Male

Unknown.

Female (IZCAS-Ar 43530): PL 3.3, PW 2.4, AW 1.6, OL 3.1, OW 1.6. Eye diameters and interdistances: AME 0.10, ALE 0.13, PME 0.11, PLE 0.11, AME–AME 0.04, AME–ALE 0.11, PME–PME 0.06, PME–PLE 0.24, AME–PME 0.06, ALE–PLE 0.08, clypeus AME 0.12, clypeus ALE 0.17. Palp and leg measurements: palp 3.8 (1.3, 0.7, 0.8, -, 1.0), I 10.3 (2.8, 1.6, 2.8, 2.1, 1.0), II 9.2 (2.4, 1.4, 2.4, 2.0, 1.0), III 9.0 (2.4, 1.3, 2.0, 2.2, 1.1), IV 12.2 (3.1, 1.4, 2.9, 3.5, 1.3). Leg formula 4123. Spination of palp and legs: palp 130, 100, 1111, 1212; femora I p002, d111, r010, II p010, d111, r010, III p111, d111, r012, IV p002, d111, r102; patellae I–IV 001; tibiae I–II v22222, III p11, d111, r11, v222, IV p111, d11, r11, v222; metatarsi I–II v222, III p112, d010, r112, v222, IV p112, r112, v222. Chelicerae with 3 promarginal, 4 + 1 retromarginal teeth, without denticles. Retromargin of chelicerae close to fang base without bristle. Tarsi and metatarsi without scopula. Claw tufts arising separately, but intermingle with each other distally. Palpal claw with 3 secondary teeth, leg claws I–II with 3, III with 2 and IV with 3 secondary teeth. Position of tarsal organ: I 0.76, II 0.72, III 0.68.

Copulatory organ (Fig. 2a, b, Fig. 3a and b). Epigynal plate width/length: 9.8/6.5; anterior width/posterior width: 9.8/7.5; heart-shaped and with a mating plug, the anterior part with a pair of pointed apophyses ventrally. Lateral teeth pointing postero-medially. Internal duct system with small oval spermathecae not fully visible, separated from each other by more than their diameter; fertilisation ducts elongate and laminar, pointing postero-medially.

Figure 1.  

Distribution records of ctenid spiders from Asia in this study. 1. Amauropelma krabi sp. n.; 2. Am. phangnga sp. n.; 3. Am. saraburi sp. n.; 4. Anahita medog sp. n.; 5. An. popa; 6. Bowie fascination; 7. B. ninhbinh sp. n.; 8. B. vinhphuc sp. n.; 9. B. borneo sp. n.; 10. B. engkilili sp. n.; 11. B. sabah sp. n.

Figure 2.  

Amauropelma krabi sp. n., holotype female. a: Epigyne, ventral view, arrow points at pointed apophysis; b: Vulva, dorsal view; c: Habitus, dorsal view; d: Habitus, ventral view. FD = fertilisation duct, IF = internal fold, LT = lateral teeth, SP = spermathecae. Scale bars: 0.2 mm (a, b), 1.0 mm (c, d).

Figure 3.  

Amauropelma krabi sp. n., holotype female. a: Epigyne, ventral view, arrow points at pointed apophysis; b: Vulva, dorsal view. FD = fertilisation duct, IF = internal fold, LT = lateral teeth, SP = spermathecae. Scale bars: 0.2 mm (a, b).

Colour (Fig. 2c and d). Reddish-brown to yellowish without patterns. Dorsal prosoma slightly reddish-brown to yellowish, with eyes marked with black rings, fovea distinct, reddish-brown. Chelicerae reddish-brown. Sternum, ventral coxae, labium yellowish-brown without patterns. Gnathocoxae yellowish-brown with lighter distal lips. Legs yellowish-brown. Opisthosoma yellowish. Spinnerets yellowish.

Small Ctenidae (total length female 6.4). The new species can be distinguished from all known congeners by the median plate roughly heart-shaped and with a mating plug (Fig. 2a and Fig. 3a), by the anterior part of median plate with a pair of pointed apophyses ventrally (arrowed in Fig. 2a, arrowed in Fig. 3a), by the internal duct system with small oval spermathecae not fully visible (Fig. 2b and Fig. 3b) and by the fertilisation ducts which are elongate and laminar, almost twice as long as the spermathecae (Fig. 2b and Fig. 3b).

The specific name refers to the type locality and is a noun in apposition.

Thailand (Krabi, type locality; Fig. 1).

Female (IZCAS-Ar 43530):

TGTTTGGAGCTTGAGCTGCTATAGCAGGAACTGGAATAAGAGTGTTGATTCGAATAGAGTTAGGTCATCCTGGTAGATTGTTAGGAGATGATCATTTATATAATGTTATTGTAACTGCTCATGCTTTTGTAATGATTTTTTTTATAGTAATACCAATTTTGATTGGTGGATTTGGAAATTGATTAGTTCCGTTGAGATTGGAGCACCTGATATATCATTTCCTCGAATAAATAATTTGTCGTTTTGATTACTACCTCCTTCTTTATTTTTATTAATAATATCATCAATAGTAGAAATAGGTGTTGGAGCGGGATGAACTGTTTATCCTCCTTTAGCATCTAGTATTGGGCATATAGGAAGATCTATAGATTTTGCTATTTTTTCTCTTCATTTGGCTGGAGCTTCTTCTATTATAGGAGCAGTAAATTTTATTTCTACTATTATTAATATACGGTTGTATGGAATGAGTATAGAAAAGGTTCCTTTGTTTGTGTGGTCTGTTTTTATTACTGCTATTTTGTTATTATTGTCGTTACCTGTGTTAGCAGGTGCTATTACTATATTATTGACTGATCGAAATTTTAATACTTCTTTTTTTGACCCTGCGGGAGGGGGAGATCCTATTTTGTTTCAACATTTATTTTGATTTTTTG (GenBank accession number OP561682).

Male (IZCAS-Ar 43531): PL 3.3, PW 2.8, AW 1.2, OL 3.0, OW 1.9. Eye diameters and interdistances: AME 0.09, ALE 0.12, PME 0.10, PLE 0.10, AME–AME 0.04, AME–ALE 0.13, PME–PME 0.08, PME–PLE 0.21, AME–PME 0.05, ALE–PLE 0.08, clypeus AME 0.17, clypeus ALE 0.22. Palp and leg measurements: palp 3.8 (0.9, 0.6, 0.8, -, 1.5), I 13.5 (3.3, 1.6, 3.7, 3.2, 1.7), II 11.6 (3.1, 1.6, 3.0, 2.6, 1.3), III 10.9 (2.9, 1.4, 2.7, 2.6, 1.3), IV 14.8 (3.5, 1.5, 3.9, 4.2, 1.7). Leg formula 4123. Spination of palp and legs: palp 131, 100, 1101; femora I p021, d211, r112, II–III p012, d111, r012, IV p102, d111, r012; patellae I–IV 001; tibiae I p010, v22222, II p100, r100, v22222, III p11, d111, r11, v222, IV p11, d11, r11, v222; metatarsi I v222, II p112, r010, v222, III p112, d010, r112, v222, IV p112, r112, v2222. Chelicerae with 3 promarginal, 4 retromarginal teeth, without denticles. Retromargin of chelicerae close to fang base without bristle. Tarsi and metatarsi without scopula. Claw tufts arising separately, but intermingle with each other distally. Leg claws I with 7 and II with 6 secondary teeth. Position of tarsal organ: I 1.37, II 0.92, III 0.85.

Palp (Fig. 4a–c). Patella with distinct retrolateral apophysis. RTA protruding at an almost right angle from tibia in ventral view, with broad base and two short apices, both dorso-distad. Cymbium tip conical, with prolatero-proximal outgrowth. Embolus (Fig. 9a) arising at 8 o’clock position, its tip with an extension. Conductor arising at 3 o’clock position, long and laminar, running around tegulum anti-clockwise, its tip situated subdistally. Tegular apophysis arising subcentrally, strongly concave on ventral side and distinctly excavated on prolateral side.

Figure 4.  

Amauropelma phangnga sp. n., palp, holotype. a: Prolateral view; b: Ventral view; c: Retrolateral view. C = conductor, E = embolus, PCO = prolatero-proximal cymbial outgrowth, RPA = retrolateral patellar apophysis, RTA = retrolateral tibial apophysis, TA = tegular apophysis. Scale bar: 0.5 mm (a–c).

Figure 5.  

Amauropelma phangnga sp. n., habitus, holotype male. a: Dorsal view; b: Ventral view. Scale bar: 1.0 mm (a, b).

Figure 6.  

Amauropelma saraburi sp. n., flipped right palp, holotype. a: Prolateral view; b: Ventral view; c: Retrolateral view. C = conductor, E = embolus, PCO = prolatero-proximal cymbial outgrowth, RPA = retrolateral patellar apophysis, RPO = retro-proximal cymbial outgrowth, TA = tegular apophysis. Scale bar: 0.5 mm (a–c).

Figure 7.  

Amauropelma saraburi sp. n. a: Paratype female, epigyne, ventral view; b: Same, vulva, dorsal view; c: Holotype male, habitus, dorsal view; d: Same, habitus, ventral view; e: Paratype female, habitus, dorsal view; f: Same, habitus, ventral view. FD = fertilisation duct, IF = internal fold, LT = lateral teeth, SP = spermathecae. Scale bars: 0.2 mm (a, b), 1.0 mm (c–f).

Figure 8.  

Amauropelma saraburi sp. n., paratype female. a: Epigyne, ventral view; b: Vulva, dorsal view. FD = fertilisation duct, IF = internal fold, LT = lateral teeth, SP = spermathecae. Scale bars: 0.2 mm (a, b).

a

b

Figure 9.

Amauropelma spp., embolic tips, ventral view.

a: Am. phangnga sp. n., paratype. Scale bar: 0.05 mm.   
b: Am. saraburi sp. n., holotype. Scale bar: 0.02 mm.  

