Biodiversity Data Journal : Taxonomic Paper
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Taxonomic Paper
Checklist of British and Irish Hymenoptera - Braconidae
expand article infoGavin R. Broad, Mark R. Shaw§, H. Charles J. Godfray|
‡ The Natural History Museum, London, United Kingdom
§ National Museums of Scotland, Edinburgh, United Kingdom
| University of Oxford, Oxford, United Kingdom
Open Access

Abstract

Background

The checklist of British and Irish Braconidae is revised, based in large part on the collections of the National Museums of Scotland, Edinburgh, and the Natural History Museum, London. Distribution records are provided at the country level together with extensive synonymy and bibliography.

New information

Of the 1,338 species regarded as valid, presumed native and certainly identified, 83 are here recorded for the first time from the British Isles. One new synonym is established (Dyscritus suffolciensis Morley, 1933 = Syntretus splendidus (Marshall, 1887) syn. nov.)

Keywords

Britain, Ireland, fauna, Ichneumonoidea

Introduction

The Braconidae is one of two families of the superfamily Ichneumonoidea, along with the Ichneumonidae. Given the size of each family in Britain (over 1,300 braconids and 2,500 ichneumonids) we are publishing the two checklists separately. This is one part of a series of papers revising the British and Irish list of Hymenoptera, that started with Broad and Livermore (2014a), Broad and Livermore (2014b) and Liston et al. (2014). For the background and rationale behind these British Hymenoptera checklists see Broad (2014). The bases for the taxonomy are Fauna Europaea and Taxapad (Yu et al. 2012) (braconid data for both compiled by Kees van Achterberg) but we have not relied on these sources for data on species occurrence in Britain. Rather, we have relied mostly on the primary literature and collections, especially NMS and BMNH. Indeed, large parts of the checklist rely on original work, identifying specimens in these collections.

Much of the synonymy adopted here is equivalent to the German list (Belokobylskij et al. 2003) but where there are taxonomic differences of opinion, usually van Achterberg’s interpretation has been followed, largely for consistency with Fauna Europaea, but also because of van Achterberg’s experience with many groups of braconids. The biggest departure from these sources is in the generic classification of Microgastrinae, for which we largely follow Mason (1981).

We reference all additions to and deletions from the British list since Huddleston (1978) and record country-level distribution within the British Isles, but regarding Ireland as one. We made much use of O'Connor et al. (1999) for Irish records. Because the braconid literature is extensive and scattered we also provide many taxonomic references. The braconid section of the 1978 British checklist (Huddleston 1978) repeats many of Shenefelt’s (e.g. Shenefelt 1973, Shenefelt 1974) mistakes and these are rectified here.

Figs 1, 2, 3, 4 illustrate a tiny part of the morphological and biological diversity of Braconidae. Shaw and Huddleston (1991) provide an introduction to the varied biology of this family of parasitoid wasps.

Figure 1.

Representative British Braconidae: Alysiinae and Aphidiinae

aFace of Alsysia manducator (Panzer) (Alysiinae), with exodont mandibles for emerging from Diptera puparia. BMNH specimen.
bPauesia sp. (Aphidiinae) (courtesy of P. Adams)
cAphidius ervi (Haliday) (Aphidiinae): mummified aphid host that the braconid pupates within. BMNH specimen.
dPraon volucre (Haliday) (Aphidiinae): mummified aphid host with the braconid pupa beneath. BMNH specimen.
Figure 2.

Representative British Braconidae: Doryctinae, Meteorinae and Microgastrinae

aDendrosoter protuberans (Nees) (Doryctinae) male (courtesy of P. Adams)
bZele albiditarsus Curtis (Meteorinae) female (courtesy of E. Brosens)
cCotesia affinis (Nees) (Microgastrinae) larvae emerging from their Laothoe populi (L.) (Lepidoptera: Sphingidae) host and then spinning cocoons (courtesy of M. Boddington)
dCotesia gonopterygis (Marshall) (Microgastrinae) overwintering cocoon mass ex Gonepteryx rhamni (L.) (Lepidoptera: Pieridae) (M.R. Shaw)
Figure 3.

Representative British Braconidae

aCotesia spuria (Wesmael) (Microgastrinae) cocoons ex larval host Apocheima hispidaria (D. & S.) (Lepidoptera: Geometridae) (courtesy of J. Voogd)
bCotesia vanessae (Reinhard) (Microgastrinae) emerging from larval host Aglais urticae (Linnaeus) (Lepidoptera: Nymphalidae) (courtesy of N. Spring)
cMicroplitis ocellatae (Bouché) (Microgastrinae) cocoons ex larval host Smerinthus ocellatus (Linnaeus) (Lepidoptera: Sphingidae) (courtesy of M. Boddington)
dHisteromerus mystacinus Wesmael (Rhyssalinae) eggs on pupal host Stictoleptura scutellata (L.) (Coleoptera: Cerambycidae) (M.R. Shaw)
Figure 4.

Representative British Braconidae: Sigalphinae

aAcampsis alternipes (Nees) ovipositing in larval host, Alsophila aescularia (D. & S.) (Lepidoptera: Geometridae) (M.R. Shaw)
bAcampsis alternipes (Nees) larva emerging from its larval host Alsophila aescularia (D. & S.) (Lepidoptera: Geometridae) to complete its feeding externally (M.R. Shaw)

Materials and methods

For a more detailed description of the background and rationale to the Hymenoptera checklist, see Broad (2014). We have tried to account for every name on the 1978 (Huddleston 1978) checklist and have referenced all additions to and deletions from that list. We provide rather extensive synonymy and citations because if you do not have access to the Taxapad database (Yu et al. 2012) it can be very difficult to trace the fate of names in the voluminous and scattered literature. Conventions and abbreviations are listed below.

[species] taxon deleted from the British and Irish list

BMNH Natural History Museum, London

NMS National Museums of Scotland, Edinburgh

# known introductions occurring only under artificial conditions

? status (including uncertain synonymy) or identification in the British Isles uncertain

misident. has been misidentified as this name

nom. dub. nomen dubium, a name of doubtful status

nom. ob. nomen oblitum, ‘forgotten name’, does not have priority over a younger name

nom. nov. nomen novum, a replacement name

nom. nud. nomen nudum, an unavailable name, with no type specimen

preocc. name preoccupied (junior homonym)

stat. rev. status revocatus, revived status (e.g., raised from synonymy)

unavailable name unavailable under provisions of the ICZN code

var. variety, only available as a valid name under certain provisions of the ICZN code

Word document and spreadsheet versions of the checklist are available in the supplementary materials. Future updates to the British and Irish list will be incorporated in an online version of the checklist at Hymenoptera of the British Isles.

Agathidinae

Subfamily Agathidinae Haliday, 1833

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Notes: 

Except for Agathis and ‘Bassus’ species (i.e. including Lytopylus and Therophilus), distribution and synonymic data from Nixon (1986).

Tribe Agathidini Nees, 1814

Nomenclature: 

BASSINI Nees, 1812 invalid

EUMICRODINI Förster, 1863

BASSINI Förster, 1869 preocc.

MICRODINI Ashmead, 1900

MESOCOELINI Viereck, 1918

ANEUROBRACONINI Fahringer, 1936

EARININI Sharkey, 1992

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Genus Agathis Latreille, 1804

Nomenclature: 

AENIGMOSTOMUS Ashmead, 1900

METRIOSOMA Szépligeti, 1902

BAEOGNATHA Kokujev, 1903 synonymy by Achterberg (2011)

LISSAGATHIS Cameron, 1911

RHAMPHAGATHIS Tobias, 1962

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Notes: 

Distribution and synonymic data from Nixon (1986) and Simbolotti and Achterberg (1999).

Species of Agathis excluded from the British and Irish list:

[malvacearum Latreille, 1805; syn. panzeri (Jurine, 1807, Ichneumon); metzneriae Muesebeck, 1967] Listed by Huddleston (1978); probably refers to varipes.

Agathis anglica Marshall, 1885

Nomenclature: 

longicauda Kokujev, 1895 preocc.

?marshalli Fahringer, 1937

albanica Fischer, 1957

syriaca Fischer, 1957

caucasica Tobias, 1963

taiwanensis Chou & Sharkey, 1989

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Distribution: 
England, Wales

Agathis assimilis Kokujev, 1895

Nomenclature: 

propinqua Kokujev, 1895

jakowlewi Kokujev, 1895

sibirica Telenga, 1933

anchisiades Nixon, 1986

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Distribution: 
Scotland
Notes: 

added by Nixon (1986)

Agathis breviseta Nees, 1812

Nomenclature: 

achterbergi Nixon, 1986

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Distribution: 
England, Scotland, Ireland
Notes: 

Simbolotti and Achterberg (1999) synonymised Agathis achterbergi Nixon, 1986 under breviseta but it is listed as a valid species in Taxapad (Yu et al. 2012).

Agathis fuscipennis (Zetterstedt, 1838)

Nomenclature: 

Microgaster fuscipennis Zetterstedt, 1838

breviseta misident.

rostrata misident.

glabricula Thomson, 1895

schmiedeknechti Kokujev, 1895

annulata Fahringer, 1937

meridionellae Fischer, 1957

albicostellae Fischer, 1966

artemesiana Fischer, 1966

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Distribution: 
England, Ireland

Agathis griseifrons Thomson, 1895

Nomenclature: 

laticarpa Telenga, 1955

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Distribution: 
England, Ireland

Agathis lugubris (Förster, 1863)

Nomenclature: 

Cenostomus lugubris Förster, 1863

minuta Niezabitowski, 1910

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Distribution: 
England, Scotland, Wales, Ireland
Notes: 

added by Nixon (1986)

Agathis montana Shestakov, 1932

Nomenclature: 

zaykovi Nixon, 1986

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Distribution: 
England
Notes: 

added by Simbolotti and Achterberg (1999)

Agathis nigra Nees, 1812

Nomenclature: 

testaceipes Fischer, 1957

kasachstanica Tobias, 1963

nixoni Belokobylskij & Jervis, 1998

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Distribution: 
England

Agathis rufipalpis Nees, 1812

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Distribution: 
Ireland

Agathis tibialis Nees, 1814

Nomenclature: 

genualis Marshall, 1898

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Distribution: 
England
Notes: 

NMS, det. van Achterberg, added here

Agathis varipes Thomson, 1895

Nomenclature: 

simulatrix Kokujev, 1895

rufipes Ivanov, 1899

dissimilis Shestakov, 1928

rufilabialis Fahringer, 1937

glabricollis Telenga, 1957

serratulae Tobias, 1963

lederi Fischer, 1968

ariadne Nixon, 1986

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Distribution: 
England
Notes: 

added by Nixon (1986)

Genus Bassus Fabricius, 1804

Nomenclature: 

MICRODUS Nees, 1814

DIPLOZON Haliday, 1833

EURYZONA Haliday, 1838 nom. nud.

