An annotated checklist of grasshoppers (Orthoptera, Acridoidea) from Mongolia

Enkhtsetseg Gankhuyag; Altanchimeg Dorjsuren; Eun Hwa Choi; Ui Wook Hwang

doi:10.3897/BDJ.11.e96705

Biodiversity Data Journal : Taxonomy & Inventories

PDF

Taxonomy & Inventories

An annotated checklist of grasshoppers (Orthoptera, Acridoidea) from Mongolia

Enkhtsetseg Gankhuyag^‡, Altanchimeg Dorjsuren^§,|, Eun Hwa Choi^‡, Ui Wook Hwang^¶,#,¤,«

‡ Department of Biology, Teachers College, and Institute for Phylogenomics and Evolution, Kyungpook National University, Daegu 41566, South Korea

§ Institute of Biology, Mongolian Academy of Sciences, Ulaanbaatar 133330, Mongolia

| College of Life Sciences, Inner Mongolia University, Hohhot, 010031, China

¶ Institute of Phylogenomics and Evolution, and Department of Biology, Teachers College Kyungpook National University, Daegu 41566, Republic of Korea

# Institute for Korean Herb-Bio Convergence Promotion, Kyungpook National University, Daegu 41566, South Korea

¤ School of Industrial Technology Advances, Kyungpook National University, Daegu 41566, South Korea

« Phylomics Inc., Daegu 41910, South Korea

Corresponding author: Ui Wook Hwang (uwhwang@knu.ac.kr)

Academic editor: Sheryl Yap

Received: 25 Oct 2022 | Accepted: 20 Jan 2023 | Published: 13 Mar 2023

This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.

Citation: Gankhuyag E, Dorjsuren A, Choi EH, Hwang UW (2023) An annotated checklist of grasshoppers (Orthoptera, Acridoidea) from Mongolia. Biodiversity Data Journal 11: e96705. https://doi.org/10.3897/BDJ.11.e96705

Abstract

Background

Grasshoppers (Acridoidea, Orthoptera) are the dominant herbivores in grassland ecosystems worldwide. They can increase rangeland productivity by stimulating plant growth and accelerating nutrient cycling. This article presents a comprehensive checklist of grasshoppers in Mongolia. Until then, the available information was very scattered, based on old studies of Mongolian grasshoppers, recorded in a few international catalogues and databases, individual records and research work on agroecosystem communities. However, the available information on the composition of the Orthopteran fauna in Mongolia was sometimes unclear or non-existent and these dubious data were excluded from the present study. In addition, the grasshopper distribution analysis used the standardised personal collection of D. Altanchimeg. We also present a list of grasshoppers, as well as their distribution and abundance, in countries adjacent to Mongolia, such as Russia, China and South Korea. The surveys covered six types of natural zones: high mountain, taiga, forest-steppe, steppe, desert steppe and desert; desert steppe and steppe zones are the most widely distributed. We hope to have contributed significantly to the study of the distribution of grasshopper species in all these natural zones.

New information

In this study, a total of three families of Acridoidea belonging to eight subfamilies, 17 tribes, 52 genera and 128 species are reported for the various natural zones. The recorded species belong to eight subfamilies: Gomphocerinae are the most numerous with 56 species recorded, followed by Oedipodinae (51 species), Thrinchinae (nine species), Melanoplinae (six species), Calliptaminae (three species), Dericorythinae, Acridinae, Egnatiinae (one species each).

Keywords

distribution, fauna, natural zone, Pamphagidae, Acrididae, Dericorythidae, Mongolia

Introduction

Locust and grasshoppers (Orthoptera, Acridoidea) are essential herbivores in grassland ecosystems worldwide (Latchininsky et al. 2011, Fang et al. 2015). They aid in plant growth and nutrient cycling and play an important part in food chains (Kietzka et al. 2021). However, locust and grasshopper outbreaks are considered a global problem. They can destroy grasslands and crops and inflict severe economic impacts on crops and rangelands (Hewitt 1977, Gupta 1983, Lockwood and Lockwood 2008, Latchininsky et al. 2011, Zhang et al. 2019, Lecoq and Zhang 2019, Lecoq and Cease 2022). A few grasshopper species have been proposed as ecological indicators of ecosystem health as they are susceptible to changes in land use and climate (Marini et al. 2008, Fartmann et al. 2012, Bazelet and Samways 2012, Uchida and Ushimaru 2014). Climate change and geographical characteristics are critical factors that determine grasshopper population growth. The grasshoppers (Acridoidea) are the largest superfamily of the orthopterans with 28 families, 140 subfamilies, 265 tribes, 2,459 genera (57 subgenera) and 10,531 valid species (1,951 subspecies) distributed throughout the world, except Antarctica (Cigliano et al. 2022). Mongolia is a landlocked country with diverse terrain, surrounded by mountains to the north and west and the forest-steppe of the Gobi Desert to the south. The majority of its land is covered by grassland steppe. Mongolia has six main natural zones and belts (that are divided into sub-divisions): alpine (high mountain) and mountain taiga, mixed and deciduous forests, steppe, desert steppe (Gobi) and desert zones (Yembuu 2021). Mongolia has a high elevation and cold and dry climate (Worden and Savada 1991, Gombobaatar et al. 2014).

Of these species, Mongolia contains 128 species in 52 genera and three families (Pamphagidae, Dericorythidae and Acrididae). In the early 1960s, several foreign and Mongolian researchers started undertaking expeditions for the Mongolian insect species checklist. Some of these expeditions were undertaken collaboratively with other countries, such as Poland (1962–1964), Hungary (1963–1968), Germany (1962–1964), the Czech Republic (1965–1966) and Russia (1967–1987). This research facilitated the study of insect distribution (physical and geographical distribution) in the different provinces of Mongolia and collected several grasshopper specimens. Referring to Rentsendorj and Khodroi (2020), the study on the list of grasshoppers in Mongolia was undertaken from 1951 to 2019 and a list of 110 species was established. Of them, three different families and 48 genera between 1951 and 1989 (Mistshenko and Bey-Bienko 1951, Mistshenko 1968, Günther 1971, Chogsomzhav 1971, Chogsomzhav 1974b, Chogsomzhav 1977, Chogsomzhav 1989, Gorochov et al. 1989) and 81 species of two families and 33 genera between 1990–2019 (Batkhuyag 1995, Batnaran et al. 1999, Batnaran 2008, Ganbold 2009, Altanchimeg et al. 2014, Altanchimeg and Nonnaizab 2013, Batkhuyag et al. 2014, Gandulam 2016, Otgonchimeg 2017) were added to the list. Moreover, 69 short-horned grasshopper species, which belonged to two families and 31 genera, were identified between 2013 and 2019 in Mongolia (Rentsendorj and Khodroi 2020). Recently, the species list of Mongolian grasshoppers was updated to three families, 49 genera and 127 species (Batkhuyag and Batnaran 2021). However, as this species list included uncertain specimen records and unclear sources, the accuracy of these sources needed to be checked.

The present study is the first comprehensive study to update the diversity of Mongolian grasshoppers (Acridoidea), including information on species traits and distribution by natural zones. To create this annotated checklist, we reviewed all published materials related to short-horned grasshopper species that were newly discovered in Mongolia since the 1930s, followed by a taxonomic analysis using important source information from the Orthoptera Species File (Cigliano et al. 2022). The current study's findings will give fundamental information regarding the grasshopper diversity of Mongolia. Furthermore, it can inspire local scientists interested in topics such as ecology, biology, medicine, agriculture and education.

Materials and methods

The list presented in the present paper is based on literature records of grasshopper species in Mongolia available up to May 2022. The taxa that were reported from Mongolia are listed taxonomically by subfamily and alphabetically within each subfamily, tribe and genus. Each species was examined in Mongolia and the citation of the first or most reliable reference to support this record is provided. First, the references to recorded species in Mongolia reported in the Orthoptera Species File online version 5.0/5.0 (Cigliano et al. 2022), Institute of Biology, Mongolian Academy of Sciences (MAS) (Chogsomzhav 1975, Altanchimeg et al. 2014), Plant Protection Research Institute of Mongolia (Batkhuyag et al. 2014) and the grasshopper list in Mongolia from the Mongolian Journal of Agricultural Sciences were checked and, if erroneous, corrected in the current lists. Changes in systematic status and synonymies have been proposed by (Chogsomzhav (1989), Altanchimeg (2011) and Batkhuyag and Batnaran (2021).

The current work is an annotated study of grasshoppers (Acridoidea) in Mongolia, with an emphasis on a rare and unknown grasshopper species. Except for long-horned grasshoppers and crickets, this research systematises Acridoidea. Likewise, the list is used in taxonomic sequences by superfamily, family, subfamily, tribe, genus and species. Depending on the natural zone where the grasshopper was collected, they belong to that area. Grasshopper distribution in six different natural zones, distribution of grasshopper species in neighbouring countries on the northern border with Russia and south with China and comparison with the South Korean population were performed using the Biodiversity Pro 2.0 programme. In addition, in all cases, the similarity matrix of the Bray-Curtis cluster analysis dendrogram was used (single link). The grasshopper distribution in the six different natural zones was determined using ArcGIS ArcMap 10.7.1. The grasshopper's geographical distribution was drawn, based on the Mongolian steppe figure (Mohamed, A and Kimura 2014). The species registered on the European Red List were also divided into categories using IUCN Red List Categories and Criteria (IUCN 2022).

Institutional Abbreviations

(B.-Ulg.) = Bayn-Ulgii

(Zav.) = Zavkhan

(Khuvs.) = Khuvsgul

(A.-khang) = Arkhangai

(Bulg.) = Bulgan

(Orkh.) = Orkhon

(Sel.) = Selenge

(Da.) = Darkhan-Uul

(Khent.) = Khentii

(S.-baat.) = Sukhbaatar

(Dо) = Dornod

(G.-alt.) = Gobi-Altai

(B.-khong.) = Baynkhongor

(U-khang) = Uvurkhangai

(Du.-govi) = Dundgovi

(U.-govi) = Umnugovi

(Do.-govi) = Dornogovi.

(*) = Endemic species of Mongolia

(+) = Geographical distribution of the natural zones

(★) = Indicator species of the geographical natural zones

(-) = poor species

Checklist of Mongolian grasshopper

Acrida kozlovi (Mistshenko, 1951)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1111388

Native status:

Distribution in the natural zone: Desert steppe and desert.

Distribution:

in Mongolia: U.-govi., Uvs. Mistshenko (1968):489, Chogsomzhav (1989):90, Myagmar et al. (2019):56, Batkhuyag and Batnaran (2021):48.

Global distribution: China, Inner Mongolia, Ningxia, Mongolia, Russia (Mistshenko and Bey-Bienko 1951, Chogsomzhav 1989).

Arcyptera (Arcyptera) albogeniculata Ikonnikov, 1911

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1105385

Native status:

Distribution in the natural zone: Forest steppe.

Distribution:

in Mongolia: Tuv., S.-baat., Do., Khovd. Ikonnikov (1911):250, Cejchan and Maran (1966):179, Steinmann (1967):108, Mistshenko (1968):490, Chogsomzhav (1970):127, Günther (1971):114, Altanchimeg and Nonnaizab (2013), Batkhuyag and Batnaran (2021):64.

Global distribution: China, South Korea, Mongolia (Ikonnikov 1911, Storozhenko and Paik 2007).

Arcyptera (Pararcyptera) meridionalis Ikonnikov, 1911

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1105423

Nomenclature:

Arcyptera flavicosta sibirica Uvarov (1914):170.

Native status:

Distribution in the natural zone: Forest steppe, steppe and desert steppe.

Distribution:

in Mongolia: Uvs, Khuvs., Bulg., Tuv, Khent., S.-baat, B.-khong, U-khang., Du.-govi., U.-govi., Sel., Khent., A.-khang., Do. Uvarov (1914):170, Pylnov (1916):278, Chogsomzhav and Shurovenkov (1963):61-63, Cejchan and Maran (1966):180, Mistshenko (1968):490, Chogsomzhav (1968):56, Chogsomzhav (1970):127, Chogsomzhav (1974b):27, Chogsomzhav (1989):91, Storozhenko and Paik (2007):168, Sergeev et al. (2019):21, Batkhuyag and Batnaran (2021):65.

Global distribution: South Korea (Storozhenko et al. 2015), Tuva, S Siberia (from Tuva to Dauria and Yakutia), the southern part of the Russian Far East, Mongolia, NE China, N Korea (Sergeev et al. 2019).

Arcyptera (Pararcyptera) microptera (Fischer von Waldheim, 1833)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1105407

Native status:

Distribution in the natural zone: Forest steppe, steppe and desert steppe.

Distribution:

in Mongolia: Khovd. Chogsomzhav (1972):160, Sergeev (1995):245, Sergeev et al. (2009):108, Altanchimeg (2011):16, Altanchimeg and Nonnaizab (2013), Batnaran et al. (2016):32, Sergeev et al. (2019):21, Batkhuyag and Batnaran (2021):64.

Global distribution: S Europe, W Siberia, S Krasnoyarsk Region, Caucasus, Kazakhstan, NW Mongolia, NW China (Sergeev et al. 2019).

Arcyptera (Arcyptera) fusca (Pallas, 1773)

Species-ID http://orthoptera.speciesfile.org/common/basic/Taxa.aspx?TaxonNameID=47973

Nomenclature:

Gryllus cothurnatus Creutzer (1799):129.

Gryllus (Locusta) nympha Stoll (1813):23.

Arcyoptera stollii Fieber (1853):99.

Gryllus (Locusta) variegatus Sulzer (1776):84.

Gryllus versicolor Gmelin (1789):2082.

Native status:

Distribution in the natural zone: Forest steppe.