Colour (Fig. 5a and b). Yellowish-brown. Dorsal prosoma with eyes marked with black rings, fovea distinct, reddish-brown. Ventral opisthosoma grey.

Female

Unknown.

Variation: Paratype male (IZCAS-Ar 43532): PL 3.4, OL 2.4.

Small Ctenidae (total length male 5.8–6.3). The new species can be distinguished from all known congeners by the embolus tip with an extension (Fig. 4b and Fig. 9a), by the tegular apophysis strongly concave on ventral side and distinctly excavated on prolateral side (Fig. 4a and b), by the conductor arising at 3 o’clock position, long and laminar, running around tegulum anti-clockwise, its tip situated subdistally (Fig. 4b), by the RTA protruding at an almost right angle from tibia in ventral view, with broad base and two short apices, both dorso-distad (Fig. 4b) and by the patella with distinct retrolateral apophysis, pointing anteriorly (Fig. 4b). This species can also be distinguished from Am. krabi sp. n. by the COI p-distance 0.134 between them.

The specific name refers to the type locality and is a noun in apposition.

Thailand (Phang Nga, type locality; Fig. 1).

Male (IZCAS-Ar 43532):

GGTGGGTTCGGAAATTGATTGGTTCCTTTGATGTTAGGAGCTCCTGATATATCATTTCCTCGTATAAATAATTTGTCTTTTTGGTTACTTCCTCCTTCTTTATTTTTGTTATTAATATCTTCTATGGTGGAAATAGGAGTGGGAGCAGGATGAACTGTCTATCCTCCTTTAGCTTCTAGAATAGGGCATGTGGGAAGATCAATAGATTTTGCGATTTTTTCTCTTCATTTAGCTGGAGTTTCTTCTATTATGGGAGCGGTTAATTTTATTTCTACTATTATTAATATGCGATTATATGGAATAACTATAGAAAAGGTTCCTTTATTCGTTTGATCAGTTTTTATTACTGCAGTTTTGTTGTTGTTATCATTACCTGTGTTAGCAGGTGCTATTACTATATTATTGACAGATCGAAATTTTAATACTTCTTTTTTTGATCCTGCAGGGGGTGGAGATCCAATTTTATTTCAACATTTATTCTGATTTTTTGGTCACCCTGGAAAGTTTAA (GenBank accession number OP718556).

Male (IZCAS-Ar 43533): PL 4.5, PW 3.7, AW 1.6, OL 3.2, OW 2.1. Eye diameters and interdistances: AME 0.12, ALE 0.14, PME 0.14, PLE 0.14, AME–AME 0.07, AME–ALE 0.16, PME–PME 0.08, PME–PLE 0.26, AME–PME 0.07, ALE–PLE 0.13, clypeus AME 0.16, clypeus ALE 0.28. Palp and leg measurements: palp 5.2 (1.8, 0.8, 0.9, -, 1.7), I - (4.5, 2.0, 4.5, 4.0, -), II 15.2 (3.8, 2.1, 3.8, 3.7, 1.8), III 14.3 (3.6, 1.8, 3.4, 3.7, 1.8), IV 19.4 (4.9, 2.0, 4.6, 5.6, 2.3). Leg formula 4123. Spination of palp and legs: palp 131, 100, 210; femora I p112, d111, r111, II p211, d111, r211, III p112, d111, r112, IV p112, d111, r002; patellae I 001, II–IV 101; tibiae I p010, r110, v22222, II p100, r100, v22222, III–IV p11, d111, r11, v222; metatarsi I v222, II p110, r110, v222, III p112, d010, r112, v222, IV p112, d010, r112, v2222. Chelicerae with 3 promarginal, 4 retromarginal teeth, without denticles. Retromargin of chelicerae close to fang base without bristle. Claw tufts arising separately, but intermingle with each other distally. Leg claws II with 1 and III–IV with 2 secondary teeth. Position of tarsal organ: IV 1.58.

Palp (Fig. 6a–c). Patella with distinct retrolateral apophysis. Cymbium tip conical, with prolatero-proximal outgrowth and retro-proximal outgrowth. Embolus (Fig. 9b) slender, arising at 8.30 o’clock position. Conductor arising at 1 o’clock position. Tegular apophysis large and longitudinally elongated, with excavation on prolateral side.

Colour (Fig. 7c and d). Yellowish-brown. Dorsal prosoma yellowish with eyes marked with black rings, fovea distinct, brown. Sternum, ventral coxae, labium and gnathocoxae yellowish without patterns. Chelicerae brown. Legs yellowish. Dorsal opisthosoma yellowish without patterns. Lateral and ventral opisthosoma grey without patterns. Spinnerets grey.

Female (IZCAS-Ar 43534): PL 5.6, PW 4.4, AW 2.8, OL 4.9, OW 3.1. Eye diameters and interdistances: AME 0.14, ALE 0.20, PME 0.15, PLE 0.16, AME–AME 0.12, AME–ALE 0.32, PME–PME 0.16, PME–PLE 0.53, AME–PME 0.08, ALE–PLE 0.20, clypeus AME 0.13, clypeus ALE 0.21. Palp and leg measurements: palp 6.0 (1.7, 1.2, 1.4, -, 1.7), I 19.3 (4.6, 2.7, 5.6, 4.2, 2.2), II 18.3 (4.9, 2.5, 5.0, 4.0, 1.9), III 17.0 (4.6, 2.2, 4.1, 4.3, 1.8), IV 23.0 (5.8, 2.4, 5.5, 6.6, 2.7). Leg formula 4123. Spination of palp and legs: palp 131, 100, 1111, 2112; femora I p021, d111, r111, II–III p112, d111, r112, IV p112, d111, r002; patellae I–II 000, III–IV 101; tibiae I –II v22222, III–IV p11, d111, r11, v222; metatarsi I–II v222, III–IV p112, d010, r112, v222. Chelicerae with 3 promarginal, 4 retromarginal teeth, without denticles. Retromargin of chelicerae close to fang base without bristle. Sparse scopula restricted almost entirely to tarsi, only metatarsi I–II with sparse scopula hairs. Claw tufts arising separately, but intermingle with each other distally. Palpal claw with 3 secondary teeth, leg claws I with 3, II–III with 2 secondary teeth.

Copulatory organ (Fig. 7a, b, Fig. 8a and b). Epigynal plate width/length: 19.5/8.6; anterior width/posterior width: 15/6.3; with lateral wings and posterior part with distinct lateral margins and two distinct tubercles. Lateral teeth pointing anteriorly. Internal duct system with round spermathecae not fully visible and spermathecae separated from each other by more than their diameter; fertilisation ducts elongate and laminar, pointing postero-medially.

Colour (Fig. 7e and f). Reddish-brown to yellowish. Dorsal prosoma slightly reddish-brown with eyes marked with black rings, fovea distinct, reddish-brown. Sternum and ventral coxae yellowish without patterns; labium and gnathocoxae reddish-brown without patterns. Chelicerae reddish-brown. Legs yellowish-brown. Dorsal and lateral opisthosoma grey without patterns. Ventral opisthosoma yellowish without patterns. Spinnerets yellowish.

Variation: Second paratype female (IZCAS-Ar 43535): PL 4.6, OL 4.7.

Small to medium-sized Ctenidae (total length male 7.7, female 9.3–10.5). The new species can be distinguished from all known congeners by the embolus slender (Fig. 6b and Fig. 9b), by the tegular apophysis large and longitudinally elongated, with excavation on prolateral side (Fig. 6a–c), by the epigynal field wider than long, with lateral wings and distinct lateral margins (Fig. 7a and Fig. 8a) and by the lateral teeth pointing anteriorly (Fig. 7a and Fig. 8a).

The specific name refers to the type locality and is a noun in apposition.

Thailand (Saraburi, type locality; Fig. 1).

Male (IZCAS-Ar 43533):

TGTTTGGAGCTAGATCTGCTATAGCGGGAACGGCAATAAGAGTTTTAATTCGTATGGAATTAGGAAATTCTGGAAGATTATTAGGGGATGATCATTTATATAATGTAATTGTGACAGCTCATGCTTTTATTATGATTTTTTTTATAGTAATACCGATTTTGATTGGTGGTTTTGGAAATTGATTAGTGCCTTTAATGTTAGGAGCTCCTGATATATCTTTTCCTCGGATGAATAATTTGTCTTTTTGATTACTTCCACCTTCTTTGTTTTTATTATTCATATCTTCTATGGTGGAAATGGGTGTAGGAGCTGGATGAACTGTTTATCCACCTTTGGCTTCTAGAATTGGTCATGCTGGAAGATCTATGGATTTTGCTATTTTTTCTTTACATTTAGCTGGGGCTTCTTCAATTATAGGAGCGGTGAATTTTATTTCTACTATTATTAATATACGATTATCTGGAATAAGAATGGAGAAGGTTCCATTATTTGTTTGATCTGTTCTTATTACTGCAATTTTATTATTATTATCTTTGCCGGTATTAGCTGGTGCTATTACTATATTGTTGACTGATCGAAATTTTAATACTTCTTTTTTTGATCCGGCTGGGGGAGGGGATCCTATTTTATTTCAACATTTATTTTGATTTTTTG (GenBank accession number OP572100).

Female (IZCAS-Ar 43534):

TGTTTGGAGCTTGATCTGCTATAGCGGGAACGGCAATAAGAGTTTTAATTCGTATGGAATTAGGAAATTCTGGAAGATTATTAGGGGATGATCATTTATATAATGTAATTGTGACAGCTCATGCTTTTATTATGATTTTTTTTATAGTAATACCGATTTTGATTGGTGGTTTTGGAAATTGATTAGTGCCTTTAATGTTAGGAGCTCCTGATATATCTTTTCCTCGGATGAATAATTTGTCTTTTTGATTACTTCCACCTTCTTTGTTTTTATTATTCATATCTTCTATGGTGGAAATGGGTGTAGGAGCTGGATGAACTGTTTATCCACCTTTGGCTTCTAGAATTGGTCATGCTGGAAGATCTATGGATTTTGCTATTTTTTCTTTACATTTAGCTGGGGCTTCTTCAATTATAGGAGCGGTGAATTTTATTTCTACTATTATTAATATACGATTATCTGGAATAAGAATGGAGAAGGTTCCATTATTTGTTTGATCTGTTCTTATTACTGCAATTTTATTATTATTATCTTTGCCGGTATTAGCTGGTGCTATTACTATATTGTTGACTGATCGAAATTTTAATACTTCTTTTTTTGATCCGGCTGGGGGAGGGGATCCTATTTTATTTCAACATTTATTTTGATTTTTTG (GenBank accession number OP572099).