EUMICRODUS Förster, 1863

HEMIOGASTER Enderlein, 1920 synonymy by Achterberg and Long (2010)

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Notes: 

Following molecular phylogenetic analysis of Agathidinae (Sharkey et al. 2006), the genus Bassuss.l. was recognised as being a polyphyletic assemblage. Species are now being described in or reassigned to Lytopylus, Thermophilus and the non-British Camptothlipsis (Sharkey et al. 2009, Stevens et al. 2010, Stevens et al. 2011, Achterberg and Long 2010, Achterberg 2011) but very few European species have been formally transferred. The species on the British list have therefore been reassigned here on the bases of recent generic keys (e.g. Sharkey et al. 2009) in anticipation of future taxonomy. Whereas Bassuss. s. is now restricted to a small group of species (only one in Britain), Therophilus remains large and probably para- or polyphyletic. Distribution and synonymic data for Bassus, Lytopylus and Therophilus from Nixon (1986) and Simbolotti and Achterberg (1992).

Bassus calculator (Fabricius, 1798)

Nomenclature: 

Ichneumon calculator Fabricius, 1798

abscissus (Ratzeburg, 1844, Microdus)

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Distribution: 
England

Genus Earinus Wesmael, 1837

Nomenclature: 

Förster, 1863

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Earinus elator (Fabricius, 1804)

Nomenclature: 

Banchus elator Fabricius, 1804

nitidulus (Nees, 1814, Microdus)

thoracicus (Nees, 1834, Microdus)

major (Fonscolombe, 1846, Agathis)

pilosus Tobias, 1960

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Distribution: 
England, Scotland, Ireland

Earinus gloriatorius (Panzer, 1809)

Nomenclature: 

Bassus gloriatorius Panzer, 1809

gloriator (Nees, 1812, Microdus)

ochropes (Curtis, 1829, Microdus) nom. nud.

affinis (Wesmael, 1837, Microdus)

delusor (Wesmael, 1837, Microdus)

tuberculatus (Wesmael, 1837, Microdus)

varicoxis (Wesmael, 1837, Microdus)

niger (Zetterstedt, 1838, Microgaster)

bicingulatus (Thomson, 1895, Agathis)

ochropes Lyle, 1920

ruficoxis Fahringer, 1937

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Distribution: 
England, Scotland, Ireland, Isle of Man

Earinus transversus Lyle, 1920

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Distribution: 

England

Notes: 

distribution data from Shaw (2005)

Genus Lytopylus Förster, 1863

Nomenclature: 

AEROPHILUS Szépligeti, 1902 synonymy by Sharkey et al. (2006)

NEOMICRODUS Szépligeti, 1908

AEROPHILOPSIS Viereck, 1913

AEROPHILINA Enderlein, 1920

IOXIA Enderlein, 1920

HORMAGATHIS Brues, 1926

OBESOMICRODUS Papp, 1971

FACILAGATHIS van Achterberg & Chen, 2004

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Notes: 

See comments under Bassus; generic synonymy from Sharkey et al. (2009).

Lytopylus rufipes (Nees, 1814)

Nomenclature: 

Microdus rufipes Nees, 1814

germanicus (Enderlein, 1904, Braunsia)

diversus Muesebeck, 1933

amurensis (Shestakov, 1940, Microdus)

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Distribution: 
England

Genus Therophilus Wesmael, 1837

Nomenclature: 

ORGILONEURA Ashmead, 1900

AGATHIELLA Szépligeti, 1902

AEROPHILIODES Strand, 1911

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Notes: 

See comments under Bassus; generic synonymy from Sharkey et al. (2009).

Species excluded from the British and Irish list:

[brevicaudis (Reinhard, 1867, Microdus)] Listed as a ‘species inquirendae’ by Nixon (1986) but included as a German species by Simbolotti and Achterberg (1992). Transferred to Bassus by Simbolotti and Achterberg (1999) but not listed as a species of Therophilus by Sharkey and Stoelb (2012).

[nugax (Reinhard, 1867, Microdus); syn. rufiventris (Abdinbekova, 1975, Microdus)] Listed by Huddleston (1978) but no mention of British specimens by Nixon (1986) or Simbolotti and Achterberg (1992).

Therophilus arcuatus (Reinhard, 1867)

Nomenclature: 

Microdus arcuatus Reinhard, 1867

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Notes: 

Taken out of synonymy with conspicuus by Simbolotti and Achterberg (1992).

Therophilus cingulipes (Nees, 1812)

Nomenclature: 

Microdus cingulipes Nees, 1812

nantouensis Chou & Sharkey, 1989

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Distribution: 
England, Ireland

Therophilus clausthalianus (Ratzeburg, 1844)

Nomenclature: 

Ichneumon clausthalianus Ratzeburg, 1844

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Distribution: 
England, Ireland

Therophilus conspicuus (Wesmael, 1837)

Nomenclature: 

Microdus conspicuus Wesmael, 1837

zonatus (Marshall, 1885, Earinus)

carpocapsae Cushman, 1915

angustatus (Telenga, 1955, Microdus)

variabilis Chou & Sharkey, 1989

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Distribution: 
England, Ireland

Therophilus tegularis (Thomson, 1895)

Nomenclature: 

Agathis tegularis Thomson, 1895

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Distribution: 
England
Notes: 

added by Simbolotti and Achterberg (1992)

Therophilus tumidulus (Nees, 1812)

Nomenclature: 

Microdus tumidulus Nees, 1812

intermedius (Ivanov, 1899, Eumicrodus)

annae (Enderlein, 1908, Microdus)

aino (Watanabe, 1937, Microdus)

ruficoxis (Fahringer, 1937, Microdus)

rufus (Fahringer, 1938, Microdus)

bicolor (Shestakov, 1940, Microdus) preocc.

victoris (Telenga, 1955, Microdus)

shestakovi (Shenefelt, 1970, Agathis)

anuphrievi (Tobias, 1986, Microdus)

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Distribution: 
England, Scotland, Ireland

Genus Therophilus Wesmael, 1837

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Notes: 

Species incertae sedis within Agathidinae; Sharkey and Stoelb (2012) defined Therophilus as a monophyletic genus and listed the included species, but several species currently classified in Therophilus are of uncertain generic placement.

? Therophilus dimidiator (Nees, 1834)

Nomenclature: 

Microdus dimidiator Nees, 1834

cingulator (Ratzeburg, 1852, Microdus)

laticinctus (Cresson, 1873, Microdus)

ocellanae (Richardson, 1913, Microdus)

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Distribution: 

England

Notes: 

Listed by Huddleston (1978) but no mention of British specimens by Nixon (1986), who states, however, that the traditional interpretation of the species is probably correct. No British or Irish specimens were seen by Simbolotti and Achterberg (1992) but van Achterberg has identified an English specimen in BMNH as Therophilus dimidiator.

? Therophilus epinotiae (van Achterberg, 1992)

Nomenclature: 

Bassus epinotiae van Achterberg, 1992

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Distribution: 
England
Notes: 

added by Simbolotti and Achterberg (1992)

? Therophilus linguarius (Nees, 1812)

Nomenclature: 

Microdus linguarius Nees, 1812

minor (Enderlein, 1908, Microdus)

kaszabi (Papp, 1967, Vipio)

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Distribution: 
England

? Therophilus mediator (Nees, 1814)

Nomenclature: 

Microdus mediator Nees, 1814

lugubrator (Ratzeburg, 1852, Microdus)

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Distribution: 
England
Notes: 

Transferred from Agathis by Simbolotti and Achterberg (1999).

? Therophilus pumilus (Ratzeburg, 1844)

Nomenclature: 

Microdus pumilus Ratzeburg, 1844

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Distribution: 
England
Notes: 

NMS, det. Shaw, added here; listed by Huddleston (1978) but no mention of British specimens by Nixon (1986) or Simbolotti and Achterberg (1992). Transferred from Agathis by Simbolotti and Achterberg (1999) but listed as a species of Agathis in Taxapad (Yu et al. 2012).

? Therophilus rugulosus (Nees, 1834)

Nomenclature: 

Microdus rugulosus Nees, 1834

compeditus (Vollenhoven, 1878, Microdus)

punctatus (Abdinbekova, 1975, Microdus)

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Distribution: 

England, Ireland

Alysiinae

Subfamily Alysiinae Leach, 1815

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Tribe Alysiini Leach, 1815

Nomenclature: 

ALLOEINI Ashmead, 1900

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Genus Adelurola Strand, 1928

Nomenclature: 

ADELURA Förster, 1863 preocc.

NEOCARPA Fischer, 1966

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Adelurola florimela (Haliday, 1838)

Nomenclature: 

Alysia florimela Haliday, 1838

multiarticulata (Marshall, 1898, Phaenocarpa)

pentapleuroides (Fischer, 1971, Dapsilarthra) unavailable

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Distribution: 
England, Scotland, Ireland, Isle of Man

Genus Alloea Haliday, 1833

Nomenclature: 

DIASPASTA Förster, 1863

LAMADATHA Cameron, 1900

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Alloea contracta Haliday, 1833

Nomenclature: 

testaceipes (Cameron, 1900, Lamadatha)

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Distribution: 
England, Scotland, Wales, Ireland

Alloea lonchopterae Fischer, 1966

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Distribution: 

England, Isle of Man

Notes: 

Added by Godfray and Bland (2011) English record from specimen in BMNH.

Genus Alysia Latreille, 1804

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Subgenus Alysia Latreille, 1804

Nomenclature: 

CECHENUS Illiger, 1807

BASSUS Nees, 1812

GONIARCHA Förster, 1863

STROPHAEA Förster, 1863

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Notes: 

Much of the taxonomy and distribution from Wharton (1986).

Species of Alysia (Alysia) excluded from the British and Irish list:

[cingulata Nees, 1834 nom. dub.]