Distribution:

in Mongolia: Zav., Khuvs., A.-khang., Sel., Tuv., Khovd, U-khang, Du.-govi., B.-khong. Pylnov (1916):278, Cejchan and Maran (1966):179, Steinmann (1967):109, Mistshenko (1968):490, Chogsomzhav (1968):57, Chogsomzhav (1970):127, Chogsomzhav (1989):91, Batkhuyag (1995):29, Childebaev and Storozhenko (2001), Sergeev et al. (2009):108, Altanchimeg and Nonnaizab (2013), Altanchimeg et al. (2013b):65, Sergeev et al. (2019):20, Popova et al. (2020):599, Batkhuyag and Batnaran (2021):64.

Global distribution: Tuva, southern part of European Russia, S Siberia up to Sakha (Yakutia), Amur Region, mountains of S Europe, Moldova, Ukraine, Caucasus, Kazakhstan, Mongolia, NE China (Sergeev et al. 2019).

Andrea gorochovi Mistshenko, 1989

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1104197

Native status:

Distribution in the natural zone: Desert steppe.

Distribution:

in Mongolia: B.-Khong Gorochov et al. (1989):99,102, Batkhuyag and Batnaran (2021):103.

Chrysochraon dispar (Germar, 1834)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1106359

Native status:

Distribution in the natural zone: High mountain, taiga, forest-steppe, steppe, desert steppe, and desert.

Distribution:

in Mongolia: Sel. Chogsomzhav (1972):157, Chogsomzhav (1989):93, Altanchimeg and Nonnaizab (2013), Altanchimeg et al. (2013b):65, Batkhuyag and Batnaran (2021):57.

Global distribution: Tuva, N Eurasia (except the extreme North), Caucasus, a mountain of Middle Asia, Mongolia (Sergeev et al. 2019).

Euthystira brachyptera (Ocskay, 1826)

Species-ID http://orthoptera.speciesfile.org/common/basic/Taxa.aspx?TaxonNameID=1106449

Native status:

Distribution in the natural zone: High mountain, taiga, forest-steppe, steppe, desert steppe and desert.

Distribution:

in Mongolia: Sel. Chogsomzhav (1969a):77, Chogsomzhav (1972):157, Chogsomzhav (1989):93, Altanchimeg and Nonnaizab (2013), Altanchimeg et al. (2013b):65, Batkhuyag and Batnaran (2021):57.

Global distribution Tuva, N Eurasia (the southern part of the forest zone, the forest-steppe and steppe zones), Mongolia (Sergeev et al. 2019).

Mongolotettix mistshenkoi Chogsomzhav, 1974

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1107086

Native status:

Distribution in the natural zone: Steppe and desert steppe.

Distribution:

in Mongolia. Do.-govi. Chogsomzhav (1974a), Chogsomzhav (1975):42, Altanchimeg (2011):16, Batkhuyag and Batnaran (2021):58.

Mongolotettix japonicus (Bolívar, 1898)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1107087

Native status:

Distribution in the natural zone: Taiga, forest-steppe, steppe and desert steppe.

Distribution:

in Mongolia: Uvs, Khuvs., A.-khang., Bulg., Sel., Tuv., Khent., S.-baat., Do., U-khang., U.-govi. Chogsomzhav (1989):93, Batkhuyag and Batnaran (2021):58.

Global distribution: South Korea, Mongolia (Storozhenko and Paik 2007), Japan (Bolívar 1898), Inner Mongolia (Zheng et al. 2012).

Mongolotettix vittatus (Uvarov, 1914)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1107089

Nomenclature:

Chrysochraon kaszabi Steinmann (1967) :106-120.

Native status:

Distribution in the natural zone: Taiga, forest-steppe, steppe and desert steppe.

Distribution:

in Mongolia: Uvs, Khuvs., A.-khang., Bulg., Sel., Tuv, Khent., S.-baat., Do., U-khang., U.-govi. Pylnov (1916):276, Steinmann (1967):108, Steinmann (1968):240, Steinmann (1971):148, Chogsomzhav (1989):93, Sergeev et al. (2019):17, Batkhuyag and Batnaran (2021):58.

Global distribution: Tuva, S Siberia from Tuva and Krasnoyarsk Region to Dauria, S Amur Region, Mongolia, NE China (Sergeev et al. 2019).

Podismopsis altaica (Zubovski, 1900)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1106371

Native status:

Distribution in the natural zone: High mountain, forest-steppe, steppe and desert steppe.

Distribution:

in Mongolia: Khent. Zubovski (1900):2, Ikonnikov (1911):246, Bey-Bienko (1933):115, Günther (1971):114, Chogsomzhav (1969b):127, Chogsomzhav (1972):158, Chogsomzhav (1989):93, Altanchimeg and Nonnaizab (2013) , Sergeev et al. (2019):18, Batkhuyag and Batnaran (2021):59.

Global distribution: Tuva, Altai-Sayan Mts, E Kazakhstan, N Mongolia (Sergeev et al. 2019).

Podismopsis ussurensis Ikonnikov, 1911

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1106404

Native status:

Distribution in the natural zone: Forest steppe.

Distribution:

in Mongolia: Khent. Chogsomzhav (1972), Storozhenko and Paik (2007):187, Storozhenko et al. (2015):269, Batkhuyag and Batnaran (2021):60.

Global distribution: Korea (HN, PN), Russia (far east), NE China, Mongolia (Storozhenko et al. 2015).

Eremippus mistshenkoi Stebaev, 1965

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1105621

Native status:

Distribution in the natural zone: Steppe.

Distribution:

in Mongolia: Uvs. Chogsomzhav (1969a):77, Chogsomzhav (1969b):127, Chogsomzhav (1972)1:61, Chogsomzhav (1989):92, Sergeev et al. (2019):22, Batkhuyag and Batnaran (2021):67.

Global distribution: Tuva, E Kazakhstan, NW Mongolia (Sergeev et al. 2019).

Eremippus mongolicus Ramme, 1952

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1105622

Nomenclature:

Eremippus kozlov Mistshenko and Bey-Bienko (1951):452.

Native status:

Distribution in the natural zone: Steppe, desert steppe and desert.

Distribution:

in Mongolia: G.-alt., B.-khong., U-khang., U.-govi. Mistshenko and Bey-Bienko (1951):452, Mistshenko (1968):490, Chogsomzhav (1968):59, Chogsomzhav (1989):92, Sergeev et al. (2019):22, Batkhuyag and Batnaran (2021):67.

Global distribution: Tuva, SE Kazakhstan, W Mongolia, NW China (Mistshenko and Bey-Bienko 1951), SE European Russia, Kazakhstan (except the northern part) (Sergeev et al. 2019).

Eremippus simplex (Eversmann, 1859)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1105654

Native status:

Distribution in the natural zone: Steppe, desert steppe and desert.

Distribution:

in Mongolia: Uvs, Khovd, G.-alt. Günther (1971):115, Chogsomzhav (1974b):27, Chogsomzhav (1989):92, Sergeev (1995): 246, Sergeev et al. (2009):108, Popova et al. (2020):600, Batkhuyag and Batnaran (2021):68.

Global distribution: Kazakhstan, Turkmenistan, Uzbekistan, Afghanistan, Kirgizstan, Mongolia (Childebaev and Storozhenko 2001).

Notostaurus albicornis (Eversmann, 1848)

Species-ID http://orthoptera.speciesfile.org/common/basic/Taxa.aspx/common/editTaxon/Distribution/Taxa.aspx?TaxonNameID=1105674

Native status:

Distribution in the natural zone: Steppe, desert steppe and desert.

Distribution:

in Mongolia: Khovd. Günther (1971):114, Chogsomzhav (1989):92, Childebaev and Storozhenko (2001), Sergeev et al. (2009):108, Popova et al. (2020): 599, Batkhuyag and Batnaran (2021):66.

Global distribution: E Europe, Caucasus range, Mongolia, W Siberia, N Iran (Garai 2010).

Dociostaurus (Kazakia) brevicollis (Eversmann, 1848)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1105587

Native status:

Distribution in the natural zone: Forest steppe, steppe, desert steppe and desert.

Distribution:

in Mongolia: Tuv, Khovd. Chogsomzhav (1969b):127, Chogsomzhav (1972):161, Chogsomzhav (1974b):28, Günther (1971):114, Sergeev (1995):245, Childebaev and Storozhenko (2001):30, Sergeev et al. (2009):108, Altanchimeg and Nonnaizab (2013), Altanchimeg et al. (2013b):65, Batkhuyag and Batnaran (2021):66.

Global distribution: Caucasus, Transcaucasia, East Europe, Russia East and South, Kazakhstan, Kirgizstan, Mongolia, Siberia, Iran (Childebaev and Storozhenko 2001).

Dociostaurus (Kazakia) tarbinskyi (Bey-Bienko, 1933)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1105586

Native status:

Distribution in the natural zone: Desert.

Distribution:

in Mongolia: Khovd. Chogsomzhav (1969a):77, Chogsomzhav (1989):92, Childebaev and Storozhenko (2001):30, Batkhuyag and Batnaran (2021):66.

Global distribution: Kazakhstan and Mongolia (Childebaev and Storozhenko 2001).

Eclipophleps bogdanovi Tarbinsky, 1927

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1105104

Native status:

Distribution in the natural zone: High mountain, steppe and desert steppe.

Distribution:

in Mongolia: Altai. Tarbinsky (1927):495, Günther (1971):123, Chogsomzhav (1972):175, Batkhuyag (1995):27, Chogsomzhav (1989):91, Sergeev et al. (2009):108, Altanchimeg (2011):16, Altanchimeg and Nonnaizab (2013), Batnaran et al. (2016):35, Batkhuyag and Batnaran (2021):61.

Eclipophleps carinata Mistshenko, 1968

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1105105

Native status:

Distribution in the natural zone: High mountain, desert steppe and desert.

Distribution:

in Mongolia: B.-khong., G.-alt. Mistshenko (1968):493, Steinmann (1968):245, Sergeev (1995):243, Qian et al. (2021):1310, Batkhuyag and Batnaran (2021):63.

Eclipophleps confinis Mistshenko, 1951

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1105116

Nomenclature:

Oreoptygonotus mongolicus Steinmann (1968):245.

Native status:

Distribution in the natural zone: High mountain, steppe and desert steppe.

Distribution:

in Mongolia: Khovd, G.-alt., B.-khong, U.-govi., U-khang. Günther (1971):123, Mistshenko (1968):495, Chogsomzhav (1972):175, Chogsomzhav (1989):91, Sergeev (1995):243, Sergeev et al. (2009):108, Altanchimeg (2011):16, Batnaran et al. (2016):35, Batkhuyag and Batnaran (2021):63.

Eclipophleps glacialis Bey-Bienko, 1933

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1105106

Native status:

Distribution in the natural zone: High mountain, steppe and desert steppe.

Distribution:

in Mongolia: B.-Ulg., Uvs. Bey-Bienko (1933):115, Chogsomzhav (1969b):128, Chogsomzhav (1989):91, Batkhuyag (1995):27, Sergeev et al. (2009):108, Altanchimeg (2011):16, Sergeev et al. (2019):19, Batkhuyag and Batnaran (2021):62.

Eclipophleps kerzhneri Mistshenko, 1968

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1105108

Native status:

Distribution in the natural zone: High mountain, steppe, desert steppe and desert.

Distribution:

in Mongolia: G.-alt. Mistshenko (1968):493, Batkhuyag (1995):28, Altanchimeg (2011):16, Batkhuyag and Batnaran (2021):62.

Eclipophleps lucida Mistshenko, 1973

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1105110

Native status:

Distribution in the natural zone: High mountain, steppe, desert steppe and desert.

Distribution:

in Mongolia: Uvs, Khovd. Mistshenko (1973), Chogsomzhav (1977):86, Altanchimeg (2011):16, Batkhuyag and Batnaran (2021):62.

Eclipophleps similis Mistshenko, 1951

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1105111

Native status:

Distribution in the natural zone: High mountain, steppe, desert steppe and desert.

Distribution:

in Mongolia: Uvs, Khovd, B.-Ulg., Tuv. Mistshenko and Bey-Bienko (1951):549, Steinmann (1968):243, Batkhuyag (1995):27, Sergeev et al. (2009):108, Altanchimeg (2011):16, Batkhuyag and Batnaran (2021):61.

Eclipophleps tarbinskii Orishchenko, 1960

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1105112

Native status:

Distribution in the natural zone: High mountain, steppe and desert steppe.

Distribution:

in Mongolia: Altai. Orishchenko. (1960), Chogsomzhav (1989):91, Batkhuyag (1995):27, Sergeev et al. (2009):108, Altanchimeg (2011):16, Altanchimeg and Nonnaizab (2013), Batkhuyag and Batnaran (2021):61.

Chorthippus (Altichorthippus) intermedius (Bey-Bienko, 1926)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1106089

Native status:

Distribution in the natural zone: Taiga, forest-steppe and steppe.

Distribution:

in Mongolia: Uvs, Khuvs., Bulg., Sel., Tuv, Khent., S.-baat., Do., U-khang., A.-khang. Tarbinsky (1927):492, Cejchan and Maran (1966):183, Mistshenko (1968):492, Steinmann (1967):116-117, Steinmann (1968):243, Chogsomzhav (1969b):127, Storozhenko and Paik (2007), Altanchimeg et al. (2013b):65, Batkhuyag and Batnaran (2021):87.

Global distribution: Tuva, Altai-Sayan Mts, Sakha (Yakutia), S Russian Far East (including Sakhalin), Mongolia N, NE China, Tibet (Sergeev et al. 2020).

