEC
scientificName:
Odontomyia megacephala
; taxonID:
https://www.gbif.org/species/7863197
; country:
Egypt
; locality:
Kosseima
; decimalLatitude:
30.90307
; decimalLongitude:
34.3836
; samplingProtocol:
Not given
; eventDate:
09-10-1924
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
ESEC
scientificName:
Odontomyia megacephala
; taxonID:
https://www.gbif.org/species/7863197
; country:
Egypt
; locality:
Kharga Oasis
; decimalLatitude:
25.25
; decimalLongitude:
30.5833
; samplingProtocol:
Not given
; eventDate:
Oct.1924
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
ESEC
scientificName:
Odontomyia megacephala
; taxonID:
https://www.gbif.org/species/7863197
; country:
Egypt
; locality:
Fayoum
; decimalLatitude:
29.32061
; decimalLongitude:
30.818
; samplingProtocol:
Not given
; eventDate:
Sept.1924
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
ESEC
scientificName:
Odontomyia megacephala
; taxonID:
https://www.gbif.org/species/7863197
; country:
Egypt
; locality:
Alexandria
; decimalLatitude:
31.2129
; decimalLongitude:
29.9726
; samplingProtocol:
Not given
; eventDate:
07-04-1920
; sex:
1 female
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Odontomyia megacephala
; taxonID:
https://www.gbif.org/species/7863197
; country:
Egypt
; locality:
Nuzha
; decimalLatitude:
31.2001
; decimalLongitude:
29.9436
; samplingProtocol:
Not given
; eventDate:
05-07-1921
; sex:
1 female
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Odontomyia megacephala
; taxonID:
https://www.gbif.org/species/7863197
; country:
Egypt
; locality:
Nuzha
; decimalLatitude:
31.2001
; decimalLongitude:
29.9436
; samplingProtocol:
Not given
; eventDate:
07-18-1921
; sex:
1 male
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Odontomyia megacephala
; taxonID:
https://www.gbif.org/species/7863197
; country:
Egypt
; locality:
Cleopatra
; decimalLatitude:
31.22021
; decimalLongitude:
29.9348
; samplingProtocol:
Not given
; eventDate:
07-10-1921
; sex:
2 females
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Odontomyia megacephala
; taxonID:
https://www.gbif.org/species/7863197
; country:
Egypt
; locality:
Cleopatra
; decimalLatitude:
31.22021
; decimalLongitude:
29.9348
; samplingProtocol:
Not given
; eventDate:
07-22-1921
; sex:
1 male
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Odontomyia megacephala
; taxonID:
https://www.gbif.org/species/7863197
; country:
Egypt
; locality:
Cleopatra
; decimalLatitude:
31.22021
; decimalLongitude:
29.9348
; samplingProtocol:
Not given
; eventDate:
08-01-1921
; sex:
1 male
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Odontomyia megacephala
; taxonID:
https://www.gbif.org/species/7863197
; country:
Egypt
; locality:
Fayed
; decimalLatitude:
30.32382
; decimalLongitude:
32.3008
; samplingProtocol:
Not given
; eventDate:
09-24-1925
; sex:
1 male
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Odontomyia megacephala
; taxonID:
https://www.gbif.org/species/7863197
; country:
Egypt
; locality:
Fayed
; decimalLatitude:
30.32382
; decimalLongitude:
32.3008
; samplingProtocol:
Not given
; eventDate:
10-20-1925
; sex:
1 female
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Odontomyia megacephala
; taxonID:
https://www.gbif.org/species/7863197
; country:
Egypt
; locality:
Wadi El-Natroun
; decimalLatitude:
30.3814
; decimalLongitude:
30.3441
; samplingProtocol:
Not given
; eventDate:
08-06-1929
; sex:
2 females
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC

Distribution:

PA: Egypt. [Sources: Lindner (1936), Woodley (2001) and Badrawy (2006)]

Local distribution and dates of collection (Fig. 5): EGYPT: Coastal Strip: Alexandria, Cleopatra, Mariout, Max, Nuzha (May and August). Eastern Desert: Fayed (September). Fayoum: Fayoum City (September). Lower Nile Valley: El-Baragil (April). Sinai: Kosseimah, Moweileh (September). Western Desert: Kharga Oasis, Wadi El-Natroun (August to October). [Sources: original description (Olivier 1811), Badrawy (2006) and museum material]

Figure 5.

Distribution map of Odontomyia megacephala Olivier and Odontomyia xanthopus Bezzi.

Odontomyia xanthopus Bezzi, 1906

GBIF https://www.gbif.org/species/1581000

Nomenclature:

Odontomyia xanthopus Bezzi, 1906: 225. Type locality: Eritrea (Adi Ugri) [Adi Ugri belongs now to Eritrea not to Ethiopia as written in world catalogues].

Material Download as CSV

scientificName:
Odontomyia xanthopus
; taxonID:
https://www.gbif.org/species/1581000
; country:
Egypt
; locality:
Gebel Ela
; decimalLatitude:
22.2008
; decimalLongitude:
36.3331
; samplingProtocol:
Not given
; eventDate:
Apr 1928
; sex:
1 male, 3 females
; lifeStage:
Adult
; recordedBy:
M.Tewfik
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC

Distribution:

AF: Eritrea, Malawi, Oman, Yemen (this is the first time this species has been recorded from Oman and Yemen). PA: Egypt, Israel. [Sources: original description (Bezzi 1906), Woodley (2001) and collected material from Yemen and Oman]

Local distribution and dates of collection (Fig. 5): EGYPT: Gebel Elba: ? (April). Sinai: El-Tour, Wadi Hebran (May to July). [Source: Lindner (1930), Lindner (1974) and museum material]

Genus Oplodontha Rondani, 1863

GBIF https://www.gbif.org/species/1577328

Nomenclature:

Oplodontha Rondani, 1863: 78. Type species: Stratiomys viridula Fabricius, by original designation.

Oplodontha minuta Fabricius, 1794

GBIF https://www.gbif.org/species/1577386

Nomenclature:

Stratiomys minuta Fabricius, 1794: 268. Type locality: India (Tranquebar).

Nemotelus pusillus Fabricius, 1794: 271. Type locality: India (Tranquebar).

Musca minutior Turton, 1801: 631. New name for Stratiomys minuta Fabricius, 1794.

Musca minor Turton, 1801: 655. New name for Nemotelus pusillus Fabricius, 1794.

Oxycera indica Brunetti, 1907: 119. Type locality: India (Uttar Pradesh: Bareilly).