Alysia (Alysia) alticola (Ashmead, 1890)

Nomenclature: 

Pentapleura alticola Ashmead, 1890

soror Marshall, 1894

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Distribution: 
England, Scotland, Wales, Isle of Man

Alysia (Alysia) frigida Haliday, 1838

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Distribution: 
England
Notes: 

added by Godfray and Achterberg (2015)

Alysia (Alysia) incongrua Nees, 1834

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Distribution: 
England, Scotland, Ireland

Alysia (Alysia) lucia Haliday, 1838

Nomenclature: 

diversiceps Fischer, 1967

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Distribution: 
Scotland

Alysia (Alysia) lucicola Haliday, 1838

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Distribution: 
England, Scotland, Ireland

Alysia (Alysia) luciella Stelfox, 1941

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Distribution: 
Ireland

Alysia (Alysia) manducator (Panzer, 1799)

Nomenclature: 

Ichneumon manducator Panzer, 1799

haematopa (Gmelin, 1790, Ichneumon) preocc.

stercoraria Latreille, 1805

apicalis Curtis, 1826

similis Curtis, 1826

stercorator Lamarck, 1835

curtungula Thomson, 1895

bucephala Marshall, 1898

manducatrix Schulz, 1906

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Distribution: 
England, Scotland, Wales, Ireland, Isle of Man

Alysia (Alysia) truncator (Nees, 1812)

Nomenclature: 

Bassus truncator Nees, 1812

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Distribution: 
England, Scotland, Ireland

Subgenus Anarcha Förster, 1863

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Notes: 

Much of the taxonomy and distribution from Wharton (1988b).

Species of Alysia (Anarcha) excluded from the British and Irish list:

[similis (Nees, 1812, Bassus) nom. dub.]

Alysia (Anarcha) atra Haliday, 1838

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Distribution: 
Ireland

Alysia (Anarcha) fuscipennis Haliday, 1838

Nomenclature: 

obscuripes Thomson, 1895

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Distribution: 
England, Scotland, Ireland

Alysia (Anarcha) mandibulator (Nees, 1812)

Nomenclature: 

Bassus mandibulator Nees, 1812

loripes Haliday, 1838

mandibulatrix Schulz, 1906

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Distribution: 
England, Ireland

Alysia (Anarcha) rufidens Nees, 1834

Nomenclature: 

puncticollis Thomson, 1895

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Distribution: 
England, Ireland

Alysia (Anarcha) sophia Haliday, 1835

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Distribution: 
Ireland

Alysia (Anarcha) subaperta Thomson, 1895

Nomenclature: 

similis misident.

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Distribution: 
England
Notes: 

added by Wharton (1988b)

Alysia (Anarcha) thapsina Wharton, 1988

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Distribution: 
Ireland
Notes: 

added by Wharton (1988b)

Alysia (Anarcha) tipulae (Scopoli, 1763)

Nomenclature: 

Ichneumon tipulae Scopoli, 1763

abdominator (Nees, 1814, Bassus)

notabilis (Förster, 1863, Anarcha)

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Distribution: 
England, Scotland, Ireland

Alysia (Anarcha) umbrata Stelfox, 1941

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Distribution: 
Ireland

Genus Anisocyrta Förster, 1863

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Anisocyrta perdita (Haliday, 1838)

Nomenclature: 

Alysia perdita Haliday, 1838

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Distribution: 
Scotland

Genus Aphaereta Förster, 1863

Nomenclature: 

TRICHESIA Provancher, 1880

TRINARIA Provancher, 1886

APHAERETE Dalla Torre, 1898

ATOPANDRIUM Graham, 1952

TRISYNALDIS Fischer, 1958

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Notes: 

Atopandrium was synonymised under Aphaereta by Achterberg (1995) but treated as a valid genus again in Taxapad (Yu et al. 2012).

Aphaereta debilitata Morley, 1933

Nomenclature: 

loripenne (Graham, 1952, Atopandrium)

confluctum (Fischer, 1958, Trisynaldis)

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Distribution: 
England, Wales

Aphaereta falcigera Graham, 1960

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Distribution: 
England, Ireland

Aphaereta major (Thomson, 1895)

Nomenclature: 

Alysia major Thomson, 1895

major Marshall, 1898 preocc.

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Distribution: 
England, Scotland, Ireland

Aphaereta minuta (Nees, 1811)

Nomenclature: 

Stephanus minutus Nees, 1811

cephalotes (Haliday, 1833, Alysia)

fuscipes (Nees, 1834, Alysia)

confluens (Ratzeburg, 1844, Alysia)

stigmaticalis (Thomson, 1895, Alysia)

inepta Morley, 1933

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Distribution: 
England, Scotland, Ireland

Aphaereta pallipes (Say, 1829)

Nomenclature: 

Alysia pallipes Say, 1829

auripes (Provancher, 1881, Trichesia)

pilicornis (Provancher, 1886, Trinaria)

californica Ashmead, 1889

muscae Ashmead, 1889

oscinidis Ashmead, 1889

pallidipes (Dalla Torre, 1898, Aphaerete)

delosa Viereck, 1905

subtricarinata Viereck, 1905

pegomyiae Brues, 1907

sarcophagae Gahan, 1914

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Distribution: 

England

Notes: 

added by Shaw (1983)

Aphaereta tenuicornis Nixon, 1939

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Distribution: 

England, Wales, Ireland

Genus Asobara Förster, 1863

Nomenclature: 

SPANISTA Förster, 1863

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Asobara tabida (Nees, 1834)

Nomenclature: 

Alysia tabida Nees, 1834

anomala (Thomson, 1895, Alysia)

crenulata (Fahringer, 1935, Phaenocarpa)

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Distribution: 
England, Ireland

Genus Aspilota Förster, 1863

Nomenclature: 

DIPIESTA Förster, 1863

EUSYNALDIS Zaykov & Fischer, 1982

SYNALDIS misident.

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Notes: 

Some distribution data from Stelfox and Graham (1951a). Some species treated as belonging to Dinotrema by other authors have been included in Aspilota in Taxapad (Yu et al. 2012) and there is clearly much work to be done in allocating species to the current generic concepts (see Wharton 1985). The late T. Munk was preparing a revision of the European species of Aspilotas. l. (see note under Dinotrema).

Aspilota acutidentata (Fischer, 1970)

Nomenclature: 

Synaldis acutidentata Fischer, 1970

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Distribution: 
England, Scotland
Notes: 

NMS, BMNH, det. Munk, added here; identified by T. Munk as a species of Dinotrema.

Aspilota anaphoretica Fischer, 1973

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Distribution: 
England, Scotland, Ireland
Notes: 

NMS, BMNH, det. Munk, added here

Aspilota blasii Fischer, 1973

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Distribution: 
England
Notes: 
BMNH, det. Munk, added here

Aspilota compressiventris Stelfox & Graham, 1951

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Distribution: 
England

Aspilota curta Marshall, 1895

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Distribution: 
England

Aspilota daemon Stelfox and Graham, 1948

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Distribution: 
England, Scotland, Ireland

Aspilota delicata Fischer, 1973

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Distribution: 
England, Scotland, Ireland
Notes: 
BMNH, det. Munk, added here

Aspilota efoveolata Thomson, 1895

Nomenclature: 

pneumatica Fischer, 1973

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Distribution: 
England, Scotland, Wales
Notes: 

NMS, BMNH, det. Munk, added here

Aspilota flagellaris Fischer, 1973

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Distribution: 
England, Scotland, Ireland
Notes: 
BMNH, det. Munk, added here

Aspilota fuscicornis (Haliday, 1838)

Nomenclature: 

Alysia fuscicornis Haliday, 1838

?minuta (Nees, 1812, Bassus)

exile (Ruthe, 1859, Orthostigma)

dilatata (Thomson, 1895, Alysia)

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Distribution: 
England, Scotland, Ireland

Aspilota globipes (Fischer, 1962)

Nomenclature: 

Synaldis globipes Fischer, 1962

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Distribution: 
England, Scotland
Notes: 

NMS, BMNH, det. Munk, added here

Aspilota imparidens Fischer, 1974

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Distribution: 
England, Scotland, Wales
Notes: 

NMS, BMNH, det. Munk, added here

Aspilota insolita (Tobias, 1962)

Nomenclature: 

Orthostigma insolita Tobias, 1962

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Distribution: 
England, Ireland
Notes: 
BMNH, det. Munk, added here

Aspilota intermediana Fischer, 1975

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Distribution: 
England
Notes: 

added by Notton (1991)

Aspilota iocosipecta Fischer, 1974

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Distribution: 
England, Scotland, Ireland
Notes: 

NMS, BMNH, det. Munk, added here

Aspilota macrops Stelfox & Graham, 1951

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Distribution: 
England, Ireland

Aspilota nidicola Hedqvist, 1972

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Distribution: 
England
Notes: 
BMNH, det. Munk, added here

Aspilota pillerensis Fischer, 1973

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Distribution: 
England
Notes: 
BMNH, det. Munk, added here

Aspilota ruficornis (Nees, 1834)

Nomenclature: 

Alysia ruficornis Nees, 1834

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Distribution: 
England, Scotland, Wales, Ireland

Aspilota stenogaster Stelfox & Graham, 1951

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Distribution: 
England

Aspilota tetragona Fischer, 1976

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Distribution: 
England, Scotland, Ireland
Notes: 

NMS, BMNH, det. Munk, added here

Aspilota vernalis Stelfox & Graham, 1951

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Distribution: 
England, Scotland, Ireland

Genus Chasmodon Haliday, 1838

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Chasmodon apterus (Nees, 1812)

Nomenclature: 

Bassus apterus Nees, 1812

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Distribution: 
England, Scotland, Wales, Ireland, Isle of Man

Genus Cratospila Förster, 1863

Nomenclature: 

HEDYLUS Marshall, 1891; synonymy by Papp (2009b)

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Cratospila circe (Haliday, 1838)

Nomenclature: 

Alysia circe Haliday, 1838

habilis (Marshall, 1891, Hedylus); synonymy by Papp (2009b)

annellata (Thomson, 1895, Alysia)

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Distribution: 
England, Ireland

Genus Dapsilarthra Förster, 1863

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Notes: 

Dapsilarthra has been used in a very broad sense to include Adelurola, Mesocrina, Heterolexis and Grammospila, and in a more narrow sense to include the last two genera. Achterberg (2014) is followed here in according each generic rank.