Male (IZCAS-Ar 43536): PL 2.7, PW 2.3, AW 0.9, OL 2.4, OW 1.4. Eye diameters and interdistances: AME 0.13, ALE 0.10, PME 0.22, PLE 0.19, AME–AME 0.10, AME–ALE 0.22, PME–PME 0.16, PME–PLE 0.23, AME–PME 0.11, ALE–PLE 0.14, clypeus AME 0.10, clypeus ALE 0.37. Palp and leg measurements: palp 4.1 (1.5, 0.6, 0.8, -, 1.2), I missing, II 13.0 (3.6, 1.2, 3.7, 3.3, 1.2), III 10.9 (2.8, 1.1, 2.8, 3.0, 1.2), IV 16.0 (4.1, 1.1, 4.2, 5.1, 1.5). Leg formula 4123. Spination of palp and legs: palp 023, 000, 0211; femora II p112, d111, r012, III p112, d111, r112, IV p112, d111, r012; patellae II–IV 101; tibiae II p110, d101, r100, v22222, III p11, d111, r11, v222, IV p11, d111, r11, v22; metatarsi II p111, r111, v222, III p112, d010, r112, v222, IV p112, d010, r112, v2222. Chelicerae with 3 promarginal, 4 + 1 retromarginal teeth and with elongated patch of 6 tiny denticles along entire cheliceral furrow. Leg claws II with 9, III with 5 and IV with 7 secondary teeth. Position of tarsal organ: II 1.06, III 0.85, IV 1.14.

Palp (Fig. 10a–c). Palpal tibia without RTA and intrasegmental sclerite, distally with retrolateral stout spine. Cymbium tip conical. Embolus (Fig. 14a) arising at 6.30 o’clock position, with wide base and narrow tip and a membranous apophysis apically. Conductor absent. Tegular apophysis arising nearly centrally from tegulum.

Figure 10.  

Anahita medog sp. n., palp, holotype. a: Prolateral view; b: Ventral view; c: Retrolateral view. DS = distal retrolateral spine, E = embolus, TA = tegular apophysis. Scale bar: 0.2 mm (a–c).

Figure 11.  

Anahita medog sp. n. a: Paratype female, epigyne, ventral view; b: Same, vulva, dorsal view; c: Holotype male, habitus, dorsal view; d: Same, habitus, ventral view; e: Paratype female, habitus, dorsal view; f: Same, habitus, ventral view. CD = copulatory duct, CO = copulatory opening, FD = fertilisation duct, LT = lateral teeth, SP = spermathecae. Scale bars: 0.2 mm (a, b), 1.0 mm (c–f).

Figure 12.  

Anahita popa Jäger & Minn, 2015, palp, male from Sagaing Hill. a: Prolateral view; b: Ventral view; c: Retrolateral view. DS = distal retrolateral spine, E = embolus, TA = tegular apophysis. Scale bar: 0.2 mm (a–c).

Figure 13.  

Anahita popa Jäger & Minn, 2015, male and female from Sagaing Hill. a: Female, epigyne, ventral view; b: Same, vulva, dorsal view; c: Male, habitus, dorsal view; d: Same, habitus, ventral view; e: Female, habitus, dorsal view; f: Same, habitus, ventral view. FD = fertilisation duct, LT = lateral teeth, SP = spermathecae, SS = slit sensillum. Scale bars: 0.2 mm (a, b), 1.0 mm (c–f).

a

b

Figure 14.

Anahita spp., embolic tips, ventral view.

a: An. medog sp. n., holotype. Scale bar: 0.05 mm.  
b: An. popa Jäger & Minn, 2015, male from Sagaing Hill. Scale bar: 0.01 mm.   

Colour (Fig. 11c and d). Black to yellowish. Dorsal prosoma with two parallel black lateral bands and distinctly marked fovea. Sternum, ventral coxae, labium and gnathocoxae yellowish. Chelicerae yellowish with longitudinal lines. Palps and legs yellowish. Dorsal opisthosoma black with light median band. Lateral opisthosoma spotted. Ventral opisthosoma yellowish with lateral black patterns. Spinnerets dark.

Female (IZCAS-Ar 43537): PL 2.9, PW 2.4, AW 1.2, OL 3.5, OW 2.0. Eye diameters and interdistances: AME 0.13, ALE 0.11, PME 0.19, PLE 0.18, AME–AME 0.13, AME–ALE 0.26, PME–PME 0.21, PME–PLE 0.24, AME–PME 0.15, ALE–PLE 0.17, clypeus AME 0.10, clypeus ALE 0.38. Palp and leg measurements: palp 3.2 (0.9, 0.6, 0.8, -, 0.9), I 10.4 (2.9, 1.3, 3.1, 2.2, 0.9), II 9.3 (2.7, 1.2, 2.5, 2.0, 0.9), III 8.3 (2.3, 1.1, 2.1, 2.0, 0.8), IV 12.4 (3.3, 1.2, 3.1, 3.6, 1.2). Leg formula 4123. Spination of palp and legs: palp 020, 010, 010, 2012; femora I p011, d111, r021, II p011, d111, r011, III p012, d111, r112, IV p002, d111, r012; patellae I–II 000, III–IV 101; tibiae I v22212, II v22222, III–IV p11, d111, r11, v222; metatarsi I–II v222, III p112, d010, r112, v222, IV p112, r112, v2222. Chelicerae with 3 promarginal, 4 retromarginal teeth and with elongated patch of 5 tiny denticles along entire cheliceral furrow. Palpal claw with 5 secondary teeth, leg claws I–II with 5, III with 4 and IV with 7 secondary teeth. Position of tarsal organ: I 0.79, II 0.72, III 0.70, IV 1.00.

Copulatory organ (Fig. 11a and b). Lateral teeth arising posteriorly from median plate. Median plate with a large n-shaped sclerite. Copulatory openings hidden under median plate. Copulatory ducts nearly triangular. Spermathecae nearly cylindrical. Fertilisation ducts pointing anteriorly.

Colour (Fig. 11e and f). As in male.

Small Ctenidae (total length male 5.1, female 6.4). The species resembles A. maolan Zhu, Chen & Song, 1999 (see Zhu et al. 1999: figs 1–5; Yin et al. 2012: fig. 473a–e; Marusik and Omelko 2016: figs 1–13 and 32–34) by having similar distal retrolateral spine, tegular apophysis (Fig. 10a–c) and fertilisation ducts (Fig. 11b), but can be distinguished by the embolus arising at 6.30 o’clock position, its tip with membranous apophysis (Fig. 10b and Fig. 14a; embolus arising centrally from tegulum, its tip without membranous apophysis in A. maolan), by the median plate with a large n-shaped sclerite (Fig. 11a; absent in A. maolan), by the lateral teeth pointing postero-medially (Fig. 11a; lateral teeth pointing medially in A. maolan) and by the spermathecae nearly cylindrical (Fig. 11b; spermathecae nearly wavy in A. maolan).

The specific name refers to the type locality and is a noun in apposition.

China (Tibet, type locality; Fig. 1).

Male (IZCAS-Ar 43536):.

TGTTTGGAGCTTGAGCTGCTATAGCTGGAACAGCAATAAGAGTTTTAATTCGAATGGAATTAGGACATTCTGGTAGATTGTTAGGAGATGATCATTTATATAATGTAATTGTAACGGCTCATGCTTTTGTTATAATTTTTTTTATAGTAATACCTATTTTGATTGGGGGCTTTGGTAATTGGTTGGTTCCTTTAATGTTAGGGGCTCCGGATATATCTTTTCCTCGAATAAATAATTTATCCTTTTGATTATTACCGCCTTCTTTATTTTTGTTGTTTATATCTTCTATAGTTGAGATAGGGGTTGGAGCAGGTTGAACGGTTTATCCTCCTTTAGCTTCTAGAATTGGGCATATGGGAAGTTCAATGGATTTTGCTATTTTTTCTTTACATTTAGCAGGTGCTTCTTCTATTATAGGTGCTGTGAATTTTATTTCTACTATTATTAATATACGATTAATAGGAATAACAATGGAGAAGATCCCTTTATTTGTATGATCGGTTTTTATTACTGCAATTTTATTATTATTATCTTTACCTGTTTTAGCAGGAGCTATTACTATATTATTGACTGATCGAAATTTTAATACTTCTTTTTTTGACCCTGCTGGAGGTGGAGATCCTATTTTATTTCAACATTTATTTTGATTTTTTG (GenBank accession number OP572101).

Female (IZCAS-Ar 43537):

TGTTTGGAGCTTGAGCTGCTATAGCTGGAACAGCAATAAGAGTTTTAATTCGAATGGAATTAGGACATTCTGGTAGATTGTTAGGAGATGATCATTTATATAATGTAATTGTAACGGCTCATGCTTTTGTTATAATTTTTTTTATAGTAATACCTATTTTGATTGGGGGCTTTGGTAATTGGTTGGTTCCTTTAATGTTAGGGGCTCCGGATATATCTTTTCCTCGAATAAATAATTTATCCTTTTGATTATTACCGCCTTCTTTATTTTTGTTGTTTATATCTTCTATAGTTGAGATAGGGGTTGGAGCAGGTTGAACGGTTTATCCTCCTTTAGCTTCTAGAATTGGGCATATGGGAAGTTCAATGGATTTTGCTATTTTTTCTTTACATTTAGCAGGTGCTTCTTCTATTATAGGTGCTGTGAATTTTATTTCTACTATTATTAATATACGATTAATAGGAATAACAATGGAGAAGATCCCTTTATTTGTATGATCGGTTTTTATTACTGCAATTTTATTATTATTATCTTTACCTGTTTTAGCAGGAGCTATTACTATATTATTGACTGATCGAAATTTTAATACTTCTTTTTTTGACCCTGCTGGAGGTGGAGATCCTATTTTATTTCAACATTTATTTTGATTTTTTG (GenBank accession number OP572102).