Odontomyia incompleta Brunetti, 1907: 128. Nomen nudum.

Odontomyia ochracea Brunetti, 1907: 129. Type locality: India (Calcutta).

Odontomyia submutica Brunetti, 1907: 130. Type locality: India (Siliguri, Calcutta and Tollygunge).

Eulalia oasina Lindner, 1925: 150. Type locality: Egypt (Khârga Oasis and Dakhla Oasis).

Materials Download as CSV

scientificName:
Oplodontha minuta
; taxonID:
https://www.gbif.org/species/1577386
; country:
Egypt
; locality:
Dakhla Oasis
; decimalLatitude:
25.5
; decimalLongitude:
29.1667
; samplingProtocol:
Not given
; eventDate:
05-13-2018
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
ESEC
scientificName:
Oplodontha minuta
; taxonID:
https://www.gbif.org/species/1577386
; country:
Egypt
; locality:
Ein Moussa
; decimalLatitude:
29.8667
; decimalLongitude:
32.65
; samplingProtocol:
Not given
; eventDate:
09-20-1924
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
ESEC
scientificName:
as Eulalia oasina
; taxonID:
https://www.gbif.org/species/1577386
; country:
Egypt
; locality:
Kharga Oasis
; decimalLatitude:
25.25
; decimalLongitude:
30.5833
; samplingProtocol:
Not given
; eventDate:
05-10-1918
; sex:
1 male
; lifeStage:
Adult
; recordedBy:
Storey
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
as Eulalia oasina
; taxonID:
https://www.gbif.org/species/1577386
; country:
Egypt
; locality:
Kharga Oasis
; decimalLatitude:
25.25
; decimalLongitude:
30.5833
; samplingProtocol:
Not given
; eventDate:
06-12-1918
; sex:
1 female
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC

Distribution:

AF: Socotra Island, United Arab Emirates, Yemen. OR: India, Sri Lanka. PA: Afghanistan, Egypt, Israel. [Sources: original description of O. oasina (Lindner 1925), Woodley (2001) and Tkoč and Rozkošný (2014)]

Local distribution and dates of collection (Fig. 6): EGYPT: Eastern Desert: Ein Moussa (September). Western Desert: Dakhla Oasis, Kharga Oasis (May and June). [Sources: Lindner (1925), Lindner (1930) and museum material]

Figure 6.

Distribution map of Oplodontha minuta Fabricius and Oplodontha pulchriceps Loew.

Oplodontha pulchriceps Loew, 1858

GBIF https://www.gbif.org/species/1577339

Nomenclature:

Odontomyia pulchriceps Loew, 1858: 335. Type locality: South Africa (Cape of Good Hope).

Odontomyia pulchriceps Loew, 1860: 80. Type locality: South Africa (Cape of Good Hope). Preoccupied, primary homonym of Odontomyia pulchriceps Loew, 1858.

Hoplodonta madagascariensis Lindner, 1936: 42. Type locality: Madagascar (Bekily).

Materials Download as CSV

scientificName:
Oplodontha pulchriceps
; taxonID:
https://www.gbif.org/species/1577339
; country:
Saudi Arabia
; locality:
Jazan
; decimalLatitude:
16.9595
; decimalLongitude:
42.8348
; samplingProtocol:
Malaise trap
; eventDate:
05-11-2018
; sex:
1 male
; lifeStage:
Adult
; recordedBy:
Aldhafer H.M. et al.
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
KSMA
scientificName:
Oplodontha pulchriceps
; taxonID:
https://www.gbif.org/species/1577339
; country:
Saudi Arabia
; locality:
Dawmat Al-Jandal
; decimalLatitude:
29.809552
; decimalLongitude:
39.8749
; samplingProtocol:
Malaise trap
; eventDate:
05-26-2018
; sex:
1 male
; lifeStage:
Adult
; recordedBy:
Aldhafer H.M. et al.
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
KSMA
scientificName:
Oplodontha pulchriceps
; taxonID:
https://www.gbif.org/species/1577339
; country:
Saudi Arabia
; locality:
Hassan Ameen farm
; decimalLatitude:
28.36661
; decimalLongitude:
36.6297
; samplingProtocol:
Sweeping
; eventDate:
05-27-2018
; sex:
1 female
; lifeStage:
Adult
; recordedBy:
Aldhafer H.M. et al.
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
KSMA

Distribution:

AF: Democratic Republic of Congo, Lesotho, Madagascar, Mozambique, Saudi Arabia [as “south western part"], South Africa, United Arab Emirates. PA: Israel, Saudi Arabia [Jawf and Tabouk]. [Sources: Woodley (2001), Hauser (2008) and collected material]

Local distribution and dates of collection (Fig. 6): SAUDI ARABIA: Al-Jawf: Dawmat Al-Jandal (May). Jazan: Jazan (May). Tabouk: Hassan Ameen farm (May). [Source: collected material]

Notes:

This species (Fig. 7) and the genus Oplodontha are recorded herein for the first time from Saudi Arabia. It looks similar to O. minuta Fabricius; however, this case requires more clarification in a future study.

Figure 7.

Oplodontha pulchriceps Loew: A. female habitus, lateral; B. same, dorsal; C. male habitus, lateral.

Genus Stratiomys Geoffroy, 1762

GBIF https://www.gbif.org/species/1501902

Nomenclature:

Stratiomys Geoffroy, 1762: 449, 475. Type species: Musca chamaeleon Linnaeus, by designation of I.C.Z.N. (1957).

Stratiomys cenisia Meigen, 1822

GBIF https://www.gbif.org/species/1577708

Nomenclature:

Stratiomys cenisia Meigen, 1822: 136. Type locality: France (Mont Cenis).

Stratiomys flaviventris Loew, 1846: 464. Type locality: Italy (Sicily: Siracusa).

Stratiomyia ahngeri Pleske, 1901: 364. Type locality: “Transcaspian Region” [probably = Turkmenistan].

Stratiomyia cypria Pleske, 1902: 413. Type locality: Cyprus (Lárnax).

Stratiomyia kervillei Villeneuve, 1911: 4. Type locality: Syria (near “lac de Homs”).

Stratiomys hispanica ssp. planes James, 1941: 18. Type locality: Iran (Curum, 100 km from Bouchir).