Chorthippus (Chorthippus) albomarginatus (De Geer, 1773)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1105771

Native status:

Distribution in the natural zone: Taiga, forest-steppe, steppe, desert steppe and desert

Distribution:

in Mongolia: Uvs, Zav., Bulg., Khovd. Bey-Bienko (1933):115, Chogsomzhav (1969b):128, Chogsomzhav 1972:173, Günther 1971:123, Chogsomzhav 1989:92, Sergeev 1995:251, Sergeev et al. (2009):109, Altanchimeg and Nonnaizab (2013) Batnaran et al. (2016):14-15, Sergeev et al. (2020):10, Batkhuyag and Batnaran (2021):87.

Global distribution: South Korea (Storozhenko et al. 2015) Tuva, Europe (except the extreme north and the southern parts), W Siberia, N Kazakhstan, N Mongolia (Sergeev et al. 2020).

Chorthippus (Chorthippus) caliginosus Mistshenko, 1951

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1105762

Native status:

Distribution in the natural zone: Forest steppe.

Distribution:

in Mongolia: Sel. Garai (2001):751, Batkhuyag and Batnaran (2021):87.

Global distribution: Southern regions of Transbaikalia, Amur area, Khabarovsk region and south-east of China (Tishechkin and Bukhvalova 2009), Mongolia Garai (2001).

Chorthippus (Chorthippus) dorsatus (Zetterstedt, 1821)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1105795

Native status:

Distribution in the natural zone: Forest steppe, steppe and desert steppe.

Distribution:

in Mongolia: Uvs, S.-baat. Chogsomzhav (1969b):128, Chogsomzhav (1972):173, Sergeev et al. (2020):8, Batkhuyag and Batnaran (2021):86.

Global distribution: Tuva, Transbaikalia, Mongolia, NE China (Sergeev et al. 2020).

Chorthippus (Chorthippus) hammarstroemi (Miram, 1907)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1105913

Native status:

Distribution in the natural zone: Taiga, forest-steppe, steppe, desert steppe and desert.

Distribution:

in Mongolia: Bulg., A.-khang., Sel., Tuv, S.-baat., Khuvs. Cejchan and Maran (1966):183, Chogsomzhav (1971):80, Sergeev (1995):249, Storozhenko and Paik (2007):173, Storozhenko et al. (2015):242, Altanchimeg and Nonnaizab (2013), Altanchimeg et al. (2013b):65, Batnaran et al. (2016):34, Sergeev et al. (2020):7, Batkhuyag and Batnaran (2021):83.

Global distribution: South Korea (Storozhenko et al. 2015), Tuva, Altai-Sayan Mts, Transbaikalia, S Sakha (Yakutia), S Russian Far East, Mongolia, N, NE China (Sergeev et al. 2020).

Chorthippus (Chorthippus) dichrous (Eversmann, 1859)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1105796

Nomenclature:

Chorthippus dorsatus australia Predtechenskii (1928):89.

Native status:

Distribution in the natural zone: High mountain, forest-steppe, steppe, desert steppe and desert.

Distribution:

in Mongolia: B.-Ulg., S.-baat., Do., B.-khong., Sel., Khovd. Pylnov (1916):278, Chogsomzhav (1971):82, Chogsomzhav (1972):173, Sergeev (1995):251, Childebaev and Storozhenko (2001), Sergeev et al. (2009):109, Batnaran et al. (2016):33, Sergeev et al. (2020):9, Batkhuyag and Batnaran (2021):86.

Global distribution: Tuva, S, SE Europe, S Siberia (up to Tuva), Asia Minor, Caucasus, Iran, Kazakhstan, Tien Shan, Pamiro-Alay, NW China, Mongolia (Sergeev et al. 2020).

Chorthippus (Chorthippus) fallax (Zubovski, 1900)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1105903

Native status:

Distribution in the natural zone: Taiga, forest-steppe, steppe and desert steppe.

Distribution:

in Mongolia: B.-Ulg., Uvs, Zav., Khuvs., A.-khang., Bulg., Sel., Tuv, Khent., S.-baat., Da., G.-alt., B.-khong. Ikonnikov (1911):253, Pylnov (1916):278, Bey-Bienko (1933):115, Cejchan and Maran (1966):183, Mistshenko (1968):492, Günther (1971):122, Chogsomzhav (1968):57, Chogsomzhav (1969b):128, Chogsomzhav (1972):172, Munkhbat (2010):16, Altanchimeg and Nonnaizab (2013), Altanchimeg et al. (2013b):65, Storozhenko et al. (2015):244, Sergeev et al. (2020):85.

Global distribution: Tuva, Siberia (except the western part of West Siberian Plain and the extreme north), S Russian Far East, E Kazakhstan, N Mongolia, N, NE China, South Korea (Storozhenko et al. 2015, Sergeev et al. 2020).

Chorthippus (Chorthippus) turanicus Tarbinsky, 1925

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1105894

Native status:

Distribution in the natural zone: Steppe.

Distribution:

in Mongolia: G.-alt. Chogsomzhav (1974b):29, Chogsomzhav (1989):92, Sergeev et al. (2009):109, Batkhuyag and Batnaran (2021):84.

Global distribution: China, Xinjiang (Tarbinsky 1925), Turkestan, Kazakhstan, Tadzhikistan, Uzbekistan (Childebaev and Storozhenko 2001), Mongolia (Sergeev et al. 2009).

Chorthippus (Chorthippus) ilkazi Uvarov, 1934

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1105936

Native status:

Distribution in the natural zone: Steppe and desert steppe.

Distribution:

in Mongolia: Tuv, S.-baat., Do. Steinmann (1967):116, Altanchimeg et al. (2013b):65.

Global distribution: Western Asia, Mongolia, Turkey (Uvarov 1934).

Chorthippus (Glyptobothrus) apricarius (Linnaeus, 1758)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1105949

Native status:

Distribution in the natural zone: Forest steppe and steppe.

Distribution:

in Mongolia: Uvs, Bulg., Sel., Tuv, Khent., U-khang. Pylnov (1916):278, Steinmann (1967):114, Chogsomzhav (1968):57-58, Chogsomzhav (1969b):127, Chogsomzhav (1989):92, Childebaev and Storozhenko (2001):34, Sergeev et al. (2009):108, Altanchimeg and Nonnaizab (2013), Sergeev et al. (2020):6, Batkhuyag and Batnaran (2021):82.

Global distribution: Tuva, Europe (except the extreme north), S Siberia, Asia Minor, Kazakhstan, NW, N, NE China, Mongolia (Sergeev et al. 2020).

Chorthippus (Glyptobothrus) biguttulus (Linnaeus, 1758)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1106017

Native status:

Distribution in the natural zone: Taiga, forest-steppe, steppe and desert steppe.

Distribution:

in Mongolia: Uvs, Bulg., Sel., Tuv, Khent., U-khang. Pylnov (1916):278, Bey-Bienko (1933):115, Steinmann (1967):115, Mistshenko (1968):492, Steinmann (1968):243, Chogsomzhav (1969b):127, Chogsomzhav (1974b):28, Sergeev (1995):248, Sergeev et al. (2009):108, Altanchimeg and Nonnaizab (2013), Myagmar et al. (2019):56, Batkhuyag and Batnaran (2021):79.

Global distribution: France, Switzerland, Yugoslavia (Miksic 1981), Mongolia (Steinmann 1968), Inner Mongolia (Li et al. 2007).

Chorthippus (Glyptobothrus) brunneus (Thunberg, 1815)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1106050

Native status:

Distribution in the natural zone: Taiga, forest-steppe, steppe and desert steppe.

Distribution:

in Mongolia: B.-Ulg., Uvs, Khuvs., Bulg., Tuv, S.-baat., Do., Khovd., B.-khong., Du.-govi. Bolívar (1901):226, Bey-Bienko (1933):115, Steinmann (1968):243, Chogsomzhav (1970), Chogsomzhav (1989):92, Childebaev and Storozhenko (2001), Altanchimeg et al. (2013b):65, Batnaran et al. (2016):33, Sergeev et al. (2019):34, Batkhuyag and Batnaran (2021):78.

Global distribution: Tuva, S Russia up to Tuva, N Kazakhstan, N Mongolia (Benediktov 1999, Sergeev et al. 2019).

Chorthippus (Glyptobothrus) maritimus Mistshenko, 1951

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1106041

Native status:

Distribution in the natural zone: High mountain and desert steppe.

Distribution:

in Mongolia: Altanchimeg and Nonnaizab (2013):81.

Global distribution: Caucasus, Transcaucasus, East Europe, Krym and China (Ramme 1939), Mongolia (Altanchimeg and Nonnaizab 2013).

Chorthippus (Glyptobothrus) dubius (Zubovski, 1898)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1105932

Native status:

Distribution in the natural zone: Forest steppe, steppe and desert steppe.

Distribution:

in Mongolia: B.-Ulg., Uvs, Zav., Khuvs., A.-khang., Bulg., Sel., Tuv, Khent., Do., G.-alt., B.-khong., U-khang., U.-govi. Bolívar (1901):226, 231, Bey-Bienko (1933):115, Steinmann (1967):116, Mistshenko (1968):492, Chogsomzhav (1968):58, Chogsomzhav (1969b):128, Chogsomzhav (1972):171, Günther (1971):121, Sergeev (1995):248, Childebaev and Storozhenko (2001), Altanchimeg and Nonnaizab (2013), Altanchimeg et al. (2013b):65, Batnaran et al. (2016):34, Sergeev et al. (2019):38, Batkhuyag and Batnaran (2021):81.

Global distribution: Tuva, SE European Russia, S Siberia, Kazakhstan, Mongolia, NE, N, C China (Sergeev et al. 2019).

Chorthippus (Glyptobothrus) mollis (Charpentier, 1825)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1106033

Native status:

Distribution in the natural zone: Desert steppe and desert.

Distribution:

in Mongolia: Altanchimeg and Nonnaizab (2013):81, Batkhuyag and Batnaran (2021):80.

Global distribution: Tuva, Europe (except the extreme north), Siberia (except the extreme north), Asia Minor, Caucasus, Kazakhstan, Middle Asia, N Iran (Sergeev et al. 2019), Mongolia (Altanchimeg and Nonnaizab 2013).

Chorthippus (Glyptobothrus) vagans (Eversmann, 1848)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1105971

Nomenclature:

Gomphocerus subsinuatus Fischer (1849):42.

Native status:

Distribution in the natural zone: Forest steppe.

Distribution:

in Mongolia: Altanchimeg and Nonnaizab (2013):81, Batkhuyag and Batnaran (2021):87.

Global distribution: from West Europe far into temperate Asia (Willemse 2009), Mongolia (Altanchimeg and Nonnaizab 2013).

Pseudochorthippus montanus (Charpentier, 1825)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1106350

Nomenclature:

Stenobothrus (Chorthippus) longicornis Jacobson and Bianchi. (1905):182,234.

Chorthippus longicornis Jacobson and Bianchi. (1905):182.

Native status:

Distribution in the natural zone: Taiga, forest-steppe, steppe, desert steppe and desert.

Distribution:

in Mongolia: B.-Ulg., Uvs, Zav., A.-khang., Bulg., Tuv, B.-khong., Khovd. Mistshenko (1968):492, Chogsomzhav (1969b):127, Chogsomzhav (1972):173, Günther (1971):122, Sergeev (1995):251, Childebaev and Storozhenko (2001), Sergeev et al. (2009):109, Altanchimeg et al. (2013b):65, Storozhenko et al. (2015):246, Sergeev et al. (2020):15, Batkhuyag and Batnaran (2021):85.

Global distribution: South Korea (Storozhenko et al. 2015), Tuva, N, C Europe, Siberia, Russian Far East, N Kazakhstan, Mongolia, NE China, N Korea (Sergeev et al. 2020).

Pseudochorthippus parallelus (Zetterstedt, 1821)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1106327

Native status:

Distribution in the natural zone: Steppe.

Distribution:

in Mongolia: Uvs. Chogsomzhav (1977), Batkhuyag and Batnaran (2021):85.

Global distribution: Tuva, Europe (except the extreme north), Siberia (except the north, but including the central parts of Sakha (Yakutia) and the southern parts of Krasnoyarsk Region and the Republic of Khakassia (Ivanova 1967), Asia Minor, Caucasus, Kazakhstan, Tien Shan, Mongolia [including the Mongolian part of Uvs-Nuur Basin (Chogsomzhav 1977), NW China (Sergeev et al. 2020).

Aeropedellus baliolus Mistshenko, 1951

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1106225

Native status:

Distribution in the natural zone: Steppe.

Distribution:

in Mongolia: Tuv, S.-baat., U-khang. Steinmann (1967):114, Altanchimeg et al. (2014):133, Popova et al. (2020):602.

Global distribution: Kazakhstan and Mongolia (Altanchimeg et al. 2014).

Aeropedellus chogsomjavi Altanchimeg, Chen & Nonnaitzb, 2014

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1220729

Native status:

Distribution in the natural zone: Steppe.

Distribution:

in Mongolia: Khuvs. Altanchimeg and Nonnaizab (2013):81, Altanchimeg et al. (2014):133, Batkhuyag and Batnaran (2021):76.

Aeropedellus reuteri (Miram, 1907)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1106234

Native status:

Distribution in the natural zone: Steppe and desert steppe.

Distribution:

in Mongolia: Khent., U-khang. Steinmann (1967):114, Altanchimeg et al. (2014):133, Sergeev et al. (2020):4, Batkhuyag and Batnaran (2021):76.

Global distribution: Mongolia, Type localities and Khakassia (Sergeev et al. 2020).

Aeropedellus variegatus (Fischer von Waldheim, 1846)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1106239

Native status:

Distribution in the natural zone: Forest steppe and steppe.