Male (IZCAS-Ar 43538): PL 3.3, PW 2.7, AW 0.9, OL 3.3, OW 1.8. Eye diameters and interdistances: AME 0.14, ALE 0.12, PME 0.22, PLE 0.17, AME–AME 0.08, AME–ALE 0.23, PME–PME 0.16, PME–PLE 0.21, AME–PME 0.18, ALE–PLE 0.13, clypeus AME 0.10, clypeus ALE 0.55. Palp and leg measurements: palp 3.9 (1.4, 0.6, 0.8, -, 1.1), I 15.4 (3.7, 1.6, 4.3, 3.8, 2.0), II 12.7 (3.4, 1.4, 3.3, 3.1, 1.5), III 11.0 (2.9, 1.2, 2.6, 3.1, 1.2), IV 17.3 (4.3, 1.6, 4.4, 5.2, 1.8). Leg formula 4123. Spination of palp and legs: palp 151, 000, 122; femora I p021, d111, r112, II p112, d111, r112, III p111, d111, r111, IV p012, d111, r112; patellae I–IV 101; tibiae I p010, d101, r110, v22222, II p10, d101, r110, v22222, III–IV p11, d111, r11, v222; metatarsi I p111, d001, r111, v222, II p111, d111, r111, v222, III p111, d012, r111, v222, IV p112, d011, r112, v222. Chelicerae with 3 promarginal, 4 + 2 retromarginal teeth and with elongated patch of 3 tiny denticles along entire cheliceral furrow. Retromargin of chelicerae close to fang base with 2 bristles. Sparse scopula restricted almost entirely to tarsi. Leg claws I with 6, II–III with 5 and IV with 4secondary teeth. Position of tarsal organ: I 1.89, II 1.03, III 0.75, IV 1.56.

Palp (Fig. 12a–c). Palpal tibia without RTA and intrasegmental sclerite, distally with 2 stout retrolateral spines. Cymbium tip slightly conical. Embolus (Fig. 14b) arising at 12 o’clock position, long and laminar, running around tegulum, its tip situated distally. Conductor absent. Tegular apophysis arising at 12 to 12.30 o’clock position subdistally.

Colour (Fig. 13c and d). Yellowish-brown with light brown markings. Dorsal prosoma with distinct light median band and dark lateral bands, light patches partly fused, frontally with 2 light patches close to ALE. Sternum, coxae, labium and gnathocoxae pale yellowish without patterns. Chelicerae yellowish-brown with longitudinal dark lines frontally. Palps and legs yellowish-brown, legs I–III with pattern especially from femora to tibiae. Dorsal opisthosoma yellowish-brown with distinct serrated light median band. Lateral opisthosoma spotted. Ventral opisthosoma yellowish, with posteriorly converging lines of spots. Spinnerets light, anterior lateral spinnerets laterally dark, anal tubercle light.

Female (IZCAS-Ar 43539): See Fig. 13a, b, e and f; figs 1–6 in Jäger and Minn (2015).

Small Ctenidae (total length male 6.6). The species can be distinguished from all known congeners by the embolus arising at 12 o’clock position, long and laminar, running around tegulum, its tip situated distally (Fig. 12b and Fig. 14b), by the palp having no conductor (Fig. 12a–c), by the tegular apophysis arising at 12 to 12.30 o’clock position subdistally (Fig. 12b) and by the tibia distally with 2 stout retrolateral spines (Fig. 12b and c). For the diagnosis of female, see Jäger and Minn (2015).

Myanmar (Sagaing Hill, Fig. 1; Mt Popa, type locality).

Male (IZCAS-Ar 43538):

TATTTGGGGCTTGAGCTGCTATAGCGGGTACTGCAATAAGAGTTTTGATTCGAATGGAATTAGGACATCCTGGAAGATTATTAGGTGATGATCATTTATATAATGTTATTGTAACAGCTCATGCTTTTGTTATGATTTTTTTTATAGTTATACCTATTTTAATTGGTGGTTTTGGAAATTGGTTAGTTCCTTTAATATTAGGAGCTCCGGATATATCATTTCCTCGAATAAATAATTTATCTTTTTGGTTATTACCTCCTTCTTTGTTTTTATTGTTTATATCTTCTATAGTTGAAATAGGTGTAGGAGCAGGGTGAACAGTTTATCCTCCTTTAGCTTCTAGAATTGGGCATGCAGGGAGATCTATGGATTTTGCTATTTTTTCTTTACATTTAGCGGGTGCTTCTTCTATTATAGGGGCTGTAAATTTTATTTCTACTATTATTAATATACGATTAATAGGAATGACTATAGAGAAGGTTCCTTTGTTTGTTTGATCTGTTTTTATTACTGCAATTTTATTATTGTTATCTTTACCAGTGTTAGCTGGTGCTATTACAATATTATTAACTGATCGTAATTTTAATACTTCTTTTTTTGATCCTGCTGGAGGAGGAGATCCAGTTTTATTTCAGCATTTGTTTTGATTTTTTG (GenBank accession number OP572105).

Female (IZCAS-Ar 43539):

TTTTTGGAGCTTGAGCCGCTATAGCGGGTACTGCAATAAGAGTTTTAATTCGAATAGAATTAGGGCATCCTGGGAGATTATTAGGTGATGATCATTTATATAATGTTATTGTAACAGCTCATGCTTTTGTTATAATTTTTTTTATAGTTATACCTATTTTAATTGGTGGTTTTGGAAATTGGTTAGTTCCTTTAATGTTAGGAGCTCCGGATATATCATTTCCTCGAATAAATAATTTATCTTTTTGATTATTACCTCCTTCTTTGTTTTTATTGTTTATATCTTCCATGGTTGAAATAGGTGTGGGAGCAGGATGGACAGTTTATCCTCCTTTAGCTTCTAGAATTGGGCATGCGGGAAGATCTATGGATTTTGCTATTTTTTCTTTACATTTAGCGGGTGCTTCTTCTATTATAGGAGCTGTAAATTTTATTTCGACTATTATTAATATACGATTAATAGGAATGACTATAGAGAAGGTTCCCTTATTTGTTTGATCTGTTTTTATTACTGCAATTTTATTGTTATTATCTTTACCAGTATTAGCTGGTGCTATTACGATGTTGTTAACTGATCGTAATTTTAATACTTCTTTTTTTGACCCTGCTGGGGGAGGGGATCCGGTTTTATTTCAACATTTATTTTGATTTTTTG (GenBank accession number OP572104).

Male (IZCAS-Ar 43540): See Fig. 15a–c, Fig. 16c, d and Fig. 28a; figs 230–233 and 263–264 in Jäger (2022).

Figure 15.  

Bowie fascination Jäger, 2022, palp, male from Yunnan. a: Prolateral view; b: Ventral view; c: Retrolateral view. C = conductor, E = embolus, RPO = retro-proximal cymbial outgrowth, RTA = retrolateral tibial apophysis, TA = tegular apophysis. Scale bar: 0.5 mm (a–c).

Figure 16.  

Bowie fascination Jäger, 2022, male and female from Yunnan. a: Female, epigyne, ventral view; b: Same, vulva, dorsal view; c: Male, habitus, dorsal view; d: Same, habitus, ventral view; e: Female, habitus, dorsal view; f: Same, habitus, ventral view. AB = anterior band of epigynal field, FD = fertilisation duct, LT = lateral teeth, SP = spermathecae. Scale bars: 0.5 mm (a, b), 5.0 mm (c–f).

Figure 17.  

Bowie ninhbinh sp. n., palp, holotype. a: Prolateral view; b: Ventral view; c: Retrolateral view. C = conductor, E = embolus, RPO = retro-proximal cymbial outgrowth, RTA = retrolateral tibial apophysis, TA = tegular apophysis. Scale bar: 0.5 mm (a–c).

Figure 18.  

Bowie ninhbinh sp. n., habitus, holotype male. a: Dorsal view; b: Ventral view. Scale bar: 2.0 mm (a, b).

Figure 19.  

Bowie vinhphuc sp. n., palp, holotype. a: Prolateral view; b: Ventral view; c: Retrolateral view. C = conductor, E = embolus, RPO = retro-proximal cymbial outgrowth, RTA = retrolateral tibial apophysis, TA = tegular apophysis. Scale bar: 0.5 mm (a–c).

Figure 20.  

Bowie vinhphuc sp. n. a: Paratype female, epigyne, ventral view; b: Same, vulva, dorsal view; c: Holotype male, habitus, dorsal view; d: Same, habitus, ventral view; e: Paratype female, habitus, dorsal view; f: Same, habitus, ventral view. FD = fertilisation duct, HA = humped areas of median plate, LT = lateral teeth, SP = spermathecae. Scale bars: 0.2 mm (a, b), 2.0 mm (c–f).

Figure 21.  

Epigyne, posterior view. a: Bowie fascination Jäger, 2022; b: B. vinhphuc sp. n. HA = humped areas of median plate, LT = lateral teeth. Scale bars: 0.2 mm (a), 0.1 mm (b).

Figure 22.  

Bowie borneo sp. n., palp, holotype. a: Prolateral view; b: Ventral view; c: Retrolateral view. C = conductor, DPO = dorso-proximal outgrowth of cymbium, E = embolus, RTA = retrolateral tibial apophysis, TA = tegular apophysis. Scale bar: 0.2 mm (a–c).

Figure 23.  

Bowie borneo sp. n., habitus, holotype male. a: Dorsal view; b: Ventral view. Scale bar: 1.0 mm (a, b).