Distribution:

PA: Algeria, Armenia, Austria, Bulgaria, Cyprus, Czech Republic, Egypt, France, Germany, Hungary, Iran, Israel, Italy, Kazakhstan, Morocco, Poland, Romania, Russia, Slovakia, Spain, Syria, Tunisia, Turkey, Turkmenistan, Ukraine, former Yugoslavia. [Sources: Woodley (2001) and Yimlahi et al. (2017)]

Local distribution and dates of collection: Unknown.

Notes:

This species was listed by Woodley (2001), Badrawy (2006) and Yimlahi et al. (2017) as recorded from Egypt, but no specimens have been collected or preserved in Egyptian museums to verify these records.

Stratiomys deserticolor Lindner, 1930

GBIF https://www.gbif.org/species/4293811

Nomenclature:

Stratiomyia segnis form deserticolor Lindner, 1930: 27. Type locality: Egypt (Siwa Oasis).

Materials Download as CSV

scientificName:
Stratiomys deserticolor
; taxonID:
https://www.gbif.org/species/4293811
; country:
Egypt
; locality:
Kharga Oasis
; decimalLatitude:
25.2500
; decimalLongitude:
30.5833
; samplingProtocol:
Not given
; eventDate:
08-12-1926
; sex:
1 male
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Stratiomys deserticolor
; taxonID:
https://www.gbif.org/species/4293811
; country:
Egypt
; locality:
Mariout
; decimalLatitude:
31.0172
; decimalLongitude:
29.76
; samplingProtocol:
Not given
; eventDate:
Jul 1934
; sex:
1 female
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC

Distribution:

PA: Egypt, Saudi Arabia. [Sources: original description (Lindner 1930), Abu-Zoherah et al. (1993) and Woodley (2001)]

Local distribution and dates of collection (Fig. 8): EGYPT: Coastal Strip: Mariout (July). Western Desert: Kharga Oasis, Siwa Oasis (February to August) [Sources: Original description (Lindner 1930) and museum material]. SAUDI ARABIA: localities and dates unknown. [Sources: Abu-Zoherah et al. (1993)]

Figure 8.

Distribution map of Stratiomys deserticolor Lindner and Stratiomys longicornis (Scopoli).

Stratiomys longicornis (Scopoli, 1763)

GBIF https://www.gbif.org/species/1577660

Nomenclature:

Hirtea longicornis Scopoli, 1763: 367. Type locality: “Carniola” [= Slovenija?].

Musca tenebricus Harris, 1778: 45. Type locality: England.

Stratiomys strigata Fabricius, 1781: 417. Type locality: Italy.

Stratiomys tomentosa Schrank, 1803: 94. Type locality: Germany (Ingolstadt).

Stratiomys villosa Meigen, 1804: 124. Type locality: Europe.

Stratiomys nubeculosa Meigen, 1804: 125. Type locality: Europe.

Stratiomys thoracica Fabricius, 1805: 79. Type locality: France.

Stratiomys hirtuosa Meigen, 1830: 347. Type locality: Europe.

Stratiomys anubis Wiedemann, 1830: 60. Type locality: Egypt.

Stratiomyia flavifrons Macquart, 1838: 179. Type locality: “Mesopotamie” [= Iraq].

Stratiomys strigata var. pallida Loew, 1840: 25. Type locality: Poland (Poznañ).

Stratiomys strigata var. pallida Loew, 1840: 557. Type locality: Poland (Poznañ). Preoccupied, primary homonym of Stratiomys strigata var. pallida Loew, 1840.

Stratiomys lambessiana Bigot, 1879: 212. Type locality: Algeria (Lambessa).

Stratiomys flavolimbata Costa, 1893: 21. Type locality: Tunisia.

Stratiomyia pleskei Wagner, 1903: 108. Type locality: Uzbekistan (Fergana).

Stratiomyia segnis Becker, 1906: 8. Type locality: Tunisia (vicinity of Tunis).

Hirtea efflatouni Lindner, 1925: 148. Type locality: Egypt (Giza).

Stratiomyia (Hirtea) surcoufi Séguy, 1930: 63. Type locality: Algeria (Touggourt).

Hirtea surcoufi Séguy, 1932: 125. Type locality: Algeria (Touggourt). Preoccupied, secondary homonym of Stratiomyia surcoufi Séguy, 1930.

Stratiomyia longicornis ssp. palaestinensis Lindner, 1937: 64. Type locality: Israel (Upper-Galilee, Kfar-Giladi).

Stratiomyia (Hirtea) longicornis ssp. flavoscutellata Lindner, 1940: 24. Type locality: China (Shanxi Province: “Ta-tong-fou”). Preoccupied, primary homonym of Stratiomyia flavoscutellata Wulp, 1885.

Materials Download as CSV

scientificName:
Stratiomys longicornis
; taxonID:
https://www.gbif.org/species/1577660
; country:
Egypt
; locality:
Siala
; decimalLatitude:
31.20849
; decimalLongitude:
29.8805
; samplingProtocol:
Not given
; eventDate:
Nov 1913
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
ESEC
scientificName:
Stratiomys longicornis
; taxonID:
https://www.gbif.org/species/1577660
; country:
Egypt
; locality:
Fayed
; decimalLatitude:
30.32382
; decimalLongitude:
32.3008
; samplingProtocol:
Not given
; eventDate:
Oct 1924
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
ESEC
scientificName:
Stratiomys longicornis
; taxonID:
https://www.gbif.org/species/1577660
; country:
Egypt
; locality:
Marg
; decimalLatitude:
31.0667
; decimalLongitude:
30.2167
; samplingProtocol:
Not given
; eventDate:
03-24-1918
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
ESEC
scientificName:
Stratiomys longicornis
; taxonID:
https://www.gbif.org/species/1577660
; country:
Egypt
; locality:
Kosseima
; decimalLatitude:
30.90307
; decimalLongitude:
34.3836
; samplingProtocol:
Not given
; eventDate:
Aug 1924
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
ESEC
scientificName:
Stratiomys longicornis
; taxonID:
https://www.gbif.org/species/1577660
; country:
Egypt
; locality:
Moweileh
; decimalLatitude:
30.3924
; decimalLongitude:
34.1412
; samplingProtocol:
Not given
; eventDate:
08-21-1924
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
ESEC
scientificName:
Stratiomys longicornis
; taxonID:
https://www.gbif.org/species/1577660
; country:
Egypt
; locality:
Helwan
; decimalLatitude:
29.8500
; decimalLongitude:
31.3333
; samplingProtocol:
Not given
; eventDate:
11-25-1913
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
ESEC
scientificName:
Stratiomys longicornis
; taxonID:
https://www.gbif.org/species/1577660
; country:
Egypt
; locality:
Benha
; decimalLatitude:
30.46572
; decimalLongitude:
31.18121
; samplingProtocol:
Not given
; eventDate:
05-09-1918
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
ESEC
scientificName:
Stratiomys longicornis
; taxonID:
https://www.gbif.org/species/1577660
; country:
Egypt
; locality:
Gezeira
; decimalLatitude:
30.04596
; decimalLongitude:
31.22435
; samplingProtocol:
Not given
; eventDate:
09-21-1917
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
ESEC
scientificName:
Stratiomys longicornis
; taxonID:
https://www.gbif.org/species/1577660
; country:
Egypt
; locality:
Kharga Oasis
; decimalLatitude:
25.25
; decimalLongitude:
30.5833
; samplingProtocol:
Not given
; eventDate:
01-03-1914
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
ESEC