Distribution:

in Mongolia: B.-Ulg, Uvs, Zav., Khuvs., A.-khang, Bulg., Tuv., Khent., Khovd, G.-alt., B.-khong., U-khang. Cejchan and Maran (1966):182, Steinmann (1968):242, Steinmann (1967):113, Chogsomzhav (1969b):127, Günther (1971):120,Sergeev (1995):247, Sergeev et al. (2009):108, Altanchimeg (2011):16, Altanchimeg and Nonnaizab (2013), Altanchimeg et al. (2013b):65, Altanchimeg et al. (2014):133, Sergeev et al. (2020):4, Batkhuyag and Batnaran (2021):75.

Global distribution: Tuva, N Caucasus, NE European Russia, Siberia, N Europe, mountains of S Europe, E Kazakhstan, Mongolia (Sergeev et al. 2020).

Gomphocerus sibiricus (Linnaeus, 1767)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1106148&Next=Taxa.aspx

Native status:

Distribution in the natural zone: Forest steppe and steppe.

Distribution:

in Mongolia: Uvs, Zav., Khuvs., A.-khang., Bulg., Tuv, Khent., Khovd, G.-alt., U-khang., U.-govi., Cejchan and Maran (1966):180, Günther (1971):119, Mistshenko (1968):492, Chogsomzhav (1989):91, Sergeev et al. (2009):108, Altanchimeg and Nonnaizab (2013), Altanchimeg et al. (2013b):65, Batkhuyag and Batnaran (2021):74.

Global distribution: Tuva, N, NE Europe, Siberia (except the extreme north), N Kazakhstan, N Mongolia, NE China (Sergeev et al. 2020).

Gomphocerippus rufus (Linnaeus, 1758)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1106254

Nomenclature:

Acrydium clavicorne De Geer (1773):482.

Native status:

Distribution in the natural zone: Forest steppe and steppe.

Distribution:

in Mongolia: Tuv. Childebaev and Storozhenko (2001), Munkhbat (2010):168, Batkhuyag and Batnaran (2021):73.

Global distribution: Mongolia (Childebaev and Storozhenko 2001), Tuva, Europe (except the extreme north), Siberia (except the extreme north and NE parts), Amur Region, N Caucasus, W Kazakhstan, NE China (Sergeev et al. 2020).

Myrmeleotettix palpalis (Zubovski, 1900)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1106193

Native status:

Distribution in the natural zone: Forest steppe, steppe, desert steppe and desert.

Distribution:

in Mongolia: B.-Ulg., Uvs, Zav., Khuvs., Khovd, G.-alt., Sel., Tuv., Khent., A.-khang., U-khang., U.-govi. Zubovski (1900):13, Pylnov (1916):278, Bey-Bienko (1933):118, Cejchan and Maran (1966):180, Chogsomzhav (1968):56-58, Chogsomzhav (1969b):127, Chogsomzhav (1974b):28, Chogsomzhav (1989):91, Childebaev and Storozhenko (2001), Garai (2001):751, Sergeev et al. (2009):108, Altanchimeg (2011):16, Altanchimeg and Nonnaizab (2013), Altanchimeg et al. (2013b):65, Sergeev et al. (2019):29, Batkhuyag and Batnaran (2021):72.

Global distribution: Tuva, S Siberia (from the Altai Mts. to Dauria), Amur Region, E Kazakhstan, Mongolia, China (Sergeev et al. 2019).

Myrmeleotettix zaitzevi Mistshenko, 1968

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1106195

Native status:

Distribution in the natural zone: Desert steppe.

Distribution:

in Mongolia: Du.-govi., Tuv. Mistshenko (1968):490, Chogsomzhav (1989):91, Munkhbat (2010):168, Batkhuyag and Batnaran (2021):72.

Stauroderus scalaris (Fischer von Waldheim, 1846)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1106274

Native status:

Distribution in the natural zone: Forest steppe and steppe.

Distribution:

in Mongolia: Uvs, Bulg., Tuv. Pylnov (1916):278, Cejchan and Maran (1966):184, Steinmann (1964):382, Günther (1971):121, Chogsomzhav (1969b):127, Chogsomzhav (1972):168, Chogsomzhav (1989):92, Sergeev et al. (2009):108, Altanchimeg and Nonnaizab (2013), Altanchimeg et al. (2013b):65, Batnaran et al. (2016):35, Sergeev et al. (2019):32, Batkhuyag and Batnaran (2021):77.

Global distribution: Tuva, Europe (except the north), S Siberia (up to Buryatia), Asia Minor, Caucasus, Kazakhstan, Tien Shan, Pamiro-Alay, NW China, Mongolia, NW Iran (Sergeev et al. 2019).

Schmidtiacris schmidti (Ikonnikov, 1913)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1114808

Nomenclature:

Chorthippus nakazimai Furukawa et al. (1950):30.

Native status:

Distribution in the natural zone: Forest steppe.

Distribution:

in Mongolia: Bulg., Tuv. Mistshenko (1968) :492, Günther (1971):122, Altanchimeg et al. (2013b):65, Sergeev et al. (2019):39, Batkhuyag and Batnaran (2021):83.

Global distribution: Tuva, Transbaikalia, S Russian Far East, Mongolia, NE China, Korea, Japan (Sergeev et al. 2019).

Mesasippus kozhevnikovi (Tarbinsky, 1925)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1106179

Native status:

Distribution in the natural zone: Desert steppe and desert.

Distribution:

in Mongolia: B.-Ulg., Uvs, Zav., Khuvs., Tuv, A.-khang., Bulg., Tuv., Sel., Khent., S.-baat., Khovd, U-khang. Mistshenko and Bey-Bienko (1951):501, Mistshenko (1968):492, Steinmann (1968):240, 242, Chogsomzhav (1972):168, Chogsomzhav (1974b):28, Chogsomzhav (1989):91, Gorochov et al. (1989):104, Childebaev and Storozhenko (2001), Batkhuyag and Batnaran (2021):77.

Global distribution: China, Xinjiang, Mongolia, Kazakhstan (Mistshenko and Bey-Bienko 1951, Childebaev and Storozhenko 2001).

Dasyhippus barbipes (Fischer von Waldheim, 1846)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1106266

Native status:

Distribution in the natural zone: Steppe, desert steppe and desert.

Distribution:

in Mongolia: Khuvs., Sel., Tuv, S.-baat., Do., Khovd, Du.-govi. Günther (1971):120, Steinmann (1967):113, Pylnov (1916):278, Tarbinsky (1931):140, Chogsomzhav (1968):58, Chogsomzhav (1971):71, Chogsomzhav (1972):166, Chogsomzhav (1989):91, Altanchimeg and Nonnaizab (2013), Altanchimeg et al. (2013b):65, Batnaran et al. (2016):34, Sergeev et al. (2020):5, Batkhuyag and Batnaran (2021):75.

Global distribution: Tuva, SE Altai, Transbaikalia, Mongolia, N China (Sergeev et al. 2020).

Egnatioides desertus Uvarov, 1926

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1109695

Native status:

Distribution in the natural zone: Desert.

Distribution:

in Mongolia: Khovd, Altai Transaltai gobi. Günther (1971):113, Chogsomzhav (1989):90, Batkhuyag and Batnaran (2021):47.

Global distribution: Mongolia (Günther 1971), Kazakhstan and Turkestan (Childebaev and Storozhenko 2001).

Omocestus (Omocestus) haemorrhoidalis (Charpentier, 1825)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1107287

Native status:

Distribution in the natural zone: Forest steppe and steppe.

Distribution:

in Mongolia: Zav., Bulg., Tuv, Khent., S.-baat., Khovd, U-khang., Sel., Uvs, A.-khang., Khuvs. Pylnov (1916):277, Bey-Bienko (1933):118, Cejchan and Maran (1966):180, Steinmann (1967):110, Steinmann (1968):241, Chogsomzhav (1968):58, Chogsomzhav (1969b):127, Chogsomzhav (1974b):28, Sergeev (1995):246, Sergeev et al. (2009):108, Altanchimeg and Nonnaizab (2013), Altanchimeg et al. (2013b):65, Storozhenko et al. (2015):254, Sergeev et al. (2019):26, Batkhuyag and Batnaran (2021):71.

Global distribution: Tuva, Europe (except the extreme north), Siberia (except the extreme north), S Russian Far East, Asia Minor, Caucasus, Kazakhstan, Tien Shan, Pamiro-Alay, Mongolia, N China, South Korea (Sergeev et al. 2019).

Omocestus (Omocestus) petraeus (Brisout de Barneville, 1856)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1107268

Native status:

Distribution in the natural zone: Steppe.

Distribution:

in Mongolia: Uvs, Bulg., Tuv, Khent., U-khang. Steinmann (1964):382, Steinmann (1967):110, Chogsomzhav (1969b):127, Chogsomzhav (1971):66, Chogsomzhav (1972):164, Batnaran et al. (2016):35, Sergeev et al. (2020):30, 32, Batkhuyag and Batnaran (2021):71.

Global distribution: Tuva, S Europe, S Siberia (up to S Krasnoyarsk Region), Asia Minor, Caucasus, N Kazakhstan (Sergeev et al. 2019), Mongolia (Sergeev et al. 2020).

Omocestus (Omocestus) rufipes (Zetterstedt, 1821)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1107273

Native status:

Distribution in the natural zone: Steppe.

Distribution:

in Mongolia: Chogsomzhav (1969b):91, Childebaev and Storozhenko (2001):32, Altanchimeg and Nonnaizab (2013).

Global distribution: Russian C, E, S, Kazakhstan, Siberia, Mongolia (Childebaev and Storozhenko 2001).

Omocestus (Omocestus) tzendsureni Günther, 1971

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1107282

Native status:

Distribution in the natural zone: Forest steppe, steppe and desert steppe.

Distribution:

in Mongolia: Khovd, G.-alt. Günther (1971):116, Chogsomzhav (1989):91, Sergeev (1995):246, Batkhuyag and Batnaran (2021):71.

Global distribution: China, Xinjiang, Mongolia (Günther 1971).

Omocestus (Omocestus) viridulus (Linnaeus, 1758)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1107300

Native status:

Distribution in the natural zone: Forest steppe, steppe, desert steppe and desert.

Distribution:

in Mongolia: Khuvs., Bulg., Sel., S.-baat., Do., Khovd, G.-alt., Uvs. Steinmann (1964):382, Steinmann (1967):110, Steinmann (1968):241, Chogsomzhav (1968):57, Chogsomzhav (1969b):127, Chogsomzhav (1972):163, Chogsomzhav (1974b):28, Günther (1971):116, Chogsomzhav (1989):91, Sergeev (1995):108, Childebaev and Storozhenko (2001), Sergeev et al. (2009):108, Altanchimeg and Nonnaizab (2013), Altanchimeg et al. (2013b):65, Batnaran et al. (2016):36, Sergeev et al. (2019):26, Batkhuyag and Batnaran (2021):70.

Global distribution: Tuva, Europe (except the extreme north), S Siberia, Amur Region, S Khabarovsk Region, Asia Minor, Caucasus, Kazakhstan, Tien Shan, N China, Mongolia, N Korea (Sergeev et al. 2019).

Stenobothrus carbonarius (Eversmann, 1848)

Species-ID http://orthoptera.speciesfile.org/common/basic/Taxa.aspx/common/Links/Taxa.aspx?TaxonNameID=1107430

Native status:

Distribution in the natural zone: Forest steppe.

Distribution:

in Mongolia: Sel. Sergeev (1995):246, Childebaev and Storozhenko (2001):31, Batnaran (2008):47, Batnaran et al. (2016):36, Popova et al. (2020):600, Batkhuyag and Batnaran (2021):69.

Global distribution: Mongolia (Childebaev and Storozhenko 2001), Tuva, SE European Russia, S Siberia (up to Buryatia), Kazakhstan (Sergeev et al. 2019).

Stenobothrus newskii Zubovski, 1900

Species-ID http://orthoptera.speciesfile.org/common/basic/Taxa.aspx/common/Links/Taxa.aspx?TaxonNameID=1107354

Native status:

Distribution in the natural zone: Taiga and forest-steppe.

Distribution:

in Mongolia: G.-alt. Zubovski (1900):9, Bey-Bienko (1926):10, Chogsomzhav (1972):162, Sergeev et al. (2009):108, Altanchimeg (2011):16, Sergeev et al. (2019):24, Batkhuyag and Batnaran (2021):70.

Global distribution: Tuva, Altai Mts. (including S Altai), NW Mongolia (Sergeev et al. 2019).

Stenobothrus eurasius Zubovski, 1898

Species-ID http://orthoptera.speciesfile.org/common/basic/Taxa.aspx/common/Links/Taxa.aspx?TaxonNameID=1107386

Native status:

Distribution in the natural zone: Forest steppe and steppe.

Distribution:

in Mongolia: Sel., Tuv. Chogsomzhav (1971):64, Chogsomzhav (1989):91,Sergeev (1995):246, Munkhbat (2010):168, Altanchimeg and Nonnaizab (2013), Altanchimeg et al. (2013b):65, Sergeev et al. (2019):24, Batkhuyag and Batnaran (2021):70.

Global distribution: Tuva, SE European Russia, S Siberia, N Kazakhstan, Tien Shan, N Mongolia (Sergeev et al. 2019).

Stenobothrus fischeri (Eversmann, 1848)

Species-ID http://orthoptera.speciesfile.org/common/basic/Taxa.aspx/common/Links/Taxa.aspx?TaxonNameID=1107392

Native status:

Distribution in the natural zone: Desert steppe and desert.