Figure 24.  

Bowie engkilili sp. n., holotype female. a: Epigyne, ventral view; b: Vulva, dorsal view; c: Habitus, dorsal view; d: Habitus, ventral view. AB = anterior bulge of median plate, FD = fertilisation duct, LT = lateral teeth, SP = spermathecae. Scale bars: 0.2 mm (a, b), 2.0 mm (c, d).

Figure 25.  

Bowie engkilili sp. n., epigyne, posterior view. AB = anterior bulge of median plate, LT = lateral teeth. Scale bar: 0.1 mm.

Figure 26.  

Bowie sabah sp. n., palp, holotype. a: Prolateral view; b: Ventral view; c: Retrolateral view. C = conductor, DPO = dorso-proximal outgrowth of cymbium, E = embolus, RTA = retrolateral tibial apophysis, TA = tegular apophysis. Scale bar: 0.5 mm (a–c).

Figure 27.  

Bowie sabah sp. n. a: Paratype female, epigyne, ventral view; b: Same, vulva, dorsal view; c: Holotype male, habitus, dorsal view; d: Same, habitus, ventral view; e: Paratype female, habitus, dorsal view; f: Same, habitus, ventral view. FD = fertilisation duct, LT = lateral teeth, SP = spermathecae. Scale bars: 0.2 mm (a, b), 2.0 mm (c–f).

a

b

c

d

e

Figure 28.

Bowie spp., embolic tips.

a: B. fascination Jäger, 2022, male from Yunnan, ventral view. Scale bar: 0.05 mm.   
b: B. ninhbinh sp. n., paratype, ventral view. Scale bar: 0.1 mm.  
c: B. vinhphuc sp. n., paratype, dorsal view, arrow points at seam. Scale bar: 0.1 mm.  
d: B. borneo sp. n., holotype, ventral view. Scale bar: 0.02 mm.  
e: B. sabah sp. n., holotype, ventral view. Scale bar: 0.02 mm.  

Female (IZCAS-Ar 43541): PL 9.9, PW 7.8, AW 4.1, OL 11.0, OW 7.3. Eye diameters and interdistances: AME 0.29, ALE 0.27, PME 0.32, PLE 0.39, AME–AME 0.31, AME–ALE 0.71, PME–PME 0.43, PME–PLE 1.32, AME–PME 0.32, ALE–PLE 0.38, clypeus AME 0.42, clypeus ALE 0.95. Palp and leg measurements: palp 10.0 (3.3, 1.9, 2.1, -, 2.7), I 23.0 (6.7, 3.9, 5.7, 5.0, 1.7), II 21.4 (6.4, 3.7, 5.1, 4.6, 1.6), III 18.6 (5.6, 3.0, 3.8, 4.6, 1.6), IV 26.1 (7.2, 3.4, 5.8, 7.6, 2.1). Leg formula 4123. Spination of palp and legs: palp 131, 100, 131, 3020; femora I p021, d111, r111, II p112, d111, r111, III–IV p111, d111, r112; patellae I–II 000, III–IV 101; tibiae I–II v22222, III–IV p11, d111, r11, v222; metatarsi I–II v222, III p111, d012, r111, v222, IV p121, d012, r111, v2122. Chelicerae with 3 promarginal, 4 + 1 retromarginal teeth and with elongated patch of 27 tiny denticles along entire cheliceral furrow. Retromargin of chelicerae close to fang base with 11 bristles. Ventral tarsi and metatarsi I–II with sparse scopula. Palpal claw with 5 secondary teeth, leg claws I–II with 1, III with 2 and IV with 3 secondary teeth. Position of tarsal organ: I 1.41, II 1.34, III 1.25, IV 1.44.

Copulatory organ (Fig. 16a, b and Fig. 21a). Epigynal field wider than long, anterior bands separated from epigynal field. Lateral teeth arising laterally from median plate, curved and with pointed tips. Median plate with constricted posterior part and without distinct humped areas in posterior view. Internal duct system with two large vulval folds. Spermathecae separated by less than their diameter, with distinctly developed two chambers, smaller chamber with distinct external rim. Fertilisation ducts pointing laterally.

Colour (Fig. 16e and f). Deep reddish-brown with darker patterns. Dorsal prosoma with characteristic lighter median band, widened behind eyes, eye field with sparse white hairs and with distinctly marked fovea and radial markings. Sternum, labium, gnathocoxae and ventral coxae reddish-brown without patterns and gnathocoxae with lighter distal lips. Chelicerae black. Palps and legs deep reddish-brown. Dorsal opisthosoma brown with black patches, anterior margin with lighter area. Lateral opisthosoma spotted. Ventral opisthosoma brown with posteriorly converging lines of spots. Anterior lateral spinnerets laterally dark, posterior lateral and median spinnerets and anal tubercle light.

Large-sized Ctenidae (total length female 20.9). The species is assigned to the robustus-species group with the characteristics of stout tegular apophysis, simple stout embolus with broad base, presence of retro-proximal cymbial outgrowth, RTA arising medially to distally from palpal tibia, female possesses a transversally median plate with lateral teeth situated mainly laterally and not reaching the epigastric furrow. It resembles B. candidate Jäger, 2022 (see Jäger 2022: figs 254–262 and 280–284) by having similar lateral teeth (Fig. 16a and Fig. 21a), but can be distinguished by the median plate without distinct humped areas (Fig. 21a; median plate with humped areas best seen in posterior view in B. candidate), by the spermathecae separated by less than their diameter and smaller chamber with distinct external rim (Fig. 16b, spermathecae separated by about their diameter and smaller chamber without distinct external rim in B. candidate) and by the fertilisation ducts pointing laterally (Fig. 16b, fertilisation ducts pointing antero-medially in B. candidate). For the diagnosis of male, see Jäger (2022).

China (Yunnan, Fig. 1); Vietnam (Dien Bien, type locality).

Male (IZCAS-Ar 43540):

TATTTGGATCTTGGGCTGCTATAGCTGGGACAGCTATAAGAGTATTAATTCGTATAGAGCTAGGTCATTCTGGTAGATTATTTGGTGATGATCATTTATATAATGTAATTGTTACAGCTCATGCTTTTGTAATAATTTTTTTTATGGTTATGCCTATTTTAATTGGTGGTTTTGGAAACTGATTAGTTCCTTTGATATTAGGGGCTCCTGATATATCTTTTCCTCGTATAAATAATTTATCTTTTTGATTACTCCCTCCTTCATTATTTTTGTTATTTATATCTTCTATGGTTGAGATAGGGGTGGGAGCTGGTTGGACAGTGTATCCTCCTTTAGCTTCTAGTATTGGCCATATAGGAAGATCAATAGATTTTGCTATTTTTTCTTTACATTTAGCGGGAGCTTCTTCTATTATAGGGGCTGTTAATTTTATTTCTACAATTATTAATATACGTTTGTATGGAGTAAGAATAGAAAAGGTGCCTTTATTTGTATGATCTGTTCTAATTACTGCAGTATTATTGCTTTTATCTTTACCTGTATTAGCAGGTGCTATTACTATATTATTAACTGATCGTAATTTTAATACTTCTTTTTTTGACCCGGCTGGAGGAGGGGATCCAGTTTTATTTCAACATTTATTTTGATTTTTTG (GenBank accession number OP572108).

Female (IZCAS-Ar 43541):

TATTTGGATCTTGGGCTGCTATAGCTGGGACAGCTATAAGAGTATTAATTCGTATAGAGCTAGGTCATTCTGGTAGATTATTTGGTGATGATCATTTATATAATGTAATTGTTACAGCTCATGCTTTTGTAATAATTTTTTTTATGGTTATGCCTATTTTAATTGGTGGTTTTGGAAACTGATTAGTTCCTTTGATATTAGGGGCTCCTGATATATCTTTTCCTCGTATAAATAATTTATCTTTTTGATTACTCCCTCCTTCATTATTTTTGTTATTTATATCTTCTATGGTTGAGATAGGGGTGGGAGCTGGTTGGACAGTGTATCCTCCTTTAGCTTCTAGTATTGGCCATATAGGAAGATCAATAGATTTTGCTATTTTTTCTTTACATTTAGCGGGAGCTTCTTCTATTATAGGGGCTGTTAATTTTATTTCTACAATTATTAATATACGTTTGTATGGAGTAAGAATAGAAAAGGTGCCTTTATTTGTATGATCTGTTCTAATTACTGCAGTATTATTGCTTTTATCTTTACCTGTATTAGCAGGTGCTATTACTATATTATTAACTGATCGTAATTTTAATACTTCTTTTTTTGACCCGGCTGGAGGAGGGGATCCAGTTTTATTTCAACATTTATTTTGATTTTTTG (GenBank accession number OP572107).

Male (IZCAS-Ar 43542): PL 7.6, PW 5.9, AW 3.0, OL 5.8, OW 3.9. Eye diameters and interdistances: AME 0.26, ALE 0.19, PME 0.38, PLE 0.30, AME–AME 0.19, AME–ALE 0.29, PME–PME 0.22, PME–PLE 0.43, AME–PME 0.18, ALE–PLE 0.23, clypeus AME 0.11, clypeus ALE 0.55. Palp and leg measurements: palp 7.8 (2.7, 1.2, 1.3, -, 2.6), I 21.3 (5.6, 3.0, 5.8, 4.9, 2.0), II 19.6 (5.3, 2.7, 5.1, 4.7, 1.8), III 15.9 (4.3, 2.5, 3.4, 4.1, 1.6), IV 23.1 (5.9, 2.5, 5.7, 7.0, 2.0). Leg formula 4123. Spination of palp and legs: palp 151, 100, 101; femora I p021, d111, r112, II p112, d111, r112, III p212, d111, r112, IV p112, d111, r012; patellae I–IV 101; tibiae I p110, d111, r210, v22222, II p110, d111, r110, v22222, III p11, d200, r11, v222, IV p11, d111, r11, v222; metatarsi I–III p112, d010, r112, v222, IV p112, d010, r112, v2222. Chelicerae with 3 promarginal, 4 retromarginal teeth and with elongated patch of 19 tiny denticles along entire cheliceral furrow. Retromargin of chelicerae close to fang base with 6 bristles. Ventral tarsi and metatarsi I–II with sparse scopula. Right leg claws I–III with 2 and IV with 3 secondary teeth. Position of tarsal organ: I 1.27, II 1.28, III 1.06, IV 1.36.