Distribution:

PA: Afghanistan, Albania, Algeria, Armenia, Austria, Azerbaijan, Belgium, Bulgaria, China, Cyprus, Czech Republic, Denmark, Egypt, England, France, Germany, Greece, Hungary, Iran, Israel, Italy including Sardinia, Korea, Lithuania, Malta, Mongolia, Morocco, Netherlands, Poland, Portugal, Romania, Russia, Scotland, Slovakia, Slovenia, Spain, Sweden, Switzerland, Tunisia, Turkey, former Yugoslavia. [Sources: Woodley (2001), Mason et al. (2009) and Yimlahi et al. (2017)]

Local distribution and dates of collection (Fig. 8): EGYPT: Coastal Strip: El-Siala, Cleopatra, Dekhela, Nuzha (April to November). Eastern Desert: Fayed (October). Fayoum: Fayoum City, Nazla, Sanhur (April, May and September). Lower Nile Valley and Delta: Abu-Rawash, Benha, Faraskour, Gezeira, Giza, Helwan, Kerdassa, Marg, Sandoub, Shubra (March to November). Sinai: El-Arish, Kosseimah, Moweileh (April and August). Western Desert: Kharga Oasis, Siwa Oasis (January and May). [Sources: Lindner (1925) and museum material]

Stratiomys singularior (Harris, 1776)

GBIF https://www.gbif.org/species/4293822

Nomenclature:

Musca singularius Harris, 1778: 45. Type locality: England.

Stratiomys furcata Fabricius, 1794: 264. Type locality: Germany.

Stratiomys panthaleon Fallén, 1817: 7. Type locality: Sweden.

Stratiomys riparia Meigen, 1822: 138. Type locality: Europe.

Stratiomys paludosa Siebke, 1863: 149. Type locality: Norway (Dovre Mountains: Jerkin). Preoccupied, primary homonym of Stratiomys paludosa Schummel in Gravenhorst, 1837.

Materials Download as CSV

scientificName:
Stratiomys singularior
; taxonID:
https://www.gbif.org/species/4293822
; country:
Egypt
; locality:
Fayoum
; decimalLatitude:
29.32061
; decimalLongitude:
30.818
; samplingProtocol:
Not given
; eventDate:
03-01-1947
; sex:
1 male
; lifeStage:
Adult
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Stratiomys singularior
; taxonID:
https://www.gbif.org/species/4293822
; country:
Egypt
; locality:
Girza
; decimalLatitude:
29.49968
; decimalLongitude:
31.0738
; samplingProtocol:
Not given
; eventDate:
04-09-1950
; sex:
2 males
; lifeStage:
Adult
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Stratiomys singularior
; taxonID:
https://www.gbif.org/species/4293822
; country:
Egypt
; locality:
Abu-Zaabal
; decimalLatitude:
30.24098
; decimalLongitude:
31.35211
; samplingProtocol:
Not given
; eventDate:
03-19-1950
; sex:
1 male
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC

Distribution:

PA: Armenia, Austria, Belgium, Bulgaria, China, Czech Republic, Denmark, Egypt, England, Estonia, Finland, France, Germany, Hungary, Iran, Ireland, Italy, Kazakhstan, Lithuania, Mongolia, Netherlands, Norway, Poland, Romania, Russia, Slovakia, Spain, Sweden, Switzerland, Ukraine, former Yugoslavia. [Sources: Woodley (2001) and Badrawy (2006)]

Local distribution and dates of collection (Fig. 9): EGYPT: Fayoum: Fayoum City, Girza (March and April). Lower Nile Valley & Delta: Abu-Zaabal (March). [Source: Badrawy (2006) and museum material]

Figure 9.

Distribution map of Stratiomys singularior (Harris) and Nemotelus anchora Loew.

Subfamily Nemotelinae

Genus Nemotelus Geoffroy, 1762

GBIF https://www.gbif.org/species/1501964

Nomenclature:

Nemotelus Geoffroy, 1762: 450, 542. Type species: Musca pantherina Linnaeus, by designation of I.C.Z.N. (1957).

Subgenus Nemotelus Geoffroy, 1762

Nemotelus (Nemotelus) anchora Loew, 1846

GBIF https://www.gbif.org/species/1578822

Nomenclature:

Nemotelus anchora Loew, 1846: 429. Type locality: Italy (Sicily: Siracusa).

Nemotelus siculus Jaennicke, 1866: 224. Type locality: Italy (Sicily).

Nemotelus persicus Pleske in Lindner, 1937: 137. Type locality: Iran (Irak-Adzhemi: Buyun Village).

Material Download as CSV

scientificName:
Nemotelus anchora
; taxonID:
https://www.gbif.org/species/1578822
; country:
Egypt
; locality:
Fayoum
; decimalLatitude:
29.32061
; decimalLongitude:
30.818
; samplingProtocol:
Not given
; eventDate:
03-01-1947
; sex:
32 males, 5 females
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC

Distribution:

PA: Algeria, Egypt, Iran, Israel, Italy (including Sicily and Sardinia), Malta, Russia, Tunisia. [Sources: Woodley (2001), Mason et al. (2009) and Mohammad et al. (2009)]

Local distribution and dates of collection (Fig. 9): EGYPT: Fayoum: ? (March). [Source: Mohammad et al. (2009)]

Nemotelus (Nemotelus) candidus Becker, 1906

GBIF https://www.gbif.org/species/1578675

Nomenclature:

Nemotelus candidus Becker, 1906: 4. Type locality: Algeria (Biskra: Hammam-Salahin).