Distribution:

in Mongolia: Bulg., Sel., Khovd. Pylnov (1916):277, Günther (1971):116, Chogsomzhav (1972):163, Childebaev and Storozhenko (2001), Sergeev et al. (2009):108, Altanchimeg et al. (2013b):65, Sergeev et al. (2019):23, Batkhuyag and Batnaran (2021):68.

Global distribution: Tuva, S Europe, S Siberia (up to Tuva), Asia Minor, Caucasus, Kazakhstan, a mountain of Middle Asia, Mongolia (Sergeev et al. 2019).

Stenobothrus lineatus (Panzer, 1796)

Species-ID http://orthoptera.speciesfile.org/common/basic/Taxa.aspx/common/Links/Taxa.aspx?TaxonNameID=1107404

Native status:

Distribution in the natural zone: Forest steppe and steppe.

Distribution:

in Mongolia: Zav., Bulg., Tuv, Khent., S.-baat.,. Cejchan and Maran (1966):180, Steinmann (1967):110, Mistshenko (1968):490, Chogsomzhav (1969b):127, Chogsomzhav (1989):91, Sergeev (1995):246, Childebaev and Storozhenko (2001), Sergeev et al. (2019):23, Batkhuyag and Batnaran (2021):68.

Global distribution: Tuva, Europe (except the northern part), S Siberia up to Sakha (Yakutia), Caucasus, N Kazakhstan, N Mongolia, Russian Far East, Dauria (ssp. flavotobialis) (Storozhenko 1985, Sergeev et al. 2019).

Megaulacobothrus aethalinus (Tarbinsky, 1927)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1106289

Native status:

Distribution in the natural zone: Forest steppe.

Distribution:

in Mongolia: Bogdkhan Mountain, Ulaanbaatar, Altanchimeg et al. (2013a).

Global distribution: China (Li et al. 2007), Mongolia (Altanchimeg et al. 2013a).

Bryodema gebleri (Fischer von Waldheim, 1836)

Species-ID http://orthoptera.speciesfile.org/common/basic/taxa.aspx/common/editimgsnd/Taxa.aspx?TaxonNameID=1104225

Native status:

Distribution in the natural zone: Steppe, desert steppe and desert.

Distribution:

in Mongolia: Uvs. Bey-Bienko (1933):119, Chogsomzhav (1969b):128, Chogsomzhav (1972):180, Sergeev (1995):253, Sergeev et al. (2009):109, Altanchimeg et al. (2015):69, Batkhuyag et al. (2019):107, Sergeev et al. (2020):24, 25, Batkhuyag and Batnaran (2021):101, Dey et al. (2021):335.

Global distribution: Tuva, S Ural Mts, Altai-Sayan Mts (except SE Altai), Transbaikalia, Kazakhstan, Tien Shan (except the eastern part), NW China, W Mongolia (Sergeev et al. 2020).

Bryodema heptapotanicum Bey-Bienko, 1930

Species-ID http://orthoptera.speciesfile.org/common/basic/taxa.aspx/common/editimgsnd/Taxa.aspx?TaxonNameID=1104216

Native status:

Distribution in the natural zone: Desert steppe.

Distribution:

in Mongolia: G.-alt. Altanchimeg and Nonnaizab (2013):81, Batkhuyag and Batnaran (2021):101.

Global distribution: China, Xinjiang (Bey-Bienko 1930), Kazakhstan (Childebaev and Storozhenko 2001), Mongolia (Altanchimeg and Nonnaizab 2013).

Bryodema kozlovi Bey-Bienko, 1930

Species-ID http://orthoptera.speciesfile.org/common/basic/taxa.aspx/common/editimgsnd/Taxa.aspx?TaxonNameID=1104218

Native status:

Distribution in the natural zone: Desert steppe.

Distribution:

in Mongolia: Bey-Bienko (1930):101.

Global distribution: Mongolia (Bey-Bienko 1930), China, Inner Mongolia (Alashan) (Zheng et al. 2012).

Bryodema luctuosum (Stoll, 1813)

Species-ID http://orthoptera.speciesfile.org/common/basic/taxa.aspx/common/editimgsnd/Taxa.aspx?TaxonNameID=1104229

Native status:

Distribution in the natural zone: Steppe, desert steppe and desert.

Distribution:

in Mongolia: Uvs, Zav., Khuvs., A.-khang., Sel., Tuv, Khent., S.-baat., B.-khong., Du.-govi., B.-khong., U-khang. Bolívar (1901):226, 233, Pylnov (1916):279, Bey-Bienko (1930):113, Cejchan and Maran (1966):185, Steinmann (1967):118, Steinmann (1968):247, Mistshenko (1968):495, Chogsomzhav (1968):57, Chogsomzhav (1969b):128, Chogsomzhav (1972):182, Sergeev (1995):254, Altanchimeg and Nonnaizab (2013), Batnaran et al. (2016):37, Batkhuyag and Batnaran (2021):101, Dey et al. (2021):336.

Global distribution: Siberia, China and Mongolia (Benediktov 2016).

Bryodema miramae Bey-Bienko, 1930

Species-ID http://orthoptera.speciesfile.org/common/basic/taxa.aspx/common/editimgsnd/Taxa.aspx?TaxonNameID=1104236

Native status:

Distribution in the natural zone: Desert steppe.

Distribution:

in Mongolia: Altanchimeg and Nonnaizab (2013):81, Batkhuyag and Batnaran (2021):101.

Global distribution: Mongolia (Altanchimeg and Nonnaizab 2013), China (Zhang et al. 2006).

Bryodema nigripennis Mistshenko & Gorochov, 1989

Species-ID http://orthoptera.speciesfile.org/common/basic/taxa.aspx/common/editimgsnd/Taxa.aspx?TaxonNameID=1104220

Native status:

Distribution in the natural zone: Desert steppe.

Distribution:

in Mongolia. B.-khong. Mistshenko and Gorochov (1989):99, Batkhuyag and Batnaran (2021):101.

Global distribution: Mongolia (Mistshenko and Gorochov 1989), China (Zhang et al. 2006).

Compsorhipis bryodemoides Bey-Bienko, 1932

Species-ID http://orthoptera.speciesfile.org/common/basic/taxa.aspx/common/edittaxon/distribution/Taxa.aspx?TaxonNameID=1104250

Native status:

Distribution in the natural zone: Desert steppe and desert.

Distribution:

in Mongolia: Uvs, Bulg., S.-baat., Khovd, G.-alt., B.-khong., U-khang., B.-khong., Du.-govi., U.-govi. Bey-Bienko (1932):84, :606, Cejchan and Maran (1966):186, Mistshenko (1968):495, Chogsomzhav (1968):59, Chogsomzhav (1972):185, Chogsomzhav (1989):94, Sergeev (1995):254, Batnaran et al. (2016):38, Batkhuyag et al. (2019):107, Myagmar et al. (2019):56, Batkhuyag and Batnaran (2021):102, Dey et al. (2021):339.

Compsorhipis davidiana (Saussure, 1888)

Species-ID http://orthoptera.speciesfile.org/common/basic/taxa.aspx/common/edittaxon/distribution/Taxa.aspx?TaxonNameID=1104251

Native status:

Distribution in the natural zone: Desert steppe and desert.

Distribution:

in Mongolia: U.-govi., Uvs, Khovd, B.-khong., Du.-govi. Bolívar (1901):226, 235, Chogsomzhav (1969b):128, Chogsomzhav (1972):184, Chogsomzhav (1989):94, Batkhuyag et al. (2019):107, Sergeev et al. (2020):28, Batkhuyag and Batnaran (2021):102.

Global distribution: Tuva, S Transbaikalia, Mongolia, NW, N China (Sergeev et al. 2020).

Compsorhipis orientalis Chogsomzhav, 1989

Species-ID http://orthoptera.speciesfile.org/common/basic/taxa.aspx/common/edittaxon/distribution/Taxa.aspx?TaxonNameID=1104253

Native status:

Distribution in the natural zone: Steppe.

Distribution:

in Mongolia: Do.-govi. Chogsomzhav (1989):94, Batkhuyag and Batnaran (2021):103, Dey et al. (2021):340.

Global distribution: China (Yin and Wang 2005), Mongolia (Dey et al. 2021).

Bryodemella (Bryodemella) holdereri (Krauss, 1901)

Species-ID http://orthoptera.speciesfile.org/common/basic/taxa.aspx/common/specimen/SpecimensByTaxon.aspx?TaxonNameID=1104267

Nomenclature:

Bryodema occidentale Bey-Bienko (1930):87.

Native status:

Distribution in the natural zone: Forest steppe, steppe, desert steppe and desert.

Distribution:

in Mongolia: Uvs, Zav., Khuvs., A.-khang., Bulg., Sel., Tuv., Khovd, S.-baat., Do., Khovd, G.-alt., B.-khong., U-khang., Du.-govi. Bey-Bienko (1930):85, Chogsomzhav and Shurovenkov (1963):61, Günther (1971):124, Steinmann (1967):119, Steinmann (1968):246, Mistshenko (1968):495, Chogsomzhav (1968):57, Chogsomzhav (1969b):128, Chogsomzhav (1972):179, Sergeev (1995):253, Altanchimeg and Nonnaizab (2005):234, Sergeev et al. (2009):109, Altanchimeg and Nonnaizab (2013), Altanchimeg et al. (2013b):65, Batnaran et al. (2016):37, Sergeev et al. (2020):21, Batkhuyag and Batnaran (2021):99, Dey et al. (2021):337.

Global distribution: Tuva, SE Altai, Khakassia, S Krasnoyarsk Region, Transbaikalia, Mongolia, N, NE China (Sergeev et al. 2020).

Bryodemella (Bryodemella) tuberculata (Fabricius, 1775)

Species-ID http://orthoptera.speciesfile.org/common/basic/taxa.aspx/common/specimen/Taxa.aspx?TaxonNameID=1104270

Native status:

Distribution in the natural zone: Taiga, forest-steppe, steppe and desert steppe.

Distribution:

in Mongolia: B.-Ulg., Uvs, Zav., Khuvs., A.-khang., Bulg., Sel., Tuv, Khent., S.-baat., B.-khong., U-khang., U.-govi. Bolívar (1901):226, Uvarov (1914):171, Pylnov (1916):279, Bey-Bienko (1930):91, Cejchan and Maran (1966):184, Steinmann (1967):118, Steinmann (1968):247, Mistshenko (1968):495, Chogsomzhav (1968):57, Chogsomzhav (1969b):128, Chogsomzhav (1972):179, Batkhuyag and Batnaran (2021):99, Dey et al. (2021):339.

Global distribution: Tuva, Europe (except the extreme north and the southern regions of W Europe), Siberia and Far East (northwards to Magadan Region), Kazakhstan, Mongolia, Korea, China, Tibet, Himalayas (Sergeev et al. 2020).

Bryodemella (Marikovskiella) orientalis (Bey-Bienko, 1930)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1104280

Native status:

Distribution in the natural zone: Steppe, desert steppe and desert.

Distribution:

in Mongolia: Uvs, Zav., Khovd, G.-alt., B.-khong., U-khang., Du.-govi. Bey-Bienko (1930):101, Bey-Bienko (1933):119, Chogsomzhav (1968):59, Chogsomzhav (1972):180, Sergeev (1995):253, Sergeev et al. (2009):109, Altanchimeg (2011):16, Batnaran et al. (2016):38, Batkhuyag and Batnaran (2021):100.

Bryodemella (Marikovskiella) semenovi (Ikonnikov, 1911)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1104279

Native status:

Distribution in the natural zone: High mountain.

Distribution:

in Mongolia: Altanchimeg and Nonnaizab (2013):81, Batkhuyag and Batnaran (2021):100.

Global distribution: Kazakhstan (Childebaev and Storozhenko 2001), Mongolia (Altanchimeg and Nonnaizab 2013).

Bryodemella (Marikovskiella) zaisanicum fallax (Bey-Bienko, 1930)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1104285

Native status:

Distribution in the natural zone: Forest steppe.

Distribution:

in Mongolia: B.-Ulg., Uvs, Khovd, Bulg. Günther (1971):125, Bey-Bienko (1930):97, Bey-Bienko (1933):119, Chogsomzhav (1989):95, Sergeev et al. (2009):109, Sergeev et al. (2020):24, Batkhuyag and Batnaran (2021):100.

Global distribution: NW Mongolia, NW China, E Kazakhstan (Sergeev et al. 2020).

Angaracris barabensis (Pallas, 1773)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1104200

Nomenclature:

Angaracris acrohylina Bi (1986):195.

Angaracris morulimarginis Huang (1981):83.

Angaracris morulipennis Zheng and He. (1994):251.

Angaracris neimongolensis Zheng and Han. (1998):25, 28.

Angaracris nigrimarginis Zheng and Ren. (1993):427.

Angaracris nigripennis Lian Z and Zheng (1984):305.

Oedipoda rhodopa Fischer von Waldheim (1836):348.

Bryodema barabensis var. rhodoptila Karny (1908):49.

Bryodema barabense var. roseipennis Krauss (1901):237.

Angaracris ulashanicus Li (1981):173.

Oedipoda hospes Fischer von Waldheim (1846):295.

Oedipoda lugubris Fischer von Waldheim (1846):298.

Oedipoda thunbergi Stål (1861):345.

Native status:

Distribution in the natural zone: Forest steppe, steppe, desert steppe and desert.

Distribution:

in Mongolia: B.-ulg., Uvs, Zav., Khuvs., A.-khang., Bulg., Tuv., Khent., S.-baat., Do., Khovd, B.-khong., U-khang. Bolívar (1901):226, 223, Pylnov (1916):280, Bey-Bienko (1930):119, Bey-Bienko (1933):119, Chogsomzhav and Shurovenkov (1963), Cejchan and Maran (1966):186, Steinmann (1967):119, Steinmann (1968):267, Mistshenko (1968):495, Chogsomzhav (1968):57-58, Chogsomzhav (1970):128, Chogsomzhav (1972):183, Altanchimeg et al. (2013b):65, Myagmar et al. (2019):56, Sergeev et al. (2020):26-28, Batkhuyag (1995):102, Dey et al. (2021):334.