Palp (Fig. 17a–c). RTA arising from tibia subdistally, stout and distally bifurcated. Cymbium tip slightly conical, with retro-proximally outgrowth. Embolus (Fig. 28b) arising at 7.30 o’clock position, its tip wide and blunt, situated in distal half of tegulum. Conductor arising at 12 o’clock position. Tegular apophysis arising subcentrally from tegulum, nearly round.

Colour (Fig. 18a and b). Reddish-brown to yellowish with dark patterns. Dorsal prosoma with characteristic lighter median band, widened behind eyes and with some white hairs, distinctly marked fovea and indistinct radial markings. Sternum and ventral coxae yellowish, labium brown and gnathocoxae brown with dark patterns. Chelicerae reddish-brown with longitudinal lines. Legs reddish brown-yellowish. Dorsal opisthosoma yellowish with black patches. Lateral opisthosoma yellowish with darker spots. Ventral opisthosoma yellowish with dark patterns; epiandrium and muscle sigilla light. Spinnerets and anal tubercle light.

Female

Unknown.

Variation: Paratype male (IZCAS-Ar 43543): PL 7.4, OL 5.7.

Medium-sized Ctenidae (total length male 13.1–13.4). The new species is assigned to the robustus-species group with the characteristics of stout tegular apophysis, simple stout embolus with broad base, presence of retro-proximal cymbial outgrowth and RTA arising subdistally from palpal tibia. It resembles B. dodo Jäger, 2022 (see Jäger 2022: figs 241–243 and 267–268) by having similar conductor, broad embolus and retro-proximal cymbial outgrowth (Fig. 17a–c and Fig. 28b), but can be distinguished by the tegular apophysis nearly round and without concave (Fig. 17b, tegular apophysis with distinct concave on retrolateral side in B. dodo) and by the RTA distally bifurcated (Fig. 17b, RTA having no bifurcation in B. dodo).

The specific name refers to the type locality and is a noun in apposition.

Vietnam (Ninh Binh, type locality; Fig. 1).

Male (IZCAS-Ar 43542):

TACTTGGATCTTGGGCTGCTATGGCAGGGACAGCTATAAGAGTATTAATTCGGATGGAATTAGGCCATTCTGGGAGATTGTTAGGTGATGATCATTTATACAATGTAATTGTTACTGCACATGCTTTTGTAATGATTTTTTTTATAGTAATGCCTATTTTAATTGGGGGTTTTGGAAATTGGTTAGTACCTTTGATATTAGGGGCTCCTGATATATCTTTTCCTCGAATAAATAATTTGTCTTTTTGGTTACTTCCTCCTTCGTTATTTTTATTATTTATATCTTCAATAGTTGAGATAGGAGTTGGAGCTGGATGAACGGTATATCCTCCTTTAGCTTCTAGTATTGGTCATATAGGGAGATCTATAGATTTTGCTATTTTTTCTTTACATTTAGCGGGGGCTTCTTCTATTATAGGAGCGGTAAATTTTATTTCTACGATTATTAATATGCGTTTGTATGGGATGACTATAGAGAAAGTACCTTTATTTGTGTGATCTGTTTTAATTACTGCGGTATTGTTATTATTGTCTTTACCTGTTTTAGCAGGTGCTATTACTATATTGTTAACTGATCGAAATTTTAATACTTCTTTTTTTGATCCGGCTGGGGGTGGTGATCCTGTTTTGTTTCAACATTTATTTTGATTTTTTG (GenBank accession number OP572110).

Male (IZCAS-Ar 43544): PL 6.4, PW 5.0, AW 2.4, OL 3.1, OW 3.5. Eye diameters and interdistances: AME 0.23, ALE 0.24, PME 0.28, PLE 0.26, AME–AME 0.22, AME–ALE 0.42, PME–PME 0.25, PME–PLE 0.41, AME–PME 0.20, ALE–PLE 0.21, clypeus AME 0.21, clypeus ALE 0.53. Palp and leg measurements: palp 7.4 (2.8, 0.9, 1.4, -, 2.3), I 17.7 (5.2, 2.5, 4.3, 4.3, 1.4), II 15.9 (4.7, 2.4, 3.7, 3.8, 1.3), III 13.4 (3.8, 1.9, 2.9, 3.6, 1.2), IV 19.5 (5.5, 2.3, 4.3, 5.8, 1.6). Leg formula 4123. Spination of palp and legs: palp 141, 100, 1010; femora I p021, d111, r112, II p112, d111, r1111, III p1111, d111, r1111, IV p002, d111, r111; patellae I–IV 101; tibiae I p110, d111, r21, v22222, II p20, d111, r110, v22222, III–IV p11, d111, r11, v222; metatarsi I p11, d002, r111, v222, II p111, r111, v222, III p111, d012, r211, v222, IV p122, d012, r111, v2122. Chelicerae with 3 promarginal, 4 retromarginal teeth and with elongated patch of 12 tiny denticles along entire cheliceral furrow. Retromargin of chelicerae close to fang base with 4 bristles. Sparse scopula restricted almost entirely to tarsi, only metatarsi I–II with sparse scopula hairs. Leg claws I–III with 3 and IV with 4 secondary teeth. Position of tarsal organ: I with 1.17, II 1.07, III 0.90, IV 1.09.

Palp (Fig. 19a–c). RTA arising from tibia subdistally, with wide base and narrow tip. Cymbium tip slightly conical and with retrolatero-proximal outgrowth. Embolus (Fig. 28c) arising at 7 o’clock position, with a large seam from the middle to the tip. Conductor arising at 12 o’clock position. Tegular apophysis arising at 6 o’clock position, covering partly membranous extension part of embolus.

Colour (Fig. 20c and d). Deep reddish-brown with darker patterns. Dorsal prosoma with characteristic lighter median band, widened behind eyes, eye field and lateral field with white dense hairs, with distinctly marked fovea and radial markings. Sternum and ventral coxae yellowish-brown without patches, labium and gnathocoxae reddish-brown with lighter distal lips. Chelicerae deep reddish-brown without patterns. Palps and legs reddish-brown without patterns. Dorsal opisthosoma yellowish-brown with black patches and anterior margin with lighter area. Lateral opisthosoma without spots. Ventral opisthosoma dark brown with posteriorly converging lines of spots. Anterior lateral spinnerets laterally dark, posterior lateral and median spinnerets and anal tubercle light.

Female (IZCAS-Ar 43728): PL 6.7, PW 5.2, AW 3.1, OL 8.5, OW 7.0. Eye diameters and interdistances: AME 0.25, ALE 0.21, PME 0.28, PLE 0.26, AME–AME 0.22, AME–ALE 0.48, PME–PME 0.29, PME–PLE 0.57, AME–PME 0.23, ALE–PLE 0.23, clypeus AME 0.19, clypeus ALE 0.58. Palp and leg measurements: palp 6.9 (2.5, 1.3, 1.4, -, 1.7), I 16.3 (4.8, 2.6, 4.1, 3.5, 1.3), II 15.0 (4.5, 2.4, 3.5, 3.4, 1.2), III 13.1 (4.1, 2.0, 2.7, 3.1, 1.2), IV 18.8 (5.2, 2.2, 4.1, 5.6, 1.7). Leg formula 4123. Spination of palp and legs: palp 131, 000, 1111, 2101; femora I p021, d111, r021, II p112, d111, r021, III p112, d111, r112, IV p111, d111, r001; patellae I–II 000, III–IV 101, tibiae I–II v22222, III–IV p11, d111, r11, v222; metatarsi I–II v222, III p112, d010, r112, v222, IV p112, r112, v2222. Chelicerae with 3 promarginal, 4 retromarginal teeth and with elongated patch of 10 tiny denticles along entire cheliceral furrow. Retromargin of chelicerae close to fang base with 5 bristles. Sparse scopula restricted almost entirely to tarsi, only metatarsi I–II with sparse scopula hairs. Palpal claw with 6 secondary teeth, leg claws I with 1, II–III with 2 and IV with 3 secondary teeth. Position of tarsal organ: I 1.03, II 0.97, III 0.95, IV 1.42.

Copulatory organ (Fig. 20a, b and Fig. 21b). Epigynal field wider than long. Lateral teeth arising laterally from median plate, curved and with pointed tips. Median plate with constricted posterior part and distinct humped areas best seen in posterior view. Internal duct system with two large vulval folds. Spermathecae separated by less than their diameter, larger chamber round and apically swollen and smaller one spherical. Fertilisation ducts pointing medially.

Colour (Fig. 20e and f). As in male, except for: dorsal prosoma with distinct light median band and without white dense hairs. Lateral opisthosoma spotted.

Variation: Paratype male (IZCAS-Ar 43545): PL 7.4, OL 6.5. Second paratype female (IZCAS-Ar 43729): PL 7.6, OL 7.8.

Small to medium-sized Ctenidae (total length male 9.5–13.9, female 15.2–15.4). The new species is assigned to the robustus-species group with the characteristics of stout tegular apophysis, simple stout embolus with broad base and short apical part, presence of retro-proximal cymbial outgrowth, RTA arising subdistally from palpal tibia, female possesses a transversally oval median plate with lateral teeth situated laterally and not reaching the epigastric furrow. It resembles B. yassassin Jäger, 2022 (see Jäger 2022: figs 287–290 and 307–313) by having similar retro-proximal cymbial outgrowth, conductor, tegular apophysis, RTA (Fig. 19a–c) and lateral teeth (Fig. 20a and Fig. 21b), but can be distinguished by the embolus with a large seam from the middle to the tip (Fig. 19b, arrowed in Fig. 28c; absent in B. yassassin), by the median plate with distinct humped areas best seen in posterior view (Fig. 21b; absent in B. yassassin), by the spermathecae separated by less than their diameter, larger chamber round and apically swollen (Fig. 20b; spermathecae separated by about their diameter, larger chamber oval in B. yassassin) and by the fertilisation ducts pointing medially (Fig. 20b; fertilisation ducts pointing postero-medially in B. yassassin).