Material Download as CSV

scientificName:
Nemotelus candidus
; taxonID:
https://www.gbif.org/species/1578675
; country:
Egypt
; locality:
Wadi El-Natroun
; decimalLatitude:
30.3814
; decimalLongitude:
30.3441
; samplingProtocol:
Not given
; eventDate:
08-06-1929
; sex:
4 males, 1 female
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC

Distribution:

PA: Algeria, Egypt. [Sources: Woodley (2001) and Mohammad et al. (2009)]

Local distribution and dates of collection (Fig. 10): EGYPT: Coastal Strip: Dekhela (May and September). Western Desert: Wadi El-Natroun (August). [Source: Mohammad et al. (2009)]

Figure 10.

Distribution map of Nemotelus candidus Becker and Nemotelus dentatus Becker.

Nemotelus (Nemotelus) dentatus Becker, 1902

GBIF https://www.gbif.org/species/1578829

Nomenclature:

Nemotelus dentatus Becker, 1902: 7. Type localities: Egypt (Birket Qaroun, Damietta and Alexandria).

Materials Download as CSV

scientificName:
Nemotelus dentatus
; taxonID:
https://www.gbif.org/species/1578829
; country:
Egypt
; locality:
Maadi
; decimalLatitude:
29.95772
; decimalLongitude:
31.2505
; samplingProtocol:
Not given
; eventDate:
06-05-1916
; sex:
1 female
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
ESEC
scientificName:
Nemotelus dentatus
; taxonID:
https://www.gbif.org/species/1578829
; country:
Egypt
; locality:
Helwan
; decimalLatitude:
29.8500
; decimalLongitude:
31.3333
; samplingProtocol:
Not given
; eventDate:
02-11-1924
; sex:
1 female
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
ESEC
scientificName:
Nemotelus dentatus
; taxonID:
https://www.gbif.org/species/1578829
; country:
Egypt
; locality:
Helwan
; decimalLatitude:
29.8500
; decimalLongitude:
31.3333
; samplingProtocol:
Not given
; eventDate:
02-12-1924
; sex:
1 male
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
ESEC
scientificName:
Nemotelus dentatus
; taxonID:
https://www.gbif.org/species/1578829
; country:
Egypt
; locality:
Fayoum
; decimalLatitude:
29.32061
; decimalLongitude:
30.818
; samplingProtocol:
Not given
; eventDate:
03-01-1947
; sex:
2 males, 2 females
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Nemotelus dentatus
; taxonID:
https://www.gbif.org/species/1578829
; country:
Egypt
; locality:
Marg
; decimalLatitude:
31.0667
; decimalLongitude:
30.2167
; samplingProtocol:
Not given
; eventDate:
04-01-1923
; sex:
1 female
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Nemotelus dentatus
; taxonID:
https://www.gbif.org/species/1578829
; country:
Egypt
; locality:
Helwan
; decimalLatitude:
29.8500
; decimalLongitude:
31.3333
; samplingProtocol:
Not given
; eventDate:
04-02-1939
; sex:
1 female
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Nemotelus dentatus
; taxonID:
https://www.gbif.org/species/1578829
; country:
Egypt
; locality:
Kharga Oasis
; decimalLatitude:
25.2500
; decimalLongitude:
30.5833
; samplingProtocol:
Not given
; eventDate:
03-13-1932
; sex:
2 males
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Nemotelus dentatus
; taxonID:
https://www.gbif.org/species/1578829
; country:
Egypt
; locality:
Mirsa Matrouh
; decimalLatitude:
29.5696
; decimalLongitude:
26.4194
; samplingProtocol:
Not given
; eventDate:
05-08-1935
; sex:
2 males
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Nemotelus dentatus
; taxonID:
https://www.gbif.org/species/1578829
; country:
Egypt
; locality:
Ismaila
; decimalLatitude:
30.32382
; decimalLongitude:
32.3008
; samplingProtocol:
Not given
; eventDate:
04-07-1924
; sex:
2 males, 2 females
; lifeStage:
Adult
; recordedBy:
Tewfik
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Nemotelus dentatus
; taxonID:
https://www.gbif.org/species/1578829
; country:
Egypt
; locality:
Ismaila
; decimalLatitude:
30.32382
; decimalLongitude:
32.3008
; samplingProtocol:
Not given
; eventDate:
10-14-1926
; sex:
2 females
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Nemotelus dentatus
; taxonID:
https://www.gbif.org/species/1578829
; country:
Egypt
; locality:
Ismaila
; decimalLatitude:
30.32382
; decimalLongitude:
32.3008
; samplingProtocol:
Not given
; eventDate:
10-14-1926
; sex:
2 males
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Nemotelus dentatus
; taxonID:
https://www.gbif.org/species/1578829
; country:
Egypt
; locality:
Dekhela
; decimalLatitude:
31.12098
; decimalLongitude:
29.8157
; samplingProtocol:
Not given
; eventDate:
May-April 1923
; sex:
1 male
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Nemotelus dentatus
; taxonID:
https://www.gbif.org/species/1578829
; country:
Egypt
; locality:
Abu-Zaabal
; decimalLatitude:
30.24098
; decimalLongitude:
31.3521
; samplingProtocol:
Not given
; eventDate:
03-19-1950
; sex:
1 female
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC

Distribution:

PA: Egypt. [Source: Woodley (2001)]

Local distribution and dates of collection (Fig. 10): EGYPT: Coastal Strip: Alexandria, Dekhela, Mariout, Mersa Matrouh (March to May). Eastern Desert: Ismailia, El-Kantara (April, May & October). Fayoum: Birket Qaroun, El-Athar [(April & May). Lower Nile Valley and Delta: Abu-Zaabal, Damietta, El-Alag to Marg, El-Gabal El-Asfar, Ezbet El-Nakhl, Helwan, Maadi (February to April & November). Western Desert: Kharga Oasis, Wadi El-Natroun (March). [Sources: original description Becker (1902), Lindner (1925), Mohammad et al. (2009) and museum material]

Nemotelus (Nemotelus) marinus Becker, 1902

GBIF https://www.gbif.org/species/1578792

Nomenclature:

Nemotelus marinus Becker, 1902: 9. Type locality: Egypt (Suez, at the seashore).