Global distribution: Tuva, S Siberia, Amur Region, N Kazakhstan, Mongolia, N, NE China (Sergeev et al. 2020).

Aiolopus thalassinus (Fabricius, 1781)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1103315

Native status:

Distribution in the natural zone: Steppe.

Distribution:

in Mongolia: Uvs, A.-khang., Bulg., Khent., S.-baat., Do., B.-khong. Altanchimeg and Nonnaizab (2013):81 which is new registered species in Mongolia.

Global distribution S and C Europe, N-Africa, Caucasus, Turkey, Iran, Afghanistan, C Asia, Indian subcontinent, China, SW-Siberia (Garai 2010), South Korea (Storozhenko et al. 2015), Mongolia (Altanchimeg and Nonnaizab 2013).

Epacromius pulverulenthus (Fischer von Waldheim, 1846)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1103429

Native status:

Distribution in the natural zone: Forest steppe, steppe and desert steppe.

Distribution:

in Mongolia: Uvs, A.-khang., Bulg., Khent., S.-baat., Do., B.-khong. Mistshenko (1968):494, Günther (1971):124, Chogsomzhav (1971):86, Chogsomzhav (1972):176, Chogsomzhav (1989):94, Sergeev (1995):251, Childebaev and Storozhenko (2001), Storozhenko et al. (2015):280, Batnaran et al. (2016):38, Myagmar et al. (2019):56, Sergeev et al. (2020):16.

Global distribution: South Korea (Storozhenko et al. 2015), Tuva, S Europe, S Siberia, S Russian Far East, Kazakhstan, Tien Shan, Kashmir, Mongolia, China (Sergeev et al. 2020).

Epacromius tergestinus (Megerle von Mühlfeld, 1825)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1103430

Nomenclature:

Epacromia viridis Uvarov (1910):372.

Native status:

Distribution in the natural zone: Steppe, desert steppe and desert.

Distribution:

in Mongolia: Uvs, Bulg., Sel., Khovd, B.-khong., U.-govi. Pylnov (1916):279, Mistshenko (1968):494, Günther (1971):123, Chogsomzhav (1989):94, Sergeev (1995):251, Childebaev and Storozhenko 2001, Altanchimeg and Nonnaizab (2013), Altanchimeg et al. (2013b):65, Myagmar et al. (2019):56, Sergeev et al. (2020):16, Batkhuyag and Batnaran (2021):95.

Global distribution: Tuva, S Europe, S Siberia, Caucasus, Kazakhstan, Tien Shan, Pamiro-Alay, Afghanistan, NW Mongolia, NW China, Tibet (Sergeev et al. 2020).

Oedaleus asiaticus Bey-Bienko, 1941

Species-ID http://orthoptera.speciesfile.org/common/basic/Taxa.aspx?TaxonNameID=1103223

Native status:

Distribution in the natural zone: Forest steppe, steppe and desert steppe.

Distribution:

in Mongolia: Uvs, Bulg., Sel., Tuv, Khent., S.-baat., Do., Khovd, G.-alt., B.-khong., U-khang., Du.-govi., U.-govi. Bey-Bienko (1941):152, Chogsomzhav and Shurovenkov (1963):61, Chogsomzhav (1968):57, Chogsomzhav (1969b):128, Chogsomzhav (1972):177, Steinmann (1967):118, Steinmann (1968):246, Mistshenko (1968):495, Günther (1971):124, Childebaev and Storozhenko (2001), Altanchimeg and Nonnaizab (2013), Batnaran et al. (2016):38, Myagmar et al. (2019):56, Sergeev et al. (2020):17.

Global distribution: Siberia, Kazakhstan, China, Inner Mongolia, Mongolia, Russia (Bey-Bienko 1941, Childebaev and Storozhenko 2001, Sergeev et al. 2020).

Oedaleus decorus (Germar, 1825)

Species-ID http://orthoptera.speciesfile.org/common/basic/Taxa.aspx?TaxonNameID=1103222

Native status:

Distribution in the natural zone: High mountain.

Distribution:

in Mongolia: Chogsomzhav (1989):93, Sergeev (1995):252, Altanchimeg et al. (2013b):66, Batkhuyag and Batnaran (2021):96, Dey et al. (2021):342.

Global distribution: N Africa, Caucasus range, W Asia, C Asia, W Pakistan, Afghanistan, N India (Garai 2010), Mongolia (Dey et al. 2021).

Oedaleus infernalis Saussure, 1884

Species-ID http://orthoptera.speciesfile.org/common/basic/Taxa.aspx?TaxonNameID=1103218

Nomenclature:

Oedaleus infernalis montanus Mistshenko and Bey-Bienko (1951):221.

Microgastrimargus taeguensis Lee and Park. (1992):61-64.

Oedaleus infernalis amurensis Ikonnikov, 1911:25

Native status:

Distribution in the natural zone: Steppe and desert steppe.

Distribution:

in Mongolia: G.-alt., Uvs, Sel. Steinmann (1968):246, Pylnov (1916):279, Chogsomzhav (1971):88, Childebaev and Storozhenko (2001), Sergeev et al. (2020):20, Batkhuyag and Batnaran (2021):96.

Global distribution: South Korea (Storozhenko et al. 2015), S Russian Far East, NE, E China, Tibet, Japan, Mongolia (Sergeev et al. 2020).

Locusta migratoria (Linnaeus, 1758)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1103074

Native status:

Distribution in the natural zone: Desert steppe and desert.

Distribution:

in Mongolia: Khovd, B.-khong., U.-govi., Uvs. Chogsomzhav (1968):59, Mistshenko (1968):495, Günther (1971):124, Chogsomzhav (1989):93, Sergeev et al. (2009):109, Altanchimeg and Nonnaizab (2013), Storozhenko et al. (2015):285, Myagmar et al. (2019):56, Sergeev et al. (2020):17, Batkhuyag and Batnaran (2021):96.

Global distribution: South Korea (Storozhenko et al. 2015), Tuva, Eurasia (except the north), Africa, Australia and many islands, Mongolia (Sergeev et al. 2020).

Psophus stridulus (Linnaeus, 1758)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1103244

Nomenclature:

Psophus stridulus var. ebneri Karny (1908):57-58.

Nocarodes femoralis Fischer von Waldheim (1846):270.

Acrydium fuliginosum Olivier (1791):223.

Acrydium rubripenne De Geer (1773):472.

Psophus stridulus samniticus Baccetti (1959):397.

Native status:

Distribution in the natural zone: Forest steppe and steppe.

Distribution:

in Mongolia: Du.-govi., Sel. Bolívar (1901):226, Chogsomzhav 1972:178, Chogsomzhav (1989):93, Childebaev and Storozhenko (2001), Altanchimeg and Nonnaizab (2013), Altanchimeg et al. (2013b):65, Batnaran et al. (2016):39, Batkhuyag and Batnaran (2021):98.

Global distribution: Mongolia (Altanchimeg et al. 2013b), Tuva, Europe (except the extreme North), S Siberia, S Russian Far East, N Kazakhstan, NE China, Korea (Sergeev et al. 2020).

Celes skalozubovi Adelung, 1906

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1103549

Native status:

Distribution in the natural zone: Forest steppe, steppe, desert steppe and desert.

Distribution:

in Mongolia: Uvs, Zav., Khuvs., Bulg., Tuv. Bey-Bienko and Mistshenko (1951):587, Mistshenko (1968):495, Chogsomzhav (1969b):128, Chogsomzhav (1972):178, Chogsomzhav (1989):94, Sergeev (1995):252, Altanchimeg and Nonnaizab (2013), Altanchimeg et al. (2013b):65, Batnaran et al. (2016):39, Sergeev et al. (2020):20, Batkhuyag and Batnaran (2021):98, Dey et al. (2021):341.

Global distribution: Tuva, S Siberia, N Kazakhstan, Mongolia, N China (Sergeev et al. 2020).

Stethophyma grossum (Linnaeus, 1758)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1104012

Nomenclature:

Gryllus flavipes Gmelin (1789):2088.

Gryllus (Locusta) germanicus Stoll (1813):41.

Acrydium rubripes De Geer (1773):477.

Native status:

Distribution in the natural zone: Taiga, forest-steppe and steppe.

Distribution:

in Mongolia: B.-Ulg., Uvs, Zav., Khuvs., Tuv, A.-khang., Bulg., Sel., Khent., S.-baat., Khovd, U-khang. Pylnov (1916):279, Bey-Bienko (1933):118, Steinmann (1967):109, Mistshenko (1968):494, Chogsomzhav (1968):57, Chogsomzhav (1969b):127, Chogsomzhav (1972):162, Chogsomzhav (1974b):27, Günther (1971):115, Chogsomzhav (1989):93, Sergeev (1995):251, Childebaev and Storozhenko (2001), Altanchimeg et al. (2013b):65, Storozhenko et al. (2015):272, Batnaran et al. (2016):39, Sergeev et al. (2020):15, Batkhuyag and Batnaran (2021):95.

Global distribution: Mongolia, Tuva and almost all temperate Eurasia (except the extreme north) (Sergeev et al. 2020).

Sphingoderus carinatus (Saussure, 1888)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1103651

Native status:

Distribution in the natural zone: Desert steppe and desert.

Distribution:

in Mongolia: Khovd, U.-govi. Mistshenko (1937):186, Steinmann (1968):247, Chogsomzhav (1968):59, Chogsomzhav (1972):187, Chogsomzhav (1989):95, Childebaev and Storozhenko (2001), Sergeev et al. (2009): 109, Batkhuyag and Batnaran (2021):109.

Global distribution: China, Xinjiang, Kazakhstan, Afghanistan, Iran, Mongolia (Childebaev and Storozhenko 2001).

Sphingonotus (Sphingonotus) tzaidamicus Mistshenko, 1937

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1103783

Native status:

Distribution in the natural zone: Desert steppe and desert.

Distribution:

in Mongolia: Do.-govi. Chogsomzhav (1975):45, Chogsomzhav (1989):95, Altanchimeg (2011):16, Myagmar et al. (2019):56, Batkhuyag and Batnaran (2021):104, Dey et al. (2021):350.

Global distribution: China (Mistshenko 1937), Mongolia (Dey et al. 2021).

Sphingonotus (Sphingonotus) beybienkoi Mistshenko, 1937

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1103810

Native status:

Distribution in the natural zone: Steppe, desert steppe and desert.

Distribution:

in Mongolia: Uvs, Tuv, G.-alt., B.-khong., U-khang., Khovd, Du.-govi., U.-govi. Mistshenko (1937):148, Mistshenko (1968):495, Günther (1971):128, Chogsomzhav (1972):185, Chogsomzhav (1989):95, Sergeev (1995):255, Sergeev et al. (2009): 109, Altanchimeg et al. (2015):69, Batnaran et al. (2016):39, Myagmar et al. (2019):56, Sergeev et al. (2020):28, Dey et al. (2021):343.

Global distribution: Tuva, S Transbaikalia, C, E Kazakhstan, N Kyrgyzstan, Mongolia, NW China (Sergeev et al. 2020).

Sphingonotus (Sphingonotus) coerulipes Uvarov, 1922

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1103827

Native status:

Distribution in the natural zone: Desert steppe, taiga and forest-steppe.

Distribution:

in Mongolia: Northwest Mongolia Chogsomzhav (1989):95, Sergeev et al. (2009):109, Popova et al. (2020):604, Batkhuyag and Batnaran (2021):106, Dey et al. (2021):344.

Global distribution: Crimea, Lower Volga range, Caucasus, Turkey, Iran, Kazakhstan, Mongolia, S-Siberia (Garai 2010).

Sphingonotus (Sphingonotus) elegans Mistshenko, 1937

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1103719

Native status:

Distribution in the natural zone: Desert steppe and desert.

Distribution:

in Mongolia: Uvs, Khovd, B.-khong., U.-govi., U-khang. Mistshenko (1937):165, Mistshenko (1968):495, Chogsomzhav (1968):59, Chogsomzhav (1972):185, Günther (1971):128, Chogsomzhav (1989):95, Garai (2001):752, Sergeev et al. (2009):109, Batnaran et al. (2016):40, Myagmar et al. (2019):56, Sergeev et al. (2020):29, Dey et al. (2021):343.

Global distribution: The Mongolian part of Uvs-Nuur Intermountain Basin (Chogsomzhav 1974b), Middle Asia, NW China (Sergeev et al. 2020).

Sphingonotus (Sphingonotus) gobicus Chogsomzhav, 1975

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1103705

Native status:

Distribution in the natural zone: Desert steppe and desert.

Distribution:

in Mongolia: Khovd, G.-alt., B.-khong., Do.-govi, Chogsomzhav (1975):44, Chogsomzhav (1989):95, Batkhuyag and Batnaran (2021):105, Dey et al. (2021):345.

Sphingonotus (Sphingonotus) salinus (Pallas, 1773)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1103776

Nomenclature:

Sphingonotus suschkini Adelung (1906):86.

Oedipoda zinini Kittary (1849):470.

Native status:

Distribution in the natural zone: Desert steppe and desert.

Distribution:

in Mongolia: Khovd. Chogsomzhav (1969a):78, Chogsomzhav (1972):186, Chogsomzhav (1989):95, Sergeev (1995):255, Sergeev et al. (2009):109, Myagmar et al. (2019):56, Sergeev et al. (2020):29, Batkhuyag and Batnaran (2021):108.