The specific name refers to the type locality and is a noun in apposition.

Vietnam (Vinh Phuc, type locality; Fig. 1).

Male (IZCAS-Ar 43730): PL 3.8, PW 2.9, AW 1.2, OL 3.2, OW 2.2. Eye diameters and interdistances: AME 0.17, ALE 0.15, PME 0.22, PLE 0.21, AME–AME 0.14, AME–ALE 0.11, PME–PME 0.24, PME–PLE 0.27, AME–PME 0.11, ALE–PLE 0.13, clypeus AME 0.09, clypeus ALE 0.37. Palp and leg measurements: palp 5.1 (2.5, 0.5, 0.8, -, 1.3), I 12.1 (2.8, 1.5, 3.2, 3.3, 1.3), II 10.9 (3.2, 1.3, 2.6, 2.7, 1.1), III 9.6 (2.7, 1.1, 2.1, 2.6, 1.1), IV 15.1 (4.0, 1.4, 3.3, 4.7, 1.7). Leg formula 4123. Spination of palp and legs: palp 151, 100, 1110; femora I p012, d111, r012, II–III p112, d111, r112, IV p112, d111, r002; patellae I–II 100, III–IV 101; tibiae I p010, r010, v212222, II p100, d001, r110, v22222, III–IV p11, d111, r11, v222; metatarsi I p111, r011, v222, II p111, d002, r111, v222, III p111, d012, r111, v222, IV p111, d012, r111, v2212. Chelicerae with 3 promarginal, 4 retromarginal teeth and with elongated patch of 8 tiny denticles along entire cheliceral furrow. Retromargin of chelicerae close to fang base with 4 bristles. Sparse scopula on all tarsi. Leg claws I–II with 5 and III–IV with 4 secondary teeth. Position of tarsal organ: I 1.07, II 0.89, III 0.72, IV 1.06.

Palp (Fig. 22a–c). RTA protruding at an almost right angle from tibia in ventral view, with slightly hooked tip. Cymbium tip slightly conical and with weakly developed dorso-proximal outgrowth. Embolus (Fig. 28d) arising at 7 o’clock position. Conductor arising at 11 o’clock position. Tegular apophysis arising from tegulum sub-proximally, with narrow base and wide tip.

Colour (Fig. 23a and b). Yellowish-brown with darker patterns. Dorsal prosoma with characteristic lighter median band, widened behind eyes, eye field with sparse white hairs, with distinctly marked fovea and distinct radial markings. Sternum, labium, gnathocoxae and ventral coxae yellowish-brown without patterns. Chelicerae reddish-brown with longitudinal patterns. Palps and legs yellowish-brown, legs II–IV with distinct patterns. Dorsal opisthosoma yellowish with black patches, anterior margin and central region light. Lateral opisthosoma spotted. Ventral opisthosoma yellowish-brown with posteriorly converging lines of spots. Anterior lateral spinnerets laterally dark, posterior lateral and median spinnerets and anal tubercle light.

Female

Unknown.

Small Ctenidae (total length male 7.0). The new species is assigned to the chinagirl-species group with the characteristics of compact embolus, thick-walled, with rounded edges and with an entire margin, the tegular apophysis is longitudinally elongated, not covering the embolus. It resembles B. abdulmajid Jäger, 2022 (see Jäger 2022: figs 638–643 and 696–699) by having similar dorso-proximal cymbial outgrowth, embolus and RTA (Fig. 22a–c and Fig. 28d), but can be distinguished by the base of tegular apophysis having no bulge (Fig. 22b; the base of tegular apophysis bulging beyond tegulum margin in B. abdulmajid), by the tibia having no distinct broad longitudinal ridge ventrally (Fig. 22b; tibia with distinct broad longitudinal ridge ventrally in B. abdulmajid) and by the conductor nearly quadrilateral (Fig. 22b; conductor nearly oval in B. abdulmajid).

The specific name refers to the type locality and is a noun in apposition.

Malaysia (Borneo, type locality; Fig. 1).

Male (IZCAS-Ar 43730):

GGTTTGGTGCTTGGGCTTCTATAGCAGGTACGTCTATAAGAGTTTTGATTCGAATGGAATTAGGACATTCTGGAAGATTATTAGGGGATGATCATTTATATAATGTTGTTGTTACTGCTCATGCTTTTGTTATAATTTTTTTTATAGTGATACCTATTTTAATTGGTGGTTTTGGAAATTGGTTGGTTCCTTTAATATTAGGAGCTCCTGATATATCTTTTCCTCGTATAAATAATTTGTCGTTTTGATTACTTCCTCCTTCTTTATTTTTATTGTTTATATCTTCTATGACTGAGATAGGGGTGGGAGCTGGTTGGACGGTTTATCCACCTTTGGCTTCTGGAATTGGTCATGCAGGAAGATCGATAGATTTTGCTATCTTTTCTCTCCATTTAGCAGGTGCTTCTTCTATTATAGGAGCTATTAATTTTATTTCTACGATTATTAATATACGATTATTAGGAATGAGAATGGAAAAGGTTCCTTTGTTTGTATGGTCTGTTTTTATTACTGCAGTTTTATTATTATTATCTTTACCTGTTTTAGCGGGTGCTATTACTATATTATTAACGGATCGTAATTTTAATACTTCTTTTTTCGATCCTGCTGGAGGAGGGGATCCAATTTTATTTCAACATTTATTTTGATTTTTTG (GenBank accession number OP572103).

Male

Unknown.

Female (IZCAS-Ar 43731): PL 4.5, PW 3.6, AW 2.3, OL 3.9, OW 2.3. Eye diameters and interdistances: AME 0.21, ALE 0.11, PME 0.26, PLE 0.23, AME–AME 0.15, AME–ALE 0.31, PME–PME 0.27, PME–PLE 0.35, AME–PME 0.13, ALE–PLE 0.17, clypeus AME 0.14, clypeus ALE 0.41. Palp and leg measurements: palp 4.7 (1.6, 0.9, 1.1, -, 1.1), I missing, II 10.8 (3.0, 1.7, 2.7, 2.5, 0.9), III 10.2 (2.7, 1.7, 2.2, 2.5, 1.1), IV 14.4 (3.7, 1.6, 3.4, 4.2, 1.5). Leg formula 4123. Spination of palp and legs: palp 131, 001, 112, 2012; femora II p110, d111, r112, III p012, d111, r112, IV p002, d111, r112; patellae II 000, III–IV 101; tibiae II v22222, III–IV p11, d111, r11, v222; metatarsi II v222, III p112, d010, r112, v222, IV p112, d010, r112, v2222. Chelicerae with 3 promarginal, 4 retromarginal teeth and with elongated patch of 5 tiny denticles along entire cheliceral furrow. Retromargin of chelicerae close to fang base with 5 bristles. Sparse scopula restricted almost entirely to tarsi. Palpal claw with 7 secondary teeth, leg claws II–IV with 3 secondary teeth. Position of tarsal organ: II 0.75, IV 0.85.

Copulatory organ (Fig. 24a, b and Fig. 25). Epigynal field slightly longer than wide, with narrow separated anterior bands laterally. Median plate with a median bulge in anterior half, laterally with two separate sclerotised patches, constricted at lateral teeth, the latter moderately pointed. Spermathecae small, separated by more than twice their diameter, roundish, smaller chamber bend at right angle, laterad.

Colour (Fig. 24c and d). Reddish-brown to yellowish with dark patterns. Dorsal prosoma with characteristic lighter median band, widened behind eyes, distinctly marked fovea and indistinct radial markings. Sternum and ventral coxae yellowish, labium and gnathocoxae reddish-brown. Chelicerae black. Leg reddish-brown. Dorsal and lateral opisthosoma black. Ventral opisthosoma black with light patterns. Spinnerets and anal tubercle black.

Small Ctenidae (total length female 8.4). The new species resembles B. withinyou Jäger, 2022 (see Jäger 2022: figs 629–630 and 634–636) by having similar fertilisation ducts (Fig. 24b), but can be distinguished by the lateral teeth pointing medially (Fig. 24a; lateral teeth pointing postero-medially in B. withinyou), by the median plate with two separate sclerotised patches laterally and a longer median bulge in anterior half (Fig. 24a and Fig. 25; median plate with shorter median bulge in anterior half in B. withinyou) and by the spermathecae separated by more than twice their diameter (Fig. 24b; spermathecae separated by 1.5 times their diameter in B. withinyou).

The specific name refers to the type locality and is a noun in apposition.

Malaysia (Engkilili, type locality; Fig. 1).

Female (IZCAS-Ar 43731):

TATTTGGGGCTTGAGCTTCTATAGCTGGTACATCTATAAGTGTTTTGATTCGTATGGAGTTGGGACATTCGGGGAGAATATTGGGAGATGATCATCTCTATAATGTTATTGTTACTGCTCATGCTTTTGTTATAATTTTTTTTATAGTTATACCTATTTTAATTGGAGGTTTTGGTAATTGGTTGGTTCCTTTAATGTTAGGGGCTCCTGATATGTCGTTTCCTCGAATAAATAATTTGTCTTTTTGGTTACTTCCTCCTTCTTTATTTTTATTGTTTATGTCTTCTATAACTGAAATAGGGGTAGGAGCTGGTTGAACGGTGTATCCTCCTTTGGCTTCAAGAATGGGTCATGCTGGTAGATCTATGGATTTTGCTATTTTTTCTCTTCATTTAGCTGGGGCGTCTTCTATTATAGGTGCTATTAATTTTATTTCTACTATTATTAATATGCGTTTATTAGGAATGAGAATAGAGAAAGTTCCTTTGTTTGTATGGTCTGTTTTTATTACTGCGGTATTGTTATTATTGTCTCTTCCTGTTTTGGCAGGAGCTATTACTATATTGTTAACTGATCGTAATTTTAATACTTCTTTTTTTGATCCGGCTGGAGGGGGAGATCCGATTTTATTTCAACATTTATTTTGATTTTTT (GenBank accession number OP572106).