Materials Download as CSV

scientificName:
Nemotelus marinus
; taxonID:
https://www.gbif.org/species/1578792
; country:
Egypt
; locality:
Fayed
; decimalLatitude:
30.32382
; decimalLongitude:
32.3008
; samplingProtocol:
Not given
; eventDate:
09-24-1925
; sex:
1 male
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Nemotelus marinus
; taxonID:
https://www.gbif.org/species/1578792
; country:
Egypt
; locality:
Wadi Hoff
; decimalLatitude:
29.8821
; decimalLongitude:
31.311
; samplingProtocol:
Not given
; eventDate:
08-09-1927
; sex:
1 female
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Nemotelus marinus
; taxonID:
https://www.gbif.org/species/1578792
; country:
Egypt
; locality:
Wadi El-Natroun
; decimalLatitude:
30.3814
; decimalLongitude:
30.3441
; samplingProtocol:
Not given
; eventDate:
08-06-1929
; sex:
1 female
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC

Distribution:

PA: Egypt. [Source: original description Becker (1902) and Woodley (2001)]

Local distribution and dates of collection (Fig. 11): EGYPT: Eastern Desert: El-Ferdan, Fayed, Ismailia, Suez, Wadi Hoff (April to October). Western Desert: Wadi El Natroun (August). [Sources: original description (Becker 1902), Mohammad et al. (2009) and museum material]

Figure 11.

Distribution map of Nemotelus marinus Becker and Nemotelus matrouhensis Mohammad et al.

Nemotelus (Nemotelus) matrouhensis Mohammad et al., 2009

Nomenclature:

Nemotelus matrouhensis Mohammad, Fadl, Gadalla & Badrawy, 2009: 103. Type locality: Egypt (Mersa Matrouh).

Material Download as CSV

Lectotype:

scientificName:
Nemotelus matrouhensis
; country:
Egypt
; locality:
Mersa Matrouh
; decimalLatitude:
29.5696
; decimalLongitude:
26.4194
; samplingProtocol:
Not given
; eventDate:
July & Aug 1931
; sex:
male
; lifeStage:
Adult
; recordedBy:
H.C.E & M.T.
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC

Distribution:

PA: Egypt. [Source: Woodley (2001)]

Local distribution and dates of collection (Fig. 11): EGYPT: Coastal Strip: Mersa Matrouh (July and August). [Source: original description (Mohammad et al. (2009)]

Notes:

Mohammad et al. (2009) published this species and inaccurately cited 13 males and 8 females as "holotype". They did not explicitly select a single specimen as holotype, so all cited specimens are considered as syntypes and a lectotype should be designated. As the illustration of the male genitalia was made from an intact specimen and did not show all the important structures, we are providing an illustration of dissected genitalia (Fig. 12) and a habitus photograph of the specimen (Fig. 13). When one of us (MH) requested a specimen, we were told that this would be a paratype, but because the specimen had no identification or type label at all, a red label was generated and attached to this specimen. This label reads: Paratype ♂/ Nemotelus matrouhensis/ det.: Haitham Badrawy 2009. However, now it is clear that none of the specimens had identification labels and also that this was not a paratype, but a syntype. The male genitalia resemble N. crenatus Egger, 1859 and N. obscuripes Loew, 1871. Lectotype designation: a male specimen in EFC is herein designated as lectotype and is labelled: Mirsa Matruh, July & Aug 1931 (leg. H.C.E. & M.T.), with a red lectotype label. Paralectotypes: 12 males and 8 females, same data.

Figure 12.

Nemotelus matrouhensis Mohammad et al.: A. Male terminalia, dorsal view; B. Phallic complex, dorsal view; C. same, lateral view.

Figure 13.

Nemotelus matrouhensis Mohammad et al.: A. Male habitus, dorsal view; B. Head, frontal view; C. Head and thorax, frontolateral view; D. Male habitus, lateral view.

Nemotelus (Nemotelus) niloticus Olivier, 1811

GBIF https://www.gbif.org/species/1578794

Nomenclature:

Nemotelus niloticus Olivier, 1811: 183. Type locality: Egypt.

Nemotelus fasciatus Olivier, 1811: 183. Type locality: Egypt (“bords du Nil & des canaux qui en dérivent”). Preoccupied by Nemotelus fasciatus Geoffroy in Fourcroy, 1785.

Nemotelus albifacies Becker, 1902: 9. Type locality: Egypt (Alexandria).

Nemotelus oasis Becker, 1906: 6. Type locality: Algeria (Biskra).

Nemotelus theodori Lindner, 1974: 95. Type locality: Israel (Arava Valley: Hazeva).

Nemotelus duofasciatus Woodley 2001. Replacement name for N. fasciatus Olivier, 1811.