Global distribution: Tuva, SE Europe, Caucasus (deserts), Kazakhstan (semi-deserts and deserts), Middle Asia, NW China, NW Mongolia (Sergeev et al. 2020).

Sphingonotus (Sphingonotus) halophilus Bey-Bienko, 1929

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1103732

Native status:

Distribution in the natural zone: Desert and desert steppe.

Distribution:

in Mongolia: Khovd. Günther (1971):128, Batkhuyag and Batnaran (2021):104.

Global distribution: Mongolia (Günther 1971), SE part of European Russia, Kazakhstan (Sergeev 2021).

Sphingonotus (Sphingonotus) mongolicus Saussure, 1888

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1103759

Native status:

Distribution in the natural zone: Taiga, forest-steppe and steppe.

Distribution:

in Mongolia: Tuv, B.-khong., U.-govi. Saussure (1888):77, 82, Mistshenko (1937):229, Mistshenko (1968):496, Chogsomzhav (1971):106, Chogsomzhav (1972):186, Chogsomzhav (1989):95, Nonnaizab et al. (1999):15, Altanchimeg and Nonnaizab 2013, Altanchimeg et al. (2013b):65, Batkhuyag and Batnaran (2021):108, Dey et al. (2021):346.

Sphingonotus (Sphingonotus) nebulosus (Fischer von Waldheim, 1846)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1103863

Native status:

Distribution in the natural zone: Desert.

Distribution:

in Mongolia: Khovd, U.-govi. Chogsomzhav (1968):58, Chogsomzhav (1971):106, Chogsomzhav (1972):186, Chogsomzhav (1975):44, Chogsomzhav (1989):95, Günther (1971):128, Sergeev (1995):255, Myagmar et al. (2019):56, Sergeev et al. (2020):29, Batkhuyag and Batnaran (2021):107.

Global distribution: Tuva, SE Altai, Asia Minor, Caucasus, Kazakhstan (except the north), Tien Shanzubo, Pamiro-Alay, Iran, Pakistan, NW Mongolia, NW China (Sergeev et al. 2020).

Sphingonotus (Sphingonotus) obscuratus latissimus Uvarov, 1925

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1103875

Native status:

Distribution in the natural zone: Desert steppe and desert.

Distribution:

in Mongolia: Khovd, B.-khong., U.-govi. Mistshenko (1968):491, Chogsomzhav (1968):59, Chogsomzhav (1971):107, Chogsomzhav (1972):186, Günther (1971):129, Chogsomzhav (1989):95, Childebaev and Storozhenko (2001), Batkhuyag and Batnaran (2021):108, Dey et al. (2021):348.

Global distribution: Kazakhstan and Mongolia (Childebaev and Storozhenko 2001).

Sphingonotus (Sphingonotus) rubescens (Walker, 1870)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1103882

Native status:

Distribution in the natural zone: High mountain and desert steppe.

Distribution:

in Mongolia: Uvs, U.-govi. Chogsomzhav (1969a):77, Chogsomzhav (1969b):128, Chogsomzhav (1972):185, Chogsomzhav (1989):95, Sergeev (1995):255, Sergeev et al. (2009):109, Batnaran et al. (2016):40, Myagmar et al. (2019):56, Sergeev et al. (2020):29, Batkhuyag and Batnaran (2021):105, Dey et al. (2021):348.

Global distribution: The Mongolian part of Uvs-Nuur Intermountain Basin (Chogsomzhav 1974b). Arid part of N Caucasus, N Africa, SW Asia, deserts of Middle Asia (including mountains), NW Mongolia, NW China (Sergeev et al. 2020).

Helioscirtus moseri Saussure, 1884

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1103593

Native status:

Distribution in the natural zone: Desert.

Distribution:

in Mongolia: G.-alt. Chogsomzhav (1969b), Batkhuyag and Batnaran (2021):109.

Global distribution: Mongolia (Chogsomzhav 1969b), northern Africa and western Asia (Hodjat et al. 2018).

Leptopternis gracilis (Eversmann, 1848)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1104641

Nomenclature:

Hyalorrhipis maculipennis Saussure (1884):195.

Sphingonotus angustipennis Werner (1905):201.

Sphingonotus grobbeni Chopard (1949):361.

Native status:

Distribution in the natural zone: Desert steppe and desert.

Distribution:

in Mongolia: U.-govi. Mistshenko (1968):496, Chogsomzhav (1972):187, Chogsomzhav (1989):95, Garai (2001):753, Childebaev and Storozhenko (2001), Batkhuyag et al. (2019):107, Myagmar et al. (2019):57, Batkhuyag and Batnaran (2021):109.

Global distribution: N Africa, Lower Volga range, Caucasus range, Iran, Afghanistan, C Asia, W China, Mongolia (Garai 2001).

Leptopternis iliensis Uvarov, 1925

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1104645

Native status:

Distribution in the natural zone: Desert steppe.

Distribution:

in Mongolia: Khovd. Günther (1971):129, Chogsomzhav (1989):95, Childebaev and Storozhenko (2001), Batkhuyag and Batnaran (2021):109.

Global distribution: Kazakhstan and Mongolia (Childebaev and Storozhenko 2001).

Bohemanella frigida (Boheman, 1846)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1110374

Native status:

Distribution in the natural zone: Taiga and forest-steppe.

Distribution:

in Mongolia: Uvs, Zav., Khuvs., Tuv. Uvarov (1914):17, Mistshenko (1952):425, Mistshenko (1968):489, Steinmann (1968):240, Cejchan and Maran (1966):178, Chogsomzhav (1969b):127, Chogsomzhav (1972):154, Chogsomzhav (1989):90, Günther (1971):113, Munkhbat (2010):169, Altanchimeg et al. (2013b):64, Sergeev et al. (2019):14, Batkhuyag and Batnaran (2021):44.

Global distribution: Tuva, N Eurasia (in the southern parts of Europe in mountains), Alaska, N Canada, Mongolia (Sergeev et al. 2019).

Podisma pedestris (Linnaeus, 1758)

Species-ID http://orthoptera.speciesfile.org/Common/Basic/Taxa.aspx?TaxonNameID=1111031

Native status:

Distribution in the natural zone: Taiga and forest-steppe.

Distribution:

in Mongolia: Sel., Khuvs. Mistshenko (1952), Chogsomzhav (1972):154, Chogsomzhav (1989):90, Altanchimeg et al. (2013b):64, Sergeev et al. (2019):13, Batkhuyag and Batnaran (2021):43.

Global distribution: Tuva, Europe, W Siberia (forest steppes and steppes), E Siberia (up to the central parts of Yakutia), mountains of S Siberia, N Caucasus, NW and N Kazakhstan, Dzungarian Alatau, E Tien Shan, Tarbagatai Mts, N Mongolia (Sergeev et al. 2019).

Prumna primnoa (Motschulsky, 1846)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1111290

Nomenclature:

Podisma sachalinensis Matsumura (1911):5.

Podisma viridis Fischer von Waldheim (1846):248.

Native status:

Distribution in the natural zone: Forest steppe and steppe.

Distribution:

in Mongolia: Khent., Sel., Tuv. Pylnov (1916):280, Cejchan and Maran (1966): 178, Chogsomzhav (1989):90, Batnaran (2008):43, Batnaran et al. (2016):36, Sergeev et al. (2019):13, Batkhuyag and Batnaran (2021):43.

Global distribution: Tuva, S Siberia, the southern part of Russian Far East (except the southern part of Primorsky Region), including Sakhalin and Kunashir, N Mongolia (Sergeev et al. 2019).

Ognevia longipennis (Shiraki, 1910)

Species-ID http://orthoptera.speciesfile.org/Common/Basic/Taxa.aspx?TaxonNameID=1111098

Nomenclature:

Eirenephilus debilis Ikonnikov (1911):265.

Podisma alpina niphona Furukawa (1929):171,177.

Native status:

Distribution in the natural zone: Taiga and forest steppe.

Distribution:

in Mongolia: Khent., Sel. Bey-Bienko and Mistshenko (1951):236, Chogsomzhav (1989):90, Storozhenko and Paik (2007):157, Altanchimeg et al. (2013b):64, Storozhenko et al. (2015):220, Sergeev et al. (2019):13, Batkhuyag and Batnaran (2021):45.

Global distribution: Tuva, S Siberia (mainly in mountains), S Russian Far East, including Sakhalin and S Kurile Islands, E Kazakhstan, N Mongolia, N, NE China, Korea, Japan (Sergeev et al. 2019).

Zubovskya koeppeni (Zubovski, 1900)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1111120

Native status:

Distribution in the natural zone: Forest steppe.

Distribution:

in Mongolia: Khuvs. Chogsomzhav (1989):90, Batkhuyag and Batnaran (2021):43.

Global distribution: Mongolia (Chogsomzhav 1989) , Tuva, WSE, Altay-Sayan Mts. including W Altay (Sergeev et al. 2019).

Zubovskya mongolica Storozhenko, 1986

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1111118

Native status:

Distribution in the natural zone: Forest steppe.

Distribution:

in Mongolia: Khuvs. Storozhenko (1986):53, Sergeev et al. (2019):12.

Global distribution: Mongolia and Siberia (Storozhenko 1986, Sergeev et al. 2019).

Calliptamus abbreviatus Ikonnikov, 1913

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1112890

Nomenclature:

Calliptamus doii Lee and Lee (1985):24.

Calliptamus abbreviatus f. holoptera Ramme (1952):308.

Calliptamus sibiricus Wnukowskij (1926):91.

Native status:

Distribution in the natural zone: High mountain, forest steppe, desert steppe and desert.

Distribution:

in Mongolia: Uvs, Zav., Khuvs., Bulg., Sel., Khent., Do., Khovd, B.-khong, U-khang, Du.-govi., U.-govi. Pylnov (1916):280, Mistshenko (1952):531, Mistshenko (1968):489, Steinmann (1968):240, Chogsomzhav (1968):57, Chogsomzhav (1969b):127, Chogsomzhav (1972):155, Chogsomzhav (1975):38, Günther (1971):113, Batnaran (2008):44, Sergeev (1995):240, Childebaev and Storozhenko (2001):25, Altanchimeg (2011):16, Altanchimeg and Nonnaizab 2013, Altanchimeg et al. (2013b):65, Storozhenko et al. (2015):225, Batnaran et al. (2016):31, Sergeev et al. (2019):15, Myagmar et al. (2019):56, Batkhuyag and Batnaran (2021):45.

Global distribution: Tuva, S Siberia (from the south-eastern part of W Siberian Plain to Daura), the southern part of the Russian Far East, NE, E Kazakhstan, N Mongolia, N, NE, E China, South Korea (Sergeev et al. 2019).

Calliptamus barbarus subsp. cephalotes Fischer-Waldheim, 1846

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1112930

Native status:

Distribution in the natural zone: Steppe, desert steppe and desert.

Distribution:

in Mongolia: Khovd, G.-alt., B.-khong., U.-govi. Mistshenko (1952):544, Mistshenko (1968):489, Chogsomzhav (1968):59, Chogsomzhav (1989):90, Childebaev and Storozhenko (2001):25, Garai (2001):402, Altanchimeg and Nonnaizab (2013), Batnaran et al. (2016):31, Myagmar et al. (2019):56, Batkhuyag and Batnaran (2021):46.

Global distribution: N Africa, Caucasus, Turkey, Iran, N-Afghanistan, Kazakhstan, Mongolia, W China, Siberia (Garai 2001).

Calliptamus italicus (Linnaeus, 1758)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1112938

Native status:

Distribution in the natural zone: Steppe.

Distribution:

in Mongolia: Khuvs. Mistshenko and Bey-Bienko (1951):256, Chogsomzhav (1971):53, Chogsomzhav (1989):90, Childebaev and Storozhenko (2001):25, Sergeev et al. (2009):108, Altanchimeg and Nonnaizab 2013, Batkhuyag and Batnaran (2021):46.

Global distribution: N Africa, Turkey, from the Caucasus through Iran, Afghanistan, W Pakistan, to NW Mongolia, W China and W Siberia (Garai 2001).

Dericorys annulata (Fieber, 1853)

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1117733

Nomenclature:

Dericorys lazurescens Uvarov (1914):142, 146.

Derocorystes (Cyphophorus) roseipennis Redtenbacher (1889):30.

Native status:

Distribution in the natural zone: Desert steppe.

Distribution:

in Mongolia: Khovd, B.-khong., U.-govi. Mistshenko and Bey-Bienko (1951):151, Mistshenko (1952):97, Mistshenko (1968):489, Chogsomzhav (1968):59, Chogsomzhav (1972):153, Chogsomzhav (1974b):26, Steinmann (1971):146, Childebaev and Storozhenko (2001):21, Altanchimeg et al. (2015):69, Myagmar et al. (2019):57, Batkhuyag and Batnaran (2021):39.

Global distribution: Kazakhstan, Afghanistan, China, Mongolia (Childebaev and Storozhenko 2001).

Beybienkia lithophila Gorochov & Mistshenko, 1989

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1117444

Native status:

Distribution in the natural zone: Desert steppe.

Distribution:

in Mongolia: B.-khong. Podgornaya and Gorochov 1989:105, Batkhuyag and Batnaran 2021:37.

Global distribution: Mongolia (Ünal 2016).

Beybienkia songorica Tsyplenkov, 1956

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1117445

Native status:

Distribution in the natural zone: Desert.

Distribution:

in Mongolia: Altai, Transaltai gobi Chogsomzhav (1989):89, Batkhuyag and Batnaran (2021):37.

Global distribution: Mongolia (Chogsomzhav 1989), China (Tsyplenkov 1956).