Male (IZCAS-Ar 43732): PL 6.4, PW 5.1, AW 2.3, OL 4.8, OW 3.6. Eye diameters and interdistances: AME 0.22, ALE 0.17, PME 0.27, PLE 0.23, AME–AME 0.25, AME–ALE 0.40, PME–PME 0.34, PME–PLE 0.49, AME–PME 0.17, ALE–PLE 0.29, clypeus AME 0.11, clypeus ALE 0.52. Palp and leg measurements: palp 8.2 (3.0, 1.2, 1.6, -, 2.4), I 20.5 (5.6, 2.6, 5.4, 5.1, 1.8), II 17.4 (5.1, 2.5, 4.3, 4.2, 1.3), III 14.5 (4.3, 2.2, 3.2, 3.7, 1.1), IV 22.4 (6.1, 2.2, 5.4, 6.8, 1.9). Leg formula 4123. Spination of palp and legs: palp 161, 001, 111; femora I p021, d111, r1111, II p112, d111, r112, III p001, d222, r112, IV p012, d111, r112; patellae I–IV 101; tibiae I p110, d111, r11, v22222, II p110, d11, r11, v22222, III–IV p11, d111, r11, v222; metatarsi I–II p111, d012, r111, v222, III p111, d002, r111, v222, IV p111, d012, r111, v222. Chelicerae with 3 promarginal, 4 retromarginal teeth and with elongated patch of 23 tiny denticles along entire cheliceral furrow. Retromargin of chelicerae close to fang base with 6 bristles. Sparse scopula on all tarsi and metatarsi I–III. Leg claws I with 5, II with 4, III with 5 and IV with 6 secondary teeth. Position of tarsal organ: I with 1.64, II 1.15, III 0.96, IV 1.28.

Palp (Fig. 26a–c). RTA arising from tibia subdistally, with slightly hooked tip. Cymbium tip slightly conical and with weakly-developed dorso-proximal outgrowth. Embolus (Fig. 28e) arising at 8 o’clock position, with membranous extension at its base and with spermophor opening situated subapically. Conductor arising at 10 o’clock position. Tegular apophysis arising at 6 o’clock position, its base situated on a slightly proximally protruding part of tegulum.

Colour (Fig. 27c and d). Yellowish-brown with darker patterns. Dorsal prosoma with characteristic lighter median band, widened behind eyes, eye field with sparse white hairs, with distinctly marked fovea and radial markings. Sternum and ventral coxae III + IV yellowish-brown with patches, coxae I + II yellowish-brown without patterns, labium and gnathocoxae yellowish-brown with lighter distal lips. Chelicerae dark yellowish-brown with longitudinal patterns. Palps and legs yellowish-brown, legs III + IV with distinct patterns. Dorsal opisthosoma yellowish-brown with black patches, anterior margin and central region light. Lateral opisthosoma spotted. Ventral opisthosoma dark brown with posteriorly converging lines of spots. Anterior lateral spinnerets laterally dark, posterior lateral and median spinnerets and anal tubercle light.

Female (IZCAS-Ar 43733): PL 5.9, PW 4.4, AW 3.0, OL 5.2, OW 3.5. Eye diameters and interdistances: AME 0.23, ALE 0.19, PME 0.27, PLE 0.25, AME–AME 0.20, AME–ALE 0.44, PME–PME 0.33, PME–PLE 0.52, AME–PME 0.17, ALE–PLE 0.28, clypeus AME 0.13, clypeus ALE 0.51. Palp and leg measurements: palp 5.8 (2.1, 1.1, 1.2, -, 1.4), I 13.9 (4.0, 2.1, 3.6, 3.1, 1.1), II 12.8 (3.8, 2.1, 3.0, 2.9, 1.0), III 11.3 (3.5, 1.7, 2.4, 2.7, 1.0), IV 16.7 (4.6, 1.9, 3.8, 4.9, 1.5). Leg formula 4123. Spination of palp and legs: palp 131, 001, 112, 203; femora I p021, d111, r011, II p012, d111, r111, III p112, d111, r112, IV p002, d111, r112; patellae I–II 000, III–IV 101; tibiae I–II v22222, III–IV p11, d111, r11, v222; metatarsi I–II v222, III–IV p111, d012, r111, v222. Chelicerae with 3 promarginal, 4 retromarginal teeth and with elongated patch of 12 tiny denticles along entire cheliceral furrow. Retromargin of chelicerae close to fang base with 7 bristles. Sparse scopula restricted almost entirely to tarsi, only metatarsi I–II with sparse scopula hairs. Palpal claw with 5 secondary teeth, leg claws I with 2, II with 3, III with 4 and IV with 3 secondary teeth. Position of tarsal organ: I 0.97, II 0.84, III 0.75, IV 1.06.

Copulatory organ (Fig. 27a and b). Epigynal field roughly as long as wide; constrictive anterior width/widest width: 0.53/1.15. Lateral teeth originating at posterior margin of median plate. Internal duct system with two large vulval folds laterally. Spermathecae bottle gourd-shaped. Vulval folds separated by less than the spermathecae length and subparallel medially. Fertilisation ducts pointing antero-medially.

Colour (Fig. 27e and f). As in male, except for being darker, reddish-brown. Chelicerae dark reddish-brown without pattern.

Medium-sized Ctenidae (total length male 11.2, female 11.1). The new species is assigned to the scarymonsters-species group with the characteristics of embolus with a basal, ventral bulge (best seen in prolateral view), tegulum bulging proximally at tegular apophysis base, the subdistally arising, apically pointed RTA in males, transversally oval median plate and lateral teeth situated at posterior margin of epigyne and spermathecae bottle gourd-shaped. It resembles B. neukoeln Jäger, 2022 (see Jäger 2022: figs 440–446 and 448–460) by having similar dorso-proximal cymbial outgrowth, RTA (Fig. 26a–c) and lateral teeth (Fig. 27a), but can be distinguished by the embolus tip visible in ventral view (Fig. 26b and Fig. 28e; tegular apophysis covering the embolus tip in ventral view in B. neukoeln), by the tegular apophysis longitudinally orientated (Fig. 26b; tegular apophysis diagonally orientated in B. neukoeln), by the conductor nearly quadrilateral (Fig. 26b; conductor nearly elliptic in B. neukoeln) and by the constrictive anterior width/widest width: 1/2 (Fig. 27a; constrictive anterior width/widest width: 1/3 in B. neukoeln).

The specific name refers to the type locality and is a noun in apposition.

Malaysia (Borneo, type locality; Fig. 1).

Male (IZCAS-Ar 43732):

GGTTTGGAGCTTGAGCTTCTATAGTAGGAACATCTATAAGAGTATTAATTCGTATAGAATTAGGACATTCTGGAAGATTATTAGGAGATGATCATTTATATAATGTAGTTGTTACTGCTCATGCTTTTGTTATGATTTTTTTTATAGTAATGCCTATTTTAATTGGAGGTTTTGGAAATTGATTAGTTCCTTTAATATTAGGGGCTCCTGATATATCTTTTCCTCGAATAAATAATTTATCATTTTGATTACTTCCTCCTTCTTTATTCTTATTATTTATGTCTTCTATAACTGAGATAGGAGTAGGAGCTGGTTGAACAGTATATCCTCCCTTAGCTTCTAGAATAGGACATATGGGAAGATCAATGGATTTTGCTATTTTTTCTCTTCATTTAGCTGGGGCTTCCTCTATTATAGGAGCTATTAATTTTATTTCTACAATTATTAATATACGATTATTGGGAATAAGAATAGAGAAAGTTCCATTATTTGTGTGGTCTGTTTTTATTACTGCGGTATTGTTGTTATTGTCTTTACCTGTTTTAGCAGGTGCTATTACTATATTATTAACGGATCGTAATTTTAATACTTCTTTTTTTGACCCAGCTGGGGGGGGGGATCCCATTTTATTTCAACATTTATTTTGATTTTTGC (GenBank accession number OP572111).

Female (IZCAS-Ar 43733):

GGTTTGGAGCTTGAGCTTCTATAGTAGGAACATCTATAAGAGTATTAATTCGTATAGAATTAGGACATTCTGGAAGATTATTAGGAGATGATCATTTATATAATGTAGTTGTTACTGCTCATGCTTTTGTTATGATTTTTTTTATAGTAATGCCTATTTTAATTGGAGGTTTTGGAAATTGATTAGTTCCTCTAATATTAGGGGCTCCTGATATATCTTTTCCTCGAATAAATAATTTATCATTTTGATTACTTCCTCCTTCTTTATTCTTATTATTTATGTCTTCTATAACTGAGATAGGAGTAGGAGCTGGTTGAACAGTATATCCTCCCTTAGCTTCTAGAATAGGACATATGGGAAGATCAATGGATTTTGCTATTTTTTCTCTTCATTTAGCTGGGGCTTCCTCTATTATAGGAGCTATTAATTTTATTTCTACAATTATTAATATACGATTATTGGGAATAAGAATAGAGAAAGTTCCATTATTTGTGTGGTCTGTTTTTATTACTGCGGTATTGTTGTTATTGTCTTTACCTGTTTTAGCAGGTGCTATTACTATATTATTAACGGATCGTAATTTTAATACTTCTTTTTTTGACCCAGCTGGGGGGGGGGATCCCATTTTATTTCAACATTTATTTTGATTTTTGC (GenBank accession number OP572109).

B. sabah sp. n. belongs to the scarymonsters-species group because of the abovementioned characteristics, so the “diagonally orientated TA” in the diagnosis of the scarymonsters-species group in Jäger (2022) should be changed to “tegulum bulging proximally at TA base”.