Dapsilarthra apii (Curtis, 1826)

Nomenclature: 

Alysia apii Curtis, 1826

laevipectus (Thomson, 1895, Alysia)

americana (Brues, 1907, Orthostigma)

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Distribution: 
England, Scotland, Ireland

Dapsilarthra sylvia (Haliday, 1839)

Nomenclature: 

Alysia sylvia Haliday, 1839

carpathica van Achterberg, 1983; synonymy by Achterberg (1997)

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Distribution: 

England, Scotland, Ireland

Genus Dinotrema Förster, 1863

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Notes: 

Wharton (1980) summarises the arguments against recognition of the genus Synaldis; Achterberg (1988b) likewise notes that the genus is defined only by the absence of fore wing vein 2-SR, which is known to be intraspecifically variable (Wharton 1980). Despite this, various authors attach great taxonomic weight to this venational character and maintain Synaldis as a valid genus (as reflected by the classification in Yu et al. 2012). Species that would be classified in Synaldis (acutidentata, concolor, distracta and globipes) are listed here in Aspilota and Dinotrema, according to Fauna Europaea. Some distribution data from Stelfox and Graham (1951b), Stelfox and Graham (1951a) and Achterberg (1988a). The late T. Munk was revising the European species of Aspilotas.l. and gave much helpful advice on the generic placements of the species occurring in Britain and Ireland, some of which is unpublished and will therefore differ from the generic combinations found in, e.g. Belokobylskij et al. (2003) and Taxapad (Yu et al. 2012). Munk also advised that although Aspilota is a well-defined genus, Dinotrema is not defined by any apomorphies and will be split up.

Species excluded from the British and Irish list:

[pusillum (Nees, 1812, Bassus)] T. Munk (pers. comm.) regarded this as an unidentified species; the type has been destroyed.

Subgenus Dinotrema Förster, 1863

Nomenclature: 

ASPILOTA misident.

SYNALDIS Förster, 1863

COLOBOMA Förster, 1863

SPANOMERIS Förster, 1863

SCOTIONEURUS Provancher, 1886

PTERUSA Fischer, 1958; synonymy by Achterberg and Vikberg (2014)

EUDINOSTIGMA Tobias, 1986; synonymy by Wharton (2002)

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Dinotrema (Dinotrema) aluum (Stelfox & Graham, 1950)

Nomenclature: 

Aspilota alua Stelfox & Graham, 1950

alva misspelling

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Distribution: 
Ireland

Dinotrema (Dinotrema) alysiae Munk & Peris-Felipo, 2013

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Distribution: 
England
Notes: 

added by Munk et al. (2013)

Dinotrema (Dinotrema) areolatum (Stelfox & Graham, 1950)

Nomenclature: 

Aspilota areolata Stelfox & Graham, 1950

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Distribution: 
England, Scotland

Dinotrema (Dinotrema) brevicorne (Nees, 1814)

Nomenclature: 

Bassus brevicornis Nees, 1814

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Distribution: 
England

Dinotrema (Dinotrema) brevissimicorne (Stelfox & Graham, 1948)

Nomenclature: 

Aspilota brevissimicornis Stelfox & Graham, 1948

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Distribution: 
Scotland, Ireland

Dinotrema (Dinotrema) compressum (Haliday, 1838)

Nomenclature: 

Alysia compressa Haliday, 1838

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Distribution: 
England

Dinotrema (Dinotrema) concinnum (Haliday, 1838)

Nomenclature: 

Alysia concinna Haliday, 1838

maximum (Fischer, 1962, Synaldis)

tyrrhena (Masi, 1933, Aspilota)

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Distribution: 
England, Ireland
Notes: 

Synaldis maximum Fischer, 1962 has been treated as a junior synonym by König (1972) and subsequent authors but T. Munk (pers. comm.) regarded this as a valid species of Dinotrema that may be found to occur in Britain.

Dinotrema (Dinotrema) concolor (Nees, 1812)

Nomenclature: 

Bassus concolor Nees, 1812

distractum (Ruthe, 1859, Orthostigma) unavailable

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Distribution: 
England, Ireland

Dinotrema (Dinotrema) crassicosta (Thomson, 1895)

Nomenclature: 

Alysia crassicosta Thomson, 1895

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Distribution: 
Scotland
Notes: 

NMS, det. Munk, added here

Dinotrema (Dinotrema) cratocera (Thomson, 1895)

Nomenclature: 

Alysia cratocera Thomson, 1895

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Distribution: 
Scotland
Notes: 
BMNH, det. Munk, added here

Dinotrema (Dinotrema) denticulatum (Stelfox & Graham, 1951)

Nomenclature: 

Aspilota denticulata Stelfox & Graham, 1951

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Distribution: 
Ireland

Dinotrema (Dinotrema) dimidiatum (Thomson, 1895)

Nomenclature: 

Alysia dimidiata Thomson, 1895

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Distribution: 
England
Notes: 
BMNH, det. Munk, added here

Dinotrema (Dinotrema) distractum (Nees, 1834)

Nomenclature: 

Alysia distracta Nees, 1834

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Dinotrema (Dinotrema) divisum (Stelfox & Graham, 1950)

Nomenclature: 

Aspilota divisa Stelfox & Graham, 1950

?aureliae (Fischer, 1973, Aspilota); tentative synonymy by Papp (2004b)

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Distribution: 
England, Scotland, Ireland

Dinotrema (Dinotrema) erythropum Förster, 1863

Nomenclature: 

praecipuum (Marshall, 1895, Aspilota)

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Distribution: 
England, Wales, Ireland

Dinotrema (Dinotrema) falsificum (Stelfox & Graham, 1950)

Nomenclature: 

Aspilota falsifica Stelfox & Graham, 1950

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Distribution: 
Scotland, Ireland

Dinotrema (Dinotrema) glabrum (Stelfox & Graham, 1951)

Nomenclature: 

Aspilota glabra Stelfox & Graham, 1951

venustum (Tobias, 1962, Aspilota)

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Distribution: 
England, Scotland, Ireland

Dinotrema (Dinotrema) insidiatrix (Marshall, 1895)

Nomenclature: 

Aspilota insidiatrix Marshall, 1895

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Distribution: 
England

Dinotrema (Dinotrema) insignis (Stelfox & Graham, 1950)

Nomenclature: 

Aspilota insignis Stelfox & Graham, 1950

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Distribution: 
England

Dinotrema (Dinotrema) iuxtanaeviam (Fischer, 1980)

Nomenclature: 

Aspilota iuxtanaeviam Fischer, 1978

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Distribution: 
England
Notes: 

added by Fischer (1980)

Dinotrema (Dinotrema) jaculans (Haliday, 1838)

Nomenclature: 

Alysia jaculans Haliday, 1838

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Distribution: 
England, Scotland, Ireland

Dinotrema (Dinotrema) latistigma (Fischer, 1962)

Nomenclature: 

Synaldis latistigma Fischer, 1962

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Distribution: 
England
Notes: 

NMS, det. Munk, added here; treated as a species of Eudinostigma in Fauna Europaea and Taxapad (Yu et al. 2012).

Dinotrema (Dinotrema) lineola (Thomson, 1895)

Nomenclature: 

Alysia lineola Thomson, 1895

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Distribution: 
England

Dinotrema (Dinotrema) liosoma (Stelfox & Graham,1951)

Nomenclature: 

Aspilota liosoma Stelfox & Graham,1951

caudatum (Thomson, 1895, Aspilota) preocc.

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Dinotrema (Dinotrema) mesocaudatum van Achterberg, 1988

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Distribution: 
England
Notes: 

added by Achterberg (1988b)

Dinotrema (Dinotrema) microcera (Thomson, 1895)

Nomenclature: 

Alysia microcera Thomson, 1895

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Distribution: 
England, Scotland, Wales, Ireland
Notes: 
BMNH, det. Munk, added here

Dinotrema (Dinotrema) necrophilum (Hedqvist, 1972)

Nomenclature: 

Aspilota necrophila Hedqvist, 1972

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Distribution: 
England
Notes: 

Added by Disney and Munk (2005) and transferred from Aspilota.

Dinotrema (Dinotrema) nervosum (Haliday, 1833)

Nomenclature: 

Alysia nervosa Haliday, 1833

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Distribution: 
England, Scotland, Ireland

Dinotrema (Dinotrema) pulvinatum (Stelfox & Graham, 1949)

Nomenclature: 

Aspilota pulvinata Stelfox & Graham, 1949

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Distribution: 
England, Scotland, Wales, Ireland
Notes: 

Treated as a species of Eudinostigma in Fauna Europaea and Taxapad (Yu et al. 2012).

Dinotrema (Dinotrema) ruficollis (Stelfox & Graham, 1950)

Nomenclature: 

Aspilota ruficollis Stelfox & Graham, 1950

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Distribution: 
England, Ireland
Notes: 

Distribution data from Achterberg (1988b).

Dinotrema (Dinotrema) semicompressum (Stelfox & Graham, 1949)

Nomenclature: 

Aspilota semicompressa Stelfox & Graham, 1949

parapunctatum (Fischer, 1976, Aspilota); synonymy by Papp (2004b)

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Distribution: 
England, Scotland, Ireland

Dinotrema (Dinotrema) sphaerimembre (Fischer, 1973)

Nomenclature: 

Aspilota sphaerimembris Fischer, 1973

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Distribution: 
Scotland
Notes: 

NMS, det. Munk, added here

Dinotrema (Dinotrema) sternaulicum (Fischer, 1973)

Nomenclature: 

Aspilota sternaulica Fischer, 1973

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Distribution: 
England, Scotland
Notes: 

NMS, det. Munk, added here

Dinotrema (Dinotrema) tauricum (Telenga, 1935)

Nomenclature: 

Aspilota taurica Telenga, 1935

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Distribution: 
England
Notes: 

added by Achterberg (1988b)

Dinotrema (Dinotrema) vesparum (Stelfox, 1943)

Nomenclature: 

Aspilota vesparum Stelfox, 1943

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Distribution: 
England, Scotland

Dinotrema (Dinotrema) vituperatum (Fischer, 1974)

Nomenclature: 

Aspilota vituperata Fischer, 1974

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Distribution: 
England, Scotland
Notes: 

NMS, det. Munk, added here

Subgenus Leptotrema van Achterberg, 1988

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Notes: 

Described as a separate genus by Achterberg (1988b), who gives distribution data for the one included species; treated as a subgenus of Dinotrema by Wharton (2002).