Materials Download as CSV

scientificName:
Nemotelus niloticus
; taxonID:
https://www.gbif.org/species/1578794
; country:
Egypt
; locality:
Fayoum City
; decimalLatitude:
29.32061
; decimalLongitude:
30.818
; samplingProtocol:
Not given
; eventDate:
05-12-1918
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
ESEC
scientificName:
Nemotelus niloticus
; taxonID:
https://www.gbif.org/species/1578794
; country:
Egypt
; locality:
Beheira
; decimalLatitude:
30.62189
; decimalLongitude:
30.48755
; samplingProtocol:
Not given
; eventDate:
Apr 1924
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
ESEC
scientificName:
Nemotelus niloticus
; taxonID:
https://www.gbif.org/species/1578794
; country:
Egypt
; locality:
Damietta
; decimalLatitude:
31.34595
; decimalLongitude:
31.6317
; samplingProtocol:
Not given
; eventDate:
05-11-1918
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
ESEC
scientificName:
Nemotelus niloticus
; taxonID:
https://www.gbif.org/species/1578794
; country:
Egypt
; locality:
Quisna
; decimalLatitude:
30.53514
; decimalLongitude:
31.1117
; samplingProtocol:
Not given
; eventDate:
Mar 1924
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
ESEC
scientificName:
Nemotelus niloticus
; taxonID:
https://www.gbif.org/species/1578794
; country:
Egypt
; locality:
Mariout
; decimalLatitude:
31.0172
; decimalLongitude:
29.76
; samplingProtocol:
Not given
; eventDate:
07-09-1927
; sex:
1 female
; lifeStage:
Adult
; recordedBy:
Tewfik
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Nemotelus niloticus
; taxonID:
https://www.gbif.org/species/1578794
; country:
Egypt
; locality:
Mariout (Mallaha)
; decimalLatitude:
31.0172
; decimalLongitude:
29.76
; samplingProtocol:
Not given
; eventDate:
06-16-1929
; sex:
4 males, 11 females
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Nemotelus niloticus
; taxonID:
https://www.gbif.org/species/1578794
; country:
Egypt
; locality:
Fayoum City
; decimalLatitude:
29.32061
; decimalLongitude:
30.818
; samplingProtocol:
Not given
; eventDate:
04-23-1943
; sex:
2 males
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Nemotelus niloticus
; taxonID:
https://www.gbif.org/species/1578794
; country:
Egypt
; locality:
Fayoum City
; decimalLatitude:
29.32061
; decimalLongitude:
30.818
; samplingProtocol:
Not given
; eventDate:
04-20-1945
; sex:
2 females
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Nemotelus niloticus
; taxonID:
https://www.gbif.org/species/1578794
; country:
Egypt
; locality:
Giza-Fayoum Road
; decimalLatitude:
29.5564
; decimalLongitude:
30.8869
; samplingProtocol:
Not given
; eventDate:
04-14-1947
; sex:
1 female
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Nemotelus niloticus
; taxonID:
https://www.gbif.org/species/1578794
; country:
Egypt
; locality:
Helwan
; decimalLatitude:
29.8500
; decimalLongitude:
31.3333
; samplingProtocol:
Not given
; eventDate:
04-08-1934
; sex:
1 male
; lifeStage:
Adult
; recordedBy:
Farag
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Nemotelus niloticus
; taxonID:
https://www.gbif.org/species/1578794
; country:
Egypt
; locality:
Helwan
; decimalLatitude:
29.8500
; decimalLongitude:
31.3333
; samplingProtocol:
Not given
; eventDate:
04-14-1934
; sex:
1 female
; lifeStage:
Adult
; recordedBy:
Farag
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Nemotelus niloticus
; taxonID:
https://www.gbif.org/species/1578794
; country:
Egypt
; locality:
Helwan
; decimalLatitude:
29.8500
; decimalLongitude:
31.3333
; samplingProtocol:
Not given
; eventDate:
04-24-1934
; sex:
1 female
; lifeStage:
Adult
; recordedBy:
Farag
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Nemotelus niloticus
; taxonID:
https://www.gbif.org/species/1578794
; country:
Egypt
; locality:
WadiHoff
; decimalLatitude:
29.8821
; decimalLongitude:
31.311
; samplingProtocol:
Not given
; eventDate:
04-04-1930
; sex:
1 female
; lifeStage:
Adult
; recordedBy:
Farag
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Nemotelus niloticus
; taxonID:
https://www.gbif.org/species/1578794
; country:
Egypt
; locality:
Ramleh
; decimalLatitude:
31.2279
; decimalLongitude:
29.976
; samplingProtocol:
Not given
; eventDate:
09-17-1921
; sex:
1 female
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Nemotelus niloticus
; taxonID:
https://www.gbif.org/species/1578794
; country:
Egypt
; locality:
Dekhela
; decimalLatitude:
31.12098
; decimalLongitude:
29.8156
; samplingProtocol:
Not given
; eventDate:
06-24-1926
; sex:
1 male
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Nemotelus niloticus
; taxonID:
https://www.gbif.org/species/1578794
; country:
Egypt
; locality:
Dekhela
; decimalLatitude:
31.12098
; decimalLongitude:
29.8156
; samplingProtocol:
Not given
; eventDate:
06-24-1926
; sex:
1 female
; lifeStage:
Adult
; recordedBy:
H.C.E & M.T.
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Nemotelus niloticus
; taxonID:
https://www.gbif.org/species/1578794
; country:
Egypt
; locality:
Dekhela
; decimalLatitude:
31.12098
; decimalLongitude:
29.8156
; samplingProtocol:
Not given
; eventDate:
06-18-1929
; sex:
1 male
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Nemotelus niloticus
; taxonID:
https://www.gbif.org/species/1578794
; country:
Egypt
; locality:
Ismaila
; decimalLatitude:
30.32382
; decimalLongitude:
32.3008
; samplingProtocol:
Not given
; eventDate:
10-14-1926
; sex:
1 male
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC

Distribution:

AF: United Arab Emirates. PA: Algeria, Egypt, Israel, Italy (Sardinia), Tunisia. [Sources: Woodley (2001), Mason et al. (2009) and Hauser (2008)]

Local distribution and dates of collection (Fig. 14): EGYPT: Coastal Strip: Alexandria, Dekhela, Mariout (Mallaha), Ramleh (May to September). Eastern Desert: Fayed, Ismailia, Wadi Hoff (April, May and October). Fayoum: Fayoum City, Giza-Fayoum Road (April and May). Lower Nile Valley and Delta: Beheira, Damietta, Helwan, Quisna, Sherbin (March to September). Sinai: Zaranik Protectorate (April). [Sources: original description of N. albifacies (Becker 1902, Mohammad et al. (2009) and museum material]

Figure 14.

Distribution map of Nemotelus niloticus Olivier and Nemotelus notatus Zetterstedt.

Nemotelus (Nemotelus) notatus Zetterstedt, 1842

GBIF https://www.gbif.org/species/1578742

Nomenclature:

Nemotelus notatus Zetterstedt, 1842: 148. Type locality: Denmark (Copenhagen).

Nemotelus brachystomus Loew, 1846: 443. Type locality: Croatia (as “Dalmatien”).

Nemotelus leucorhynchus A. Costa, 1884: 61. Type locality: Italy (Sardinia: Stagno di Caliari).

Nemotelus nigroaeneus Verhoeff, 1891: 3. Type locality: Germany (Insel Norderney).

Nemotelus punctiventris Becker, 1902: 8. Type locality: Egypt (Alexandria).

Nemotelus brachystomus form aegyptiacus Lindner, 1925: 146. Type locality: Egypt (Alexandria: Nuzha, Cleopatra, Moharrem Bey, and Aboukir).

Nemotelus nigroaeneus ab. portalis Szilády, 1932: 33. Type locality: Germany (Borkum).

Nemotelus balearicus Lindner, 1937: 121. Type locality: Spain (Balearic Islands).

Nemotelus zernyi Lindner, 1937: 147. Type locality: Spain (Andalucía: Algeciras).