Mongolotmethis gobiensis Bey-Bienko, 1948

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1117481

Native status:

Distribution in the natural zone: Steppe and desert steppe.

Distribution:

in Mongolia: B.-khong., U.-govi.,U-khang. Bey-Bienko (1948):9, Bey-Bienko and Mistshenko (1951):321, Mistshenko (1968):489, Chogsomzhav (1968):58, Altanchimeg (2011):16, Batkhuyag et al. (2014):78, Myagmar et al. (2019):57, Batkhuyag and Batnaran (2021):34.

Global distribution: China, Inner Mongolia (Alashan), Mongolia (Batkhuyag et al. 2014).

Mongolotmethis kozlovi Bey-Bienko, 1948

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1117480

Native status:

Distribution in the natural zone: Steppe and desert steppe

Distribution:

in Mongolia: Tuv, Du.-govi., B.-khong., U.-govi. Bey-Bienko (1948):10, Bey-Bienko and Mistshenko (1951):321, Mistshenko (1968):489, Chogsomzhav (1968):58, Altanchimeg (2011):16, Batkhuyag et al. (2014):78, Myagmar et al. (2019):57, Batkhuyag et al. (2019):107, Batkhuyag and Batnaran (2021):34.

Mongolotmethis michidi Batkhuyag, Batnaran & Dorjderem, 2014

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1221002

Native status:

Distribution in the natural zone: Steppe and desert steppe.

Distribution:

in Mongolia: G.-alt., U.-govi. Batkhuyag et al. (2014), Batkhuyag and Batnaran (2021):35.

Rhinotmethis beybienkoi Chogsomzhav, 1975

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1117505

Native status:

Distribution in the natural zone: Desert steppe and desert.

Distribution:

in Mongolia: Du.-govi. Chogsomzhav (1975):39, Gorochov et al. (1989), Chogsomzhav (1989):89, Altanchimeg (2011):16, Batkhuyag et al. (2014):78, Batnaran et al. (2016):30, Ünal (2016):59, Batkhuyag and Batnaran (2021):36.

Global distribution: Mongolia (Ünal 2016).

Rhinotmethis hummeli Sjöstedt, 1933

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1117506

Native status:

Distribution in the natural zone: Desert steppe and desert.

Distribution:

in Mongolia: Do.-govi. Bey-Bienko (1948):12, Chogsomzhav (1975):33, Chogsomzhav (1989):89, Sergeev (1995):233, Ünal (2016):59, Myagmar et al. (2019):57.

Global distribution: China, Inner Mongolia, Mongolia (Ünal 2016, Batkhuyag and Batnaran 2021).

Asiotmethis similis Bey Bienko, 1951

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1117408

Native status:

Distribution in the natural zone: Forest steppe and desert steppe.

Distribution:

in Mongolia: G.-alt. Chogsomzhav (1989):89, Batkhuyag and Batnaran (2021):31.

Global distribution: Central Asia, Kazakhstan (Bey-Bienko and Mistshenko 1951), Mongolia (Chogsomzhav 1989).

Haplotropis brunneriana Saussure, 1888

Species-ID http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1116957

Nomenclature:

Sulcotropis cyanipes Yin and Chou. (1979):128.

Staurotylus mandshuricus Adelung (1910):344.

Haplotropis neimongolensis Jin (1994):251

Native status:

Distribution in the natural zone: Forest steppe and steppe.

Distribution:

in Mongolia: Do. Chogsomzhav (1975):41, Chogsomzhav (1989):95, Storozhenko and Paik (2007):133, Storozhenko et al. (2015):184, Batnaran et al. (2016):31, Ünal (2016):60, Batkhuyag and Batnaran (2021):38.

Global distribution: China, Inner Mongolia (Jin 1994), Manchuria (Yin and Chou. 1979), Russian Far East, eastern Asia, South Korea, China North-central, Mongolia (Storozhenko and Paik 2007, Ünal 2016).

Analysis

Result

The present study aimed to list the rare and unexplored species of grasshoppers (except long-horned grasshopper and cricket species) in Mongolia. Currently, the grasshopper fauna of Mongolia comprises 128 species, which are distributed in 52 genera and 19 tribes (Table 2). Of these, 34 species are also included in the Checklist of European Orthoptera (Acridoidea) (Suppl. material 2). In addition, Mongolia's distribution of grasshoppers was divided into six distinct natural zones (Fig. 3), (Suppl. material 3). Additionally, a combined cluster analysis was performed using the grasshopper component numbers between neighbouring countries, including Russia, China and South Korea (Suppl. material 4).

Table 1.

Download as

CSV

XLSX

Similarity matrix of grasshopper in six types of natural zones in Mongolia.

Natural zone	High Mountain (%)	Taiga (%)	Forest steppe (%)	Steppe (%)	Desert steppe (%)	Desert (%)
High Mountain	*	12.12	11.11	26.37	29.17	21.88
Taiga	*	*	46.58	28.26	20.62	12.31
Forest steppe	*	*	*	59.54	36.76	25.00
Steppe	*	*	*	*	64.52	40.65
Desert steppe	*	*	*	*	*	64.06
Desert	*	*	*	*	*	*

Cluster analysed amongst six different types of natural zone shown in %.

Table 2.

Download as

CSV

XLSX

Composition of grasshoppers (Acridoidea) in Mongolia.

No.	Subfamily	Tribe	Genus	Number of Species	Registered in the European red list
1	Acridinae MacLeay, 1821	Acridini MacLeay, 1821	Acrida Linnaeus, 1758	1	-
2	Gomphocerinae Fieber, 1853	Arcypterini Bolívar, 1914	Arcyptera Serville, 1838	4	2
6		Arcypterini Bolívar, 1914	Andrea Mistshenko, 1989	1	-
7		Chrysochraontini Brunner von Wattenwyl, 1893	Chrysochraon Fischer, 1853	1	-
8			Euthystira Fieber, 1852	1	1
9			Mongolotettix Rehn, 1928	3	-
12			Podismopsis Zubovski, 1900	2	-
14		Dociostaurini Mistshenko, 1974	Eremippus Uvarov, 1926	3	1
17		Dociostaurini Mistshenko, 1974	Notostaurus Bey-Bienko, 1933	1	1
18		Hypernephiini Mistshenko, 1973	Eclipophleps Tarbinsky, 1927	8	-
26		Gomphocerini Fieber, 1853	Chorthippus Fieber, 1852	16	8
42			Aeropedellus Hebard, 1935	4	1
46			Pseudochorthippus Defaut, 2012	2
48			Gomphocerippus Roberts, 1941	1	1
49			Gomphocerus Thunberg, 1815	1	1
50			Myrmeleotettix Bolívar, 1914	2	-
52			Stauroderus Bolívar, 1897	1	1
53			Schmidtiacris Storozhenko, 2002	1	-
54			Mesasippus Tarbinsky, 1931	1	-
55			Dasyhippus Uvarov, 1930	1	-
56			Dociostaurus Fieber, 1853	2	1
58	Oedipodinae Walker, 1871	Stenobothrini Harz, 1975	Omocestus Bolívar, 1878	5	4
63			Stenobothrus Fischer, 1853	5	3
68			Megaulacobothrus Caudell, 1921	1
69			Leptopternis Saussure, 1884	2	1
71		Bryodemini Bey-Bienko, 1930	Bryodema Fieber, 1853	6	-
77			Compsorhipis Saussure, 1889	3	-
80			Bryodema Yin, 1982	5	-
85			Angaracris Bey-Bienko, 1930	1	-
86		Epacromiini Brunner von Wattenwyl, 1893	Aiolopus Fieber, 1853	1	-
87		Epacromiini Brunner von Wattenwyl, 1893	Epacromius Uvarov, 1942	2	1
89		Locustini Kirby, 1825	Oedaleus Fieber, 1853	3	-
92			Locusta Linnaeus, 1758	1	1
93			Psophus Fieber, 1853	1	1
94		Oedipodini Walker, 1871	Celes Saussure, 1884	1	-
95		Parapleurini Brunner von Wattenwyl, 1893	Stethophyma Fischer, 1853	1	-
96		Sphingonotini Johnston, 1956	Sphingoderus Bey-Bienko, 1950	1	1
97			Sphingonotus Fieber, 1852	11	3
108			Helioscirtus Saussure, 1884	1
109	Melanoplinae Scudder, 1897	Podismini Jacobson, 1905	Bohemanella Ramme, 1951	1	-
110			Podisma Berthold, 1827	1	-
111			Prumna Motschulsky, 1859	1	-
112			Ognevia Ikonnikov, 1911	1	-
113			Zubovskya Dovnar-Zapolskij, 1932	2	-
115	Calliptaminae Jacobson, 1905	Calopteni Brunner von Wattenwyl, 1893	Calliptamus Serville, 1831	3	1
118	Egnatiinae Bey-Bienko & Mistshenko, 1951	Egnatiini Bey-Bienko & Mistshenko, 1951	Egnatioides Vosseler, 1902	1
119	Dericorythinae Jacobson and Bianchi, 1905	Derocorythini Jacobson & Bianchi, 1905	Dericorys Serville, 1838	1	-
120	Thrinchinae Stål, 1876	Thrinchini Stål, 1876	Beybienkia Tsyplenkov, 1956	2	-
123			Mongolotmethis Bey-Bienko, 1948	3	-
125			Rhinotmethis Sjöstedt, 1933	2	-
127			Asiotmethis Uvarov, 1943	1	-
128		Haplotropiidini Sergeev, 1995	Haplotropis Saussure, 1888	1	-
Total	8	19	52	128	34

Figure 1.

The grasshopper distribution of six different natural zones in Mongolia. Desert steppe - 80 species (27.30%), Steppe - 75 species (25.93%), Forest steppe - 56 species (19.11%), Desert - 48 species (16.38%), Taiga - 17 species (5.80%), High Mountain - 16 species (5.46%) distributed, respectively.

Figure 2.

Dendrogram showing the Bray-Curtis cluster analysis (Single link) of distribution of grasshopper species in six different natural zones (a); Species distribution rank abundance plot of Mongolian grasshopper by Natural Zone (b); Bray-Curtis cluster analysis (single link) neighbouring boundary countries (Russia, China and South Korea) grasshopper distribution (c); Abundance Plot of boundary countries (Russia, China including South Korea) grasshopper distribution (d).

Figure 3.

Distribution of grasshoppers in Mongolian natural zones in the High mountain natural zone (a); taiga natural zone (b); forest steppe natural zone (c); steppe natural zone (d); desert steppe natural zone (e); and desert natural zone (f).

The rich distribution of Mongolian grasshoppers was characterised by six habitats using the remnant natural habitat and forest types (Fig. 1). The total number of grasshopper species in each natural zone was as follows: 75 species (25.93%) in the steppe, 16 species (5.46%) in the high mountains, 56 species (19.11%) in the forest-steppe, 80 species (27.30%) in the desert steppe, 17 species (5.80%) in the taiga and 48 species (16.38%) in the desert. The similarity matrix between the geographical distribution of grasshopper results presented a high mountain and taiga zone of 12.1%, taiga and forest-steppe zone of 46.57%, forest-steppe and steppe zone of 59.54%, steppe and desert steppe zone of 64.51% and desert steppe and desert zone of 64.06% (Table 1). The species natural zone results used by a single-link Bray-Curtis cluster analysis dendrogram (Fig. 2a, b) considered desert and desert steppe zones, steppe and forest-steppe zones and high mountain and taiga zones as most closely related. The high mountain and taiga zones were less distributed than the other zones. In addition, the Shannon index and Berger–Parker's index showed six different natural zones (Suppl. material 3). Desert steppe and steppe zones were the most distributed. In contrast, the high mountain and taiga zones were less distributed. Grasshoppers are widely distributed in the desert steppe and steppe natural zone (Suppl. material 1). Twenty grasshopper species that are extensively dispersed in the desert-steppe natural zone have been identified as the indicator species of this zone. These species are listed as follows: Rhinotmethis hummeli Sjost.* Dericorys annulata (Fieb.), Calliptamus barbarus cephalotes F.-W., Acrida kozlovi Mistsh., Arcyptera meridionalis Ikonn., Arcyptera microptera (F.-W.), Stenobothrus fischeri Ev., Celes skalozubovi Adel., Compsorhipis bryodemoides B.-Bien.*, Leptopternis gracilis (Ev.), Sphingoderus carinatus (Sauss.) Sphingonotus beybienkoi Mistsh., Sphingonotus coerulipes Uv., Sphingonotus elegans Mistsh., Sphingonotus gobicus Chogs.*, Sphingonotus nebulosus (F.-W.), Sphingonotus obscuratus latissimus Uv., Sphingonotus rubescens (Walker), Sphingonotus salinus (Pall.) and Sphingonotus tzaidamicus Mistsh. The indicator species of each of the six types of natural zones are identified by (★) abbreviations (Suppl. material 1). In total, 16 species (13.17%) were endemic and were commonly distributed in desert steppe and desert natural zones. The similarity matrix showed the species distribution of grasshoppers in Mongolia, Russia, China and South Korea. A single-link Bray-Curtis cluster analysis dendrogram was constructed using the combined distribution data for all species. The results exhibited the relationship between a Mongolian grasshopper species and species in Russia and China that were most closely related to the Mongolian species. South Korea was reported as the country that was most distantly linked to Mongolia (Fig. 2). The number of Mongolian grasshopper species was compared with those of grasshopper species in the neighbouring countries, such as Russia and China and South Korea was included using a single-link Bray-Curtis cluster analysis dendrogram (Fig. 2c). The similarity index between the number of species of Mongolian grasshopper and those of grasshopper species in China, Russia and South Korea was 68.36%, 76.55% and 26.84%, respectively