Dinotrema (Leptotrema) dentifemur (Stelfox, 1943)

Nomenclature: 

Aspilota dentifemur Stelfox, 1943

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Distribution: 
England, Scotland, Ireland

Subgenus Prosapha Förster, 1863

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Dinotrema (Prosapha) speculum (Haliday, 1838)

Nomenclature: 

Alysia speculum Haliday, 1838

venustum (Haliday, 1838, Alysia)

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Distribution: 
England, Ireland

Genus Grammospila Förster, 1863

Nomenclature: 

PARAORTHOSTIGMA Königsmann, 1972

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Grammospila isabella (Haliday, 1838)

Nomenclature: 

Alysia isabella Haliday, 1838

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Distribution: 
England, Scotland

Grammospila rufiventris (Nees, 1812)

Nomenclature: 

Bassus rufiventris Nees, 1812

flaviventris (Haliday, 1838, Alysia)

ochrogaster (Szépligeti, 1898, Phaenocarpa)

fuscula Griffiths, 1968

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Distribution: 
England, Scotland, Ireland, Isle of Man
Notes: 

Grammospila fuscula (Griffiths, 1968, Dapsilarthra) removed from synonymy by Achterberg (2014)​.

Genus Heterolexis Förster, 1863

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Heterolexis balteata (Thomson, 1895)

Nomenclature: 

Alysia balteata Thomson, 1895

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Distribution: 
England, Scotland, Wales, Ireland

Heterolexis dictynna (Marshall, 1895)

Nomenclature: 

Adelura dictynna Marshall, 1895

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Distribution: 
England, Scotland, Ireland

Genus Idiasta Förster, 1863

Nomenclature: 

EUPHAENOCARPA Tobias, 1975

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Idiasta dichrocera Königsmann, 1960

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Distribution: 
England
Notes: 

NMS, det. Shaw, added here

Idiasta maritima (Haliday, 1838)

Nomenclature: 

Alysia maritima Haliday, 1838

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Distribution: 
England, Scotland, Ireland

Idiasta nephele (Haliday, 1838)

Nomenclature: 

Alysia nephele Haliday, 1838

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Distribution: 
England, Scotland

Genus Mesocrina Förster, 1863

Nomenclature: 

PSEUDOMESOCRINA Königsmann, 1959

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Mesocrina indagatrix Förster, 1863

Nomenclature: 

venatrix Marshall, 1895

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Distribution: 
England

Genus Orthostigma Ratzeburg, 1844

Nomenclature: 

DELOCARPA Förster, 1863

ISCHNOCARPA Förster, 1863

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Orthostigma cratospilum (Thompson, 1895)

Nomenclature: 

Alysia cratospila Thompson, 1895

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Distribution: 
England
Notes: 

added by Godfray and Achterberg (2015)

Orthostigma longicorne Königsmann, 1969

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Distribution: 
Ireland

Orthostigma maculipes (Haliday, 1838)

Nomenclature: 

Alysia maculipes Haliday, 1838

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Distribution: 
England, Ireland

Orthostigma pumilum (Nees, 1834)

Nomenclature: 

Alysia pumila Nees, 1834

flavipes (Ratzeburg, 1844, Aphidius)

fulvipes Rondani, 1876 nom. nud.

brunnipes Ratzeburg, 1852

bruneipes Dalla Torre, 1898

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Distribution: 
England, Ireland

Genus Panerema Förster, 1863

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Notes: 

Distribution data from Achterberg (1988b).

Panerema fulvicornis (Haliday, 1838)

Nomenclature: 

Alysia fulvicornis Haliday, 1838

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Distribution: 
Wales, Ireland

Panerema inops Förster, 1863

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Distribution: 
England, Wales, Ireland

Genus Pentapleura Förster, 1863

Nomenclature: 

OPISENDEA Förster, 1863

GNATHOSPILA Fischer, 1966

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Pentapleura angustula (Haliday, 1838)

Nomenclature: 

Alysia angustula Haliday, 1838

laevipleuris (Tobias, 1962, Aspilota)

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Distribution: 
England, Scotland, Ireland

Pentapleura fuliginosa (Haliday, 1838)

Nomenclature: 

Alysia fuliginosa Haliday, 1838

carinata (Thomson, 1895, Alysia)

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Distribution: 
England, Wales, Ireland

Pentapleura pumilio (Nees, 1812)

Nomenclature: 

Bassus pumilio Nees, 1812

triticaphis (Fitch, 1861, Toxares)

mesocrinoides Goidanich, 1936

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Distribution: 
England, Scotland, Wales, Ireland

Genus Phaenocarpa Förster, 1863

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Notes: 

Some taxonomic and distribution data from Achterberg (1997).

Subgenus Homophyla Förster, 1863

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Notes: 

species of Phaenocarpa (Homophyla) excluded from the British and Irish list:

[pegomyiae Marshall, 1898] Listed as British by Lyle (1933) but we have not seen any British or Irish specimens.

Phaenocarpa (Homophyla) pullata (Haliday, 1838)

Nomenclature: 

Alysia pullata Haliday, 1838

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Distribution: 
England, Ireland

Subgenus Phaenocarpa Förster, 1863

Nomenclature: 

IDIOLEXIS Förster, 1863; synonymy by Wharton (2002)

MESOTHESIS Förster, 1863

SATHRA Förster, 1863

ASYNAPHES Provancher, 1886

KAHLIA Ashmead, 1900

HOLCALYSIA Cameron, 1905

STIRALYSIA Cameron, 1910

RHOPALONEURA Stelfox, 1941

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Phaenocarpa (Phaenocarpa) canaliculata Stelfox, 1941

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Distribution: 
England, Ireland

Phaenocarpa (Phaenocarpa) conspurcator (Haliday, 1838)

Nomenclature: 

Alysia conspurcator Haliday, 1838

arctica (Thomson, 1895, Alysia)

tatrica Niezabitowski, 1910

remota Papp, 1981

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Distribution: 
England, Scotland, Wales, Ireland

Phaenocarpa (Phaenocarpa) eugenia (Haliday, 1838)

Nomenclature: 

Alysia eugenia Haliday, 1838

pectoralis (Zetterstedt, 1838, Alysia)

orbicularis Gurasashvilli, 1983

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Distribution: 
England, Scotland, Ireland

Phaenocarpa (Phaenocarpa) eunice (Haliday, 1838)

Nomenclature: 

Alysia eunice Haliday, 1838

nimia Stelfox, 1941

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Distribution: 
England, Wales, Ireland

Phaenocarpa (Phaenocarpa) flavipes (Haliday, 1838)

Nomenclature: 

Alysia flavipes Haliday, 1838

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Distribution: 
England, Ireland

Phaenocarpa (Phaenocarpa) frequentator (Zetterstedt, 1838)

Nomenclature: 

Alysia frequentator Zetterstedt, 1838

frequentatrix (Schulz, 1906, Alysia)

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Distribution: 
Wales
Notes: 

added by Godfray and Achterberg (2015)

Phaenocarpa (Phaenocarpa) galatea (Haliday, 1838)

Nomenclature: 

Alysia galatea Haliday, 1838

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Distribution: 
Scotland, Ireland

Phaenocarpa (Phaenocarpa) helophilae van Achterberg, 1998

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Distribution: 
England
Notes: 

added by Achterberg (1998)

Phaenocarpa (Phaenocarpa) livida (Haliday, 1838)

Nomenclature: 

Alysia livida Haliday, 1838

debilis (Förster, 1863, Sathra)

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Distribution: 
England, Ireland

Phaenocarpa (Phaenocarpa) luteipes Stelfox, 1950

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Distribution: 
Ireland

Phaenocarpa (Phaenocarpa) maria (Haliday, 1838)

Nomenclature: 

Alysia maria Haliday, 1838

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Distribution: 
England, Ireland

Phaenocarpa (Phaenocarpa) nina (Haliday, 1838)

Nomenclature: 

Alysia nina Haliday, 1838

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Distribution: 
Scotland
Notes: 

Not listed in Huddleston (1978), although described from Scottish material (Achterberg 1998).

Phaenocarpa (Phaenocarpa) notabilis Stelfox, 1944

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Distribution: 
Scotland, Ireland

Phaenocarpa (Phaenocarpa) picinervis (Haliday, 1838)

Nomenclature: 

Alysia picinervis Haliday, 1838

americana Ashmead, 1889

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Distribution: 
England, Scotland, Ireland

Phaenocarpa (Phaenocarpa) pratellae (Curtis, 1826)

Nomenclature: 

Alysia pratellae Curtis, 1826

piceator (Zetterstedt, 1838, Alysia)

psalliotae Telenga, 1935; synonymy by Achterberg (2014)

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Distribution: 
England, Scotland

Phaenocarpa (Phaenocarpa) punctigera (Haliday, 1838)

Nomenclature: 

Alysia punctigera Haliday, 1838

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Distribution: 

England, Ireland

Notes: 

Listed as a species of Idiolexis in Fauna Europaea.