Materials Download as CSV

scientificName:
Nemotelus notatus
; taxonID:
https://www.gbif.org/species/1578742
; country:
Egypt
; locality:
Tanta
; decimalLatitude:
30.75725
; decimalLongitude:
30.9898
; samplingProtocol:
Not given
; eventDate:
05-11-1918
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
ESEC
scientificName:
Nemotelus notatus
; taxonID:
https://www.gbif.org/species/1578742
; country:
Egypt
; locality:
Khusus
; decimalLatitude:
30.15957
; decimalLongitude:
31.3125
; samplingProtocol:
Not given
; eventDate:
Apr 1918
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
ESEC
scientificName:
Nemotelus notatus
; taxonID:
https://www.gbif.org/species/1578742
; country:
Egypt
; locality:
Alag
; decimalLatitude:
30.18009
; decimalLongitude:
31.3521
; samplingProtocol:
Not given
; eventDate:
Nov 1924
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
ESEC
scientificName:
Nemotelus notatus
; taxonID:
https://www.gbif.org/species/1578742
; country:
Egypt
; locality:
Marg
; decimalLatitude:
31.0667
; decimalLongitude:
30.2167
; samplingProtocol:
Not given
; eventDate:
Mar 1918
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
ESEC
scientificName:
Nemotelus notatus
; taxonID:
https://www.gbif.org/species/1578742
; country:
Egypt
; locality:
El-Mahareeq (Kharga)
; decimalLatitude:
25.618206
; decimalLongitude:
30.6468
; samplingProtocol:
Not given
; eventDate:
Feb 1914
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
ESEC
scientificName:
Nemotelus notatus
; taxonID:
https://www.gbif.org/species/1578742
; country:
Egypt
; locality:
Beni Sweif
; decimalLatitude:
29.07788
; decimalLongitude:
31.10713
; samplingProtocol:
Not given
; eventDate:
02-14-1914
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
ESEC
scientificName:
Nemotelus notatus
; taxonID:
https://www.gbif.org/species/1578742
; country:
Egypt
; locality:
Kharga Oasis
; decimalLatitude:
25.25
; decimalLongitude:
30.5833
; samplingProtocol:
Not given
; eventDate:
Feb 1914
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
ESEC
scientificName:
Nemotelus notatus
; taxonID:
https://www.gbif.org/species/1578742
; country:
Egypt
; locality:
Kharga Oasis
; decimalLatitude:
25.25
; decimalLongitude:
30.5833
; samplingProtocol:
Not given
; eventDate:
03-10-1924
; lifeStage:
Adult
; recordedBy:
Priesner
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
ESEC
scientificName:
Nemotelus notatus
; taxonID:
https://www.gbif.org/species/1578742
; country:
Egypt
; locality:
Cleopatra
; decimalLatitude:
31.22021
; decimalLongitude:
29.9348
; samplingProtocol:
Not given
; eventDate:
07-12-1921
; sex:
1 female
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Nemotelus notatus
; taxonID:
https://www.gbif.org/species/1578742
; country:
Egypt
; locality:
Dekhela
; decimalLatitude:
31.12098
; decimalLongitude:
29.8156
; samplingProtocol:
Not given
; eventDate:
06-24-1926
; sex:
1 male
; lifeStage:
Adult
; recordedBy:
H.C.E & M.T.
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Nemotelus notatus
; taxonID:
https://www.gbif.org/species/1578742
; country:
Egypt
; locality:
Dekhela
; decimalLatitude:
31.12098
; decimalLongitude:
29.8156
; samplingProtocol:
Not given
; eventDate:
05-24-1925
; sex:
3 males, 5 females
; lifeStage:
Adult
; recordedBy:
H.C.E & M.T.
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Nemotelus notatus
; taxonID:
https://www.gbif.org/species/1578742
; country:
Egypt
; locality:
Dekhela
; decimalLatitude:
31.12098
; decimalLongitude:
29.8156
; samplingProtocol:
Not given
; eventDate:
07-09-1927
; sex:
1 male, 1 female
; lifeStage:
Adult
; recordedBy:
Efflatoun
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Nemotelus notatus
; taxonID:
https://www.gbif.org/species/1578742
; country:
Egypt
; locality:
Fayed
; decimalLatitude:
30.32382
; decimalLongitude:
32.3008
; samplingProtocol:
Not given
; eventDate:
07-24-1924
; sex:
1 female
; lifeStage:
Adult
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Nemotelus notatus
; taxonID:
https://www.gbif.org/species/1578742
; country:
Egypt
; locality:
Helwan
; decimalLatitude:
29.8500
; decimalLongitude:
31.3333
; samplingProtocol:
Not given
; eventDate:
04-08-1934
; sex:
?
; lifeStage:
Adult
; recordedBy:
Farag
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC
scientificName:
Nemotelus notatus
; taxonID:
https://www.gbif.org/species/1578742
; country:
Egypt
; locality:
Wadi Hoff
; decimalLatitude:
29.8821
; decimalLongitude:
31.311
; samplingProtocol:
Not given
; eventDate:
06-09-1927
; sex:
?
; lifeStage:
Adult
; recordedBy:
Tewfik
; identifiedBy:
M. El-Hawagry & M. Hauser
; dateIdentified:
2020-2021
; institutionCode:
EFC

Distribution:

PA: Albania, Austria, Azerbaijan, Belgium, Bulgaria, Croatia, Cyprus, Denmark, Egypt, England, Finland, France, Germany, Greece, Hungary, Iran, Ireland, Israel, Italy (including Sardinia), Netherlands, Norway, Poland, Romania, Russia, Spain, Sweden, Turkey, former Yugoslavia. [Sources: Woodley (2001), Üstüner and Hasbenli (2005), Mason et al. (2009) and Khaghaninia et al. (2015)]

Local distribution and dates of collection (Fig. 14): EGYPT: Coastal Strip: Abu-Kir, Cleopatra, Dekhela, Mariout, Moharram Bey, Nubar Bey, Nuzha, Ramleh (April to September). Eastern Desert: Ismailia, Wadi Hoff (May to July). Lower Nile Valley and Delta: Beni Sweif, Damietta, El-Alag, Helwan, El-Marg, Khusous, Tanta (February to May and November). Sinai: Zaranik Protectorate (April). Western Desert: Kharga Oasis (February and March). [Sources: original description of N. b. aegyptiacus (Lindner 1925), Mohammad et al. (2009) and museum material]

Analysis

Acknowledgements

The authors extend their appreciation to the Deanship of Scientific Research at King Saud University for funding this work through research group No [RG-1437-009].

We are grateful to Dr Neal Evenhuis, Bishop Museum, Honolulu, Hawaii, USA and Dr Mohammed Gamal El-Den Nasser, Entomology Department, Faculty of Science, Ain Shams University, Cairo, Egypt for providing critical assistance throughout this study. We thank Haitham Badrawy for the loan of a Nemotelus matrouhensis specimen. We are also grateful to the research team in KSMA for collecting, mounting and photographing specimens.

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Supplementary material

Endnotes