Phaenocarpa (Phaenocarpa) ruficeps (Nees, 1812)

Nomenclature: 

Bassus ruficeps Nees, 1812

testacea (Nees, 1812, Bassus)

gracilis (Curtis, 1826, Alysia)

pallida (Curtis, 1826, Alysia)

agricolator (Zetterstedt, 1838, Alysia)

oculator (Ratzeburg, 1848, Alysia)

rubriceps (Provancher, 1883, Alysia) preocc.

rubricepes Provancher, 1888

testaceipes (Cameron, 1905, Holcalysia)

divergens Fischer, 1975

fervida Fischer, 1975

incerta Fischer, 1975

meritoria Papp, 1981

ferga Papp, 1982

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Distribution: 
England, Scotland, Wales, Ireland, Isle of Man

Phaenocarpa (Phaenocarpa) tacita Stelfox, 1941

Nomenclature: 

caucasica Gurasashvili, 1983 preocc.

caucasicola Tobias, 1986

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Distribution: 
Ireland

Phaenocarpa (Phaenocarpa) trisulcata Stelfox, 1950

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Distribution: 
Ireland

Genus Syncrasis Förster, 1863

Nomenclature: 

PHAENOLYTA Förster, 1863

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Syncrasis fucicola (Haliday, 1838)

Nomenclature: 

Alysia fucicola Haliday, 1838

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Distribution: 
Ireland

Syncrasis halidayi (Förster, 1863)

Nomenclature: 

Phaenolyta halidayi Förster, 1863

fuscipes preocc., unavailable

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Distribution: 
England, Ireland

Genus Tanycarpa Förster, 1863

Nomenclature: 

ACROBELA Förster, 1863; synonymy by Wharton (2002)

EPICLISTA Förster, 1863

HYPOSTROPHA Förster, 1863

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Tanycarpa bicolor (Nees, 1812)

Nomenclature: 

Bassus bicolor Nees, 1812

ancilla (Haliday, 1838, Alysia)

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Distribution: 
England, Ireland

Tanycarpa gracilicornis (Nees, 1812)

Nomenclature: 

Bassus gracilicornis Nees, 1812

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Distribution: 
Ireland

Tanycarpa mitis Stelfox, 1941

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Distribution: 
England, Ireland

Tanycarpa punctata van Achterberg, 1976

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Tanycarpa rufinotata (Haliday, 1838)

Nomenclature: 

Alysia rufinotata Haliday, 1838

carinata (Förster, 1863, Acrobela)

erythrogaster (Förster, 1863, Epiclista)

foersteri (Shenefelt, 1974, Alysia)

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Distribution: 
England, Ireland

Genus Trachyusa Ruthe, 1854

Nomenclature: 

COSMIOCARPA Förster, 1863

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Notes: 

Distribution data from Achterberg and O'Connor (1990) and the collections of NMS.

Trachyusa aurora (Haliday, 1838)

Nomenclature: 

Alysia aurora Haliday, 1838

nigriceps Ruthe, 1854

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Distribution: 
England, Ireland

Trachyusa nigrothoracica van Achterberg & O’Connor, 1990

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Distribution: 

England, Ireland

Notes: 

added by Achterberg and O'Connor (1990)

Tribe Dacnusini Förster, 1863

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Notes: 

The classification of the Dacnusini was revised by Griffiths (Griffiths 1964, Griffiths 1967a, Griffiths 1967b, Griffiths 1967c, Griffiths 1968a, Griffiths 1968b) in a pioneering treatment that was one of the first applications of explicit phylogenetic methods in taxonomy. It was also unusual in being based on extensive reared material. Griffiths’ classification is followed here although one of the consequences of his strict adherence to phylogenetic principles is that Chorebus and Dacnusa are very large genera. Also, as Griffiths realised, the definition of Exotela is unsatisfactory as it lacks clear apomorphic characters. This group of insects had previously been revised by Nixon (Nixon 1943, Nixon 1944, Nixon 1945, Nixon 1946, Nixon 1948, Nixon 1949, Nixon 1954), who recognised a greater number of genera, and some authors (Tobias 1986, Tobias 1998, Perepechayenko 2000, Fischer et al. 2004) have resurrected Nixon’s genera or used them as subgenera. This seems a retrograde step given Griffiths’ clear statements of phylogenetic hypotheses and is not used here. In both Chorebus and Dacnusa, Griffiths defined a series of species groups that with further research should form the basis of better classifications of these genera. Griffiths (1964) gives a key to genera and Wharton (1997) is also very helpful. The Coelinius genus group (Coelinius, Trachionus, Epimicta, Aristelix, Laotris, Sarops and Synelix) is generally agreed to be monophyletic and was not revised at species level by Nixon or Griffiths. Distribution data from Griffiths’ and Nixon’s revisions (loc. cit.) and NMS.

Genus Amyras Nixon, 1943

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Amyras clandestina (Haliday, 1839)

Nomenclature: 

Alysia clandestina Haliday, 1839

quadridentata (Thomson, 1895, Dacnusa)

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Distribution: 
Ireland

Genus Aristelix Nixon, 1943

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Aristelix phaenicura (Haliday, 1839)

Nomenclature: 

Alysia phaenicura Haliday, 1839

phoenicura misspelling

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Distribution: 
England, Ireland

Genus Chaenusa Haliday, 1839

Nomenclature: 

CHOREBIDEA Viereck, 1914

CHOREBIDEA Nixon, 1943 preocc.

CHOREBIDELLA Riegel, 1947

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Notes: 
No recent revision of this genus.

Chaenusa conjungens (Nees, 1811)

Nomenclature: 

Bracon conjungens Nees, 1811

conjugens misspelling

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Distribution: 
England, Scotland, Ireland

Chaenusa elongata Stelfox, 1957

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Distribution: 
Ireland

Chaenusa limoniadum (Marshall, 1896)

Nomenclature: 

Chorebus limoniadum Marshall, 1896

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Distribution: 
England
Notes: 

Transferred from Chorebus by Perepechayenko (2000).

Chaenusa lymphata (Haliday, 1839)

Nomenclature: 

Alysia lymphata Haliday, 1839

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Distribution: 
England, Ireland
Notes: 

Transferred from Chorebus by Achterberg (1997).

Chaenusa naiadum (Haliday, 1839)

Nomenclature: 

Alysia naiadum Haliday, 1839

naiadum (Curtis, 1837, Chorebus) nom. nud.

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Distribution: 
England, Scotland, Ireland

Chaenusa nereidum (Haliday, 1839)

Nomenclature: 

Alysia nereidum Haliday, 1839

nereidum (Curtis, 1837, Chorebus) nom. nud.

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Distribution: 
England, Ireland

Chaenusa opaca Stelfox, 1957

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Distribution: 
Ireland

Genus Chorebus Haliday, 1833

Nomenclature: 

AMETRIA Förster, 1863

GYROCAMPA Förster, 1863

PHAENOLEXIS Förster, 1863

STIPHROCERA Förster, 1863

DIPLUSIA Ruthe, 1882

ETRIPTES Nixon, 1943

PARAGYROCAMPA Tobias, 1962

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Chorebus abaris (Nixon, 1943)

Nomenclature: 

Dacnusa abaris Nixon, 1943

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Distribution: 
England, Scotland

Chorebus abnormiceps (Nixon, 1943)

Nomenclature: 

Dacnusa abnormiceps Nixon, 1943

quadriceps (Nixon, 1941, Dacnusa) preocc.

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Distribution: 
England

Chorebus agraules (Nixon, 1945)

Nomenclature: 

Dacnusa agraules Nixon, 1945

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Distribution: 
England

Chorebus albipes (Haliday, 1839)

Nomenclature: 

Alysia albipes Haliday, 1839

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Distribution: 
England, Scotland, Ireland

Chorebus alecto (Morley, 1924)

Nomenclature: 

Rhizarcha alecto Morley, 1924

turissa (Nixon, 1937, Dacnusa)

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Distribution: 
England, Scotland, Ireland

Chorebus alua (Nixon, 1944)

Nomenclature: 

Dacnusa alua Nixon, 1944

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Distribution: 
Ireland

Chorebus amasis (Nixon, 1945)

Nomenclature: 

Dacnusa amasis Nixon, 1945

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Distribution: 
England

Chorebus ampliator (Nees, 1834)

Nomenclature: 

Alysia ampliator Nees, 1834

nigricornis (Förster, 1863, Stiphrocera)

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Distribution: 
England, Scotland, Ireland

Chorebus anasellus (Stelfox, 1952)

Nomenclature: 

Dacnusa anasella Stelfox, 1952

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Distribution: 
Scotland, Ireland

Chorebus angelicae (Nixon, 1945)

Nomenclature: 

Dacnusa angelicae Nixon, 1945

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Distribution: 
England

Chorebus anita (Nixon, 1943)

Nomenclature: 

Dacnusa anita Nixon, 1943

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Distribution: 
England, Scotland

Chorebus aphantus (Marshall, 1895)

Nomenclature: 

Dacnusa aphanta Marshall, 1895

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Distribution: 
England, Scotland, Wales, Ireland

Chorebus apollyon (Morley, 1924)

Nomenclature: 

Dacnusa apollyon Morley, 1924

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Distribution: 
England, Ireland

Chorebus ares (Nixon, 1944)

Nomenclature: 

Dacnusa ares Nixon, 1944

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Distribution: 
England

Chorebus armida (Nixon, 1945)

Nomenclature: 

Dacnusa armida Nixon, 1945

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Distribution: 
England, Scotland, Wales

Chorebus artemisiellus Griffiths, 1968

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Distribution: 
England

Chorebus asramenes (Nixon, 1943)

Nomenclature: 

Dacnusa asramenes Nixon, 1943

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Distribution: 
England, Ireland

Chorebus avesta (Nixon, 1944)

Nomenclature: 

Dacnusa avesta Nixon, 1944

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Distribution: 
England, Scotland

Chorebus bathyzonus (Marshall, 1895)

Nomenclature: 

Dacnusa bathyzona Marshall, 1895

ornatus (Telenga, 1935, Dacnusa)

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Distribution: 
England, Ireland

Chorebus bensoni (Nixon, 1943)

Nomenclature: 

Dacnusa bensoni Nixon, 1943

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Distribution: 
England, Scotland

Chorebus bres (Nixon, 1944)

Nomenclature: 

Dacnusa bres Nixon, 1944

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Distribution: 
England, Ireland

Chorebus brevicornis (Thomson, 1895)

Nomenclature: 

Dacnusa brevicornis Thomson, 1895

chrysippe (Nixon, 1944, Dacnusa)

ea (Nixon, 1944, Dacnusa)

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Distribution: 
England, Scotland, Ireland

Chorebus caelebs (Nixon, 1944)

Nomenclature: 

Dacnusa caelebs Nixon, 1944

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Distribution: 
England

Chorebus calthae Griffiths, 1967

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Distribution: 
England

Chorebus cambricus Griffiths, 1968

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Distribution: 
Wales

Chorebus cinctus (Haliday, 1839)

Nomenclature: 

Alysia cincta Haliday, 1839

castaneiventris (Thomson, 1895, Dacnusa)

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Distribution: 
England, Ireland

Chorebus coxator (Thomson, 1895)

Nomenclature: 

Dacnusa coxator Thomson, 1895

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Distribution: 
England, Scotland, Wales

Chorebus crassipes (Stelfox, 1954)

Nomenclature: 

Dacnusa crassipes Stelfox, 1954

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Distribution: 
Scotland, Ireland

Chorebus credne (Nixon, 1944)

Nomenclature: 

Dacnusa credne Nixon, 1944

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Distribution: 
England, Scotland, Ireland

Chorebus crenulatus (Thomson, 1895)

Nomenclature: 

Dacnusa crenulata Thomson, 1895

elegantulus (Nixon, 1937, Dacnusa)

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Distribution: 
England, Scotland, Ireland

Chorebus crocale (Nixon, 1945)

Nomenclature: 

Dacnusa crocale Nixon, 1945

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Distribution: 
England

Chorebus cubocephalus (Telenga, 1934)

Nomenclature: 

Rhizarcha cubocephala Telenga, 1934

cyclops (Nixon, 1937, Dacnusa); synonymy by Tobias (1986)

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Distribution: 
Ireland

Chorebus cylindricus (Telenga, 1934)

Nomenclature: 

Dacnusa cylindrica Telenga, 1934

cybele (Nixon, 1937, Dacnusa); synonymy by Tobias (1986)

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Distribution: 
England, Scotland

Chorebus cytherea (Nixon, 1937)

Nomenclature: 

Dacnusa cytherea Nixon, 1937

calliope (Nixon, 1944, Dacnusa)

tesmia (Nixon, 1944, Dacnusa)

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Distribution: 
England, Scotland, Ireland

Chorebus dagda (Nixon, 1943)

Nomenclature: 

Dacnusa dagda Nixon, 1943

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Distribution: 
England

Chorebus daimenes (Nixon, 1945)

Nomenclature: 

Dacnusa daimenes Nixon, 1945

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Distribution: 
England, Scotland

Chorebus deione (Nixon, 1944)

Nomenclature: 

Dacnusa deione Nixon, 1944

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Distribution: 
England, Scotland

Chorebus didas (Nixon, 1944)

Nomenclature: 

Dacnusa didas Nixon, 1944

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Distribution: 
England, Scotland

Chorebus difficilis Griffiths, 1968

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Distribution: 
England

Chorebus diremtus (Nees, 1834)

Nomenclature: 

Alysia diremta Nees, 1834

diremptus (Haliday, 1839, Alysia)

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Distribution: 
England, Ireland

Chorebus dirona (Nixon, 1945)

Nomenclature: 

Dacnusa dirona Nixon, 1945

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Distribution: 
England, Ireland

Chorebus enephes (Nixon, 1945)

Nomenclature: 

Dacnusa enephes Nixon, 1945

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Distribution: 
England

Chorebus eros (Nixon, 1937)

Nomenclature: 

Dacnusa eros Nixon, 1937

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Distribution: 
England, Scotland, Ireland

Chorebus esbelta (Nixon, 1937)

Nomenclature: 

Dacnusa esbelta Nixon, 1937

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Distribution: 
England, Scotland, Ireland

Chorebus euryale (Nixon, 1944)

Nomenclature: 

Dacnusa euryale Nixon, 1944

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Distribution: 
England

Chorebus fallaciosae Griffiths, 1967

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Distribution: 
England

Chorebus fallax (Nixon, 1937)

Nomenclature: 

Dacnusa fallax Nixon, 1937

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Distribution: 
England, Scotland, Ireland

Chorebus flavipes (Goureau, 1851)

Nomenclature: 

Dacnusa flavipes Goureau, 1851

raissa (Nixon, 1937, Dacnusa)

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Distribution: 
England, Scotland, Ireland

Chorebus fordi (Nixon, 1954)

Nomenclature: 

Dacnusa fordi Nixon, 1954

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Distribution: 
England

Chorebus foveolus (Haliday, 1839)

Nomenclature: 

Alysia foveola Haliday, 1839

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Distribution: 
England, Scotland, Wales, Ireland

Chorebus fuscipennis (Nixon, 1937)

Nomenclature: 

Dacnusa fuscipennis Nixon, 1937

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Distribution: 
England

Chorebus gedanensis (Ratzeburg, 1852)

Nomenclature: 

Alysia gedanensis Ratzeburg, 1852

anguligena (Nixon, 1937, Dacnusa)

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Distribution: 
England

Chorebus glaber (Nixon, 1944)

Nomenclature: 

Dacnusa glabra Nixon, 1944

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Distribution: 
England, Ireland

Chorebus glabriculus (Thomson, 1895)

Nomenclature: 

Dacnusa glabricula Thomson, 1895

cortipalpis (Nixon, 1937, Dacnusa)

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Distribution: 
England, Scotland, Ireland

Chorebus hilaris Griffiths, 1967

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Distribution: 
England

Chorebus hirtigena Stelfox, 1957

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Distribution: 
Ireland

Chorebus humeralis Griffiths, 1968

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Distribution: 
Ireland

Chorebus incertus (Goureau, 1851)

Nomenclature: 

Dacnusa incerta Goureau, 1851

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Distribution: 
England

Chorebus iphias (Nixon, 1943)

Nomenclature: 

Dacnusa iphias Nixon, 1943

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Distribution: 
England

Chorebus kama (Nixon, 1945)

Nomenclature: 

Dacnusa kama Nixon, 1945

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Distribution: 
England

Chorebus lanigerus (Stelfox, 1957)

Nomenclature: 

Gyrocampa lanigera Stelfox, 1957

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Distribution: 
Ireland

Chorebus lar (Morley, 1924)

Nomenclature: 

Dacnusa lar Morley, 1924

innanus (Nixon, 1943, Dacnusa)

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Distribution: 
England

Chorebus larides (Nixon, 1944)

Nomenclature: 

Dacnusa larides Nixon, 1944

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Distribution: 
England, Ireland

Chorebus lateralis (Haliday, 1839)

Nomenclature: 

Alysia lateralis Haliday, 1839

fuscula (Haliday, 1839, Alysia)

albicoxa (Thomson, 1895, Dacnusa)

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Distribution: 
England, Scotland, Wales, Ireland

Chorebus leptogaster (Haliday, 1839)

Nomenclature: 

Alysia leptogaster Haliday, 1839

naenia (Morley, 1924, Dacnusa)

dinae (Burghele, 1960, Dacnusa)

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Distribution: 
England, Ireland

Chorebus longicornis (Nees, 1811)

Nomenclature: 

Bracon longicornis Nees, 1811

affinis (Nees, 1812, Bassus)

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Distribution: 
England, Scotland, Ireland
Notes: 

Synonymy first established in Shenefelt (1974) but overlooked until O'Connor et al. (1999).

Chorebus lugubris (Nixon, 1937)

Nomenclature: 

Dacnusa lugubris Nixon, 1937

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Distribution: 
England, Ireland

Chorebus luzulae Griffiths, 1966

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Distribution: 
Scotland
Notes: 

added by Godfray and Bland (2011)

Chorebus lychnidis Griffiths, 1967

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Distribution: 
England

Chorebus maculigastra Shenefelt, 1974

Nomenclature: 

maculata (Nixon, 1944, Dacnusa) preocc.

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Distribution: 
England, Ireland

Chorebus merellus (Nixon, 1937)

Nomenclature: 

Dacnusa merella Nixon, 1937

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Distribution: 
England, Scotland, Ireland

Chorebus merion (Nixon, 1945)

Nomenclature: 

Dacnusa merion Nixon, 1945

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Distribution: 
England

Chorebus miodes (Nixon, 1949)

Nomenclature: 

Gyrocampa miodes Nixon, 1949

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Distribution: 
England, Ireland

Chorebus misellus (Marshall, 1895)

Nomenclature: 

Dacnusa misella Marshall, 1895

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Distribution: 
England

Chorebus mitrus (Nixon, 1945)

Nomenclature: 

Dacnusa mitra Nixon, 1945

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Distribution: 
England, Scotland, Ireland

Chorebus nanus (Nixon, 1943)

Nomenclature: 

Dacnusa nana Nixon, 1943

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Distribution: 
England

Chorebus navicularis (Nees, 1812)

Nomenclature: 

Bassus navicularis Nees, 1812

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Distribution: 
England

Chorebus nerissus (Nixon, 1937)

Nomenclature: 

Dacnusa nerissa Nixon, 1937

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Distribution: 
England

Chorebus nigriscaposus (Nixon, 1949)

Nomenclature: 

Gyrocampa nigriscaposa Nixon, 1949

propodealis (Nixon, 1949, Gyrocampa)

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Distribution: 
Ireland

Chorebus ninella (Nixon, 1945)

Nomenclature: 

Dacnusa ninella Nixon, 1945

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Distribution: 
England, Scotland, Ireland

Chorebus nobilis Griffiths, 1968

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Distribution: 
England, Ireland

Chorebus nomia (Nixon, 1937)

Nomenclature: 

Dacnusa nomia Nixon, 1937

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Distribution: 
England, Ireland

Chorebus nydia (Nixon, 1937)

Nomenclature: 

Dacnusa nydia Nixon, 1937

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Distribution: 
England, Scotland, Ireland

Chorebus orbiculatae Griffiths, 1967

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Distribution: 
England, Ireland

Chorebus ovalis (Marshall, 1896)

Nomenclature: 

Dacnusa ovalis Marshall, 1896

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Distribution: 
England, Ireland

Chorebus parvungula (Thomson, 1895)

Nomenclature: 

Dacnusa parvungula Thomson, 1895

acco (Nixon, 1943, Dacnusa)

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Distribution: 
England, Scotland, Ireland

Chorebus perkinsi (Nixon, 1944)

Nomenclature: 

Dacnusa perkinsi Nixon, 1944

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Distribution: 
England

Chorebus petiolatus (Nees, 1834)

Nomenclature: 

Alysia petiolata Nees, 1834

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Distribution: 
England, Scotland, Ireland

Chorebus phaedra (Nixon, 1937)

Nomenclature: 

Dacnusa phaedra Nixon, 1937

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Distribution: 
England, Scotland

Chorebus pimpinellae Griffiths, 1967

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Distribution: 
England

Chorebus pione (Nixon, 1944)

Nomenclature: 

Dacnusa pione Nixon, 1944

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Distribution: 
England

Chorebus poemyzae Griffiths, 1968

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Distribution: 
England, Scotland

Chorebus posticus (Haliday, 1839)

Nomenclature: 

Alysia postica Haliday, 1839

gracilis (Nees, 1834, Alysia); synonymy by Achterberg (1997)

egregia (Marshall, 1895, Dacnusa)

dentatus (Tobias, 1962, Dacnusa); synonymy by Tobias (1986)

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Distribution: 
England, Scotland, Ireland

Chorebus pulverosus (Haliday, 1839)

Nomenclature: 

Alysia pulverosa Haliday, 1839

marsyas (Nixon, 1937, Dacnusa)

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Distribution: 
England, Ireland

Chorebus punctum (Goureau, 1851)

Nomenclature: 

Dacnusa punctum Goureau, 1851

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Distribution: 
England, Scotland, Ireland

Chorebus resa (Nixon, 1937)

Nomenclature: 

Dacnusa resa Nixon, 1937

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Distribution: 
England

Chorebus rhanis (Nixon, 1943)

Nomenclature: 

Dacnusa rhanis Nixon, 1943

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Distribution: 
Scotland

Chorebus risilis (Nixon, 1949)

Nomenclature: 

Gyrocampa risilis Nixon, 1949

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Distribution: 
Ireland

Chorebus rondanii (Giard, 1904)

Nomenclature: 

Dacnusa rondanii Giard, 1904

galbus (Nixon, 1944, Dacnusa)

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Distribution: 
England

Chorebus rotundiventris (Thomson, 1895)

Nomenclature: 

Dacnusa rotundiventris Thomson, 1895

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Distribution: 
Ireland

Chorebus rousseaui (Schulz, 1907)

Nomenclature: 

Dacnusa rousseaui Schulz, 1907

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Notes: 
The status of this taxon needs further research.

Chorebus ruficollis (Stelfox, 1957)

Nomenclature: 

Gyrocampa ruficollis Stelfox, 1957

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Distribution: 
Ireland

Chorebus rufimarginatus (Stelfox, 1954)

Nomenclature: 

Dacnusa rufimarginata Stelfox, 1954

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Distribution: 
Ireland

Chorebus scabiosae Griffiths, 1967

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Distribution: 
England

Chorebus scabrifossa Stelfox, 1957

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Distribution: 
Ireland

Chorebus selene (Nixon, 1937)

Nomenclature: 

Dacnusa selene Nixon, 1937

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Distribution: 
England, Scotland

Chorebus senilis (Nees, 1812)

Nomenclature: 

Bassus senilis Nees, 1812

tomentosus (Thomson, 1895, Dacnusa)

nemesis (Morley, 1924, Dacnusa)

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Distribution: 
England, Scotland, Wales, Ireland

Chorebus serus (Nixon, 1937)

Nomenclature: 

Dacnusa sera Nixon, 1937

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Distribution: 
England

Chorebus siniffa (Nixon, 1937)

Nomenclature: 

Dacnusa siniffa Nixon, 1937

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Distribution: 
England, Ireland

Chorebus solstitialis (Stelfox, 1952)

Nomenclature: 

Dacnusa solstitialis Stelfox, 1952

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Distribution: 
England

Chorebus spenceri Griffiths, 1964

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Distribution: 
England, Scotland

Chorebus striola Stelfox, 1957

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Distribution: 
Ireland

Chorebus sylvestris Griffiths, 1967

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Distribution: 
England, Scotland, Ireland

Chorebus talaris (Haliday, 1839)

Nomenclature: 

Alysia talaris Haliday, 1839

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Distribution: 
England, Scotland, Ireland

Chorebus tamiris (Nixon, 1943)

Nomenclature: 

Dacnusa tamiris Nixon, 1943

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Distribution: 
England

Chorebus tamsi (Nixon, 1944)

Nomenclature: 

Dacnusa tamsi Nixon, 1944

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Distribution: 
England

Chorebus tanis (Nixon, 1945)

Nomenclature: 

Dacnusa tanis Nixon, 1945

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Distribution: 
England

Chorebus tenellae Griffiths, 1967

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Distribution: 
Scotland, Ireland
Notes: 

added by Griffiths (1984)

Chorebus thecla (Nixon, 1943)

Nomenclature: 

Dacnusa thecla Nixon, 1943

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Distribution: 
England

Chorebus thisbe (Nixon, 1937)

Nomenclature: 

Dacnusa thisbe Nixon, 1937

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Distribution: 
England

Chorebus thusa (Nixon, 1937)

Nomenclature: 

Dacnusa thusa Nixon, 1937

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Distribution: 
England, Ireland

Chorebus transversus (Nixon, 1954)

Nomenclature: 

Dacnusa transversa Nixon, 1954

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Distribution: 
England, Scotland, Ireland

Chorebus trilobomyzae Griffiths, 1968

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Distribution: 
England, Scotland

Chorebus uliginosus (Haliday, 1839)

Nomenclature: 

Alysia uliginosa Haliday, 1839

thienemanni (Ruschka, 1913, Gyrocampa)

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Distribution: 
England, Ireland

Chorebus uma (Nixon, 1944)

Nomenclature: 

Dacnusa uma Nixon, 1944

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Distribution: 
England

Chorebus varuna (Nixon, 1945)

Nomenclature: 

Dacnusa varuna Nixon, 1945

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Distribution: 
England, Scotland

Chorebus vernalis Griffiths, 1968

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Distribution: 
England

Chorebus vitripennis Griffiths, 1968

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Distribution: 
Scotland, Wales, Ireland

Genus Coelinidea Viereck, 1913

Nomenclature: 

LEPTON Zetterstedt, 1838 preocc.

ERICOELINIUS Viereck, 1913

FISCHERASTRIOLUS Perepechayenko, 1999

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Notes: 

There is no recent treatment of this genus; generic synonymy follows Achterberg (2014).

Coelinidea elegans (Curtis, 1829)

Nomenclature: 

Chaenon elegans Curtis, 1829

brevicornis (Curtis, 1829, Chaenon)

cingulatus (Curtis, 1829, Chaenon)

rufinotatus (Curtis, 1829, Chaenon)

similis (Curtis, 1829, Chaenon)

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Distribution: 
England, Ireland

Coelinidea fuliginosus (Curtis, 1829)

Nomenclature: 

Chaenon fuliginosus Curtis, 1829

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Distribution: 
England

Coelinidea gracilis (Curtis, 1829)

Nomenclature: 

Chaenon gracilis Curtis, 1829

attenuator (Zetterstedt, 1838, Lepton)

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Distribution: 
England, Scotland, Ireland

Coelinidea niger (Nees, 1811)

Nomenclature: 

Stephanus niger Nees, 1811

affinis (Curtis, 1829, Chaenon)

nigricans (Westwood, 1835, Chaenon)

olivieri (Guérin-Ménéville, 1842, Alysia)

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Distribution: 
England, Ireland

Coelinidea obscurus (Curtis, 1829)

Nomenclature: 

Chaenon obscurus Curtis, 1829

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Distribution: 
England

Coelinidea podagricus (Haliday, 1839)

Nomenclature: 

Alysia podagrica Haliday, 1839

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Distribution: 
Ireland

Coelinidea ruficollis (Herrich-Schäffer, 1838)

Nomenclature: 

Coelinius ruficollis Herrich-Schäffer, 1838

procerus (Haliday, 1839, Alysia)

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Distribution: 
England

Coelinidea viduus (Curtis, 1829)

Nomenclature: 

Chaenon viduus Curtis, 1829

ater (Curtis, 1837, Chaenon) nom. nud.

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Distribution: 
England, Ireland

Genus Coelinius Nees, 1818

Nomenclature: 

CHAENON Curtis, 1829

COPISURA Schiødte,1837

COPIDURA Förster,1862

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Notes: 

Griffiths (1964) proposed that Polemochartus and Coelinidea should be included as subgenera of Coelinius. Wharton and Austin (1991), Wharton (1994) and Kula (2008) supported the concept of an enlarged Coelinius but, pointing to intermediate taxa in the Oriental Region, argued against the retention of subgenera. Achterberg (2014) retains the three genera which is current European usage and is followed here pending a modern review of the group.

Coelinius parvulus (Nees, 1811)

Nomenclature: 

Chaenon parvulus Nees, 1811

?cultriformis (Latreille, 1802, Ichneumon)

circulator (Gravenhorst, 1807, Ichneumon) preocc.

anceps (Curtis, 1829, Chaenon)

rimator (Schiødte, 1837, Copisura)

bicarinatus Herrich-Schäffer, 1838

flexuosus Herrich-Schäffer, 1838

bicolor Maréchal, 1938

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Distribution: 
England, Scotland, Ireland, Isle of Man

Genus Coloneura Förster, 1863

Nomenclature: 

ISOMERISTA Förster, 1863

TRISISA Förster, 1863

MERITES Nixon, 1943

PRIAPSIS Nixon, 1943

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Coloneura dice (Nixon, 1943)

Nomenclature: 

Priapsis dice Nixon, 1943

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Distribution: 
England

Coloneura stylata Förster, 1863

Nomenclature: 

exilis (Förster, 1863, Trisisa)

oligomera (Förster, 1863, Isomerista)

taras (Nixon, 1943, Merites)

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Distribution: 
England, Ireland

Genus Dacnusa Haliday, 1833

Nomenclature: 

AGONIA Förster, 1863

APHANTA Förster, 1863

BRACHYSTROPHA Förster, 1863

LIPOSCIA Förster, 1863

PACHYSEMA Förster, 1863

RHIZARCHA Förster, 1863

TANYSTROPHA Förster, 1863

RADIOLARIA Provancher, 1886

COLONEURELLA van Achterberg, 1976

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Dacnusa abdita Haliday, 1838

Nomenclature: 

incidens Thomson, 1895

lepida Marshall, 1896

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Distribution: 
England, Scotland, Wales, Ireland

Dacnusa adducta (Haliday, 1839)

Nomenclature: 

Alysia adducta Haliday, 1839

abducta misspelling

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Distribution: 
England, Scotland, Ireland
Notes: 

Based on its unusual venation, adducta is often placed in the monotypic genus Agonia; Griffiths’ (Griffiths 1964) argument that it is a derived species of Dacnusa is followed here.

Dacnusa alticeps Nixon, 1937

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Distribution: 
England, Scotland

Dacnusa aquilegiae Marshall, 1896

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Distribution: 
England, Scotland

Dacnusa areolaris (Nees, 1811)

Nomenclature: 

Bracon areolaris Nees, 1811

lysias Goureau, 1851

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Distribution: 
England, Scotland, Wales, Ireland

Dacnusa astarte (Nixon, 1948)

Nomenclature: 

Rhizarcha astarte Nixon, 1948

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Distribution: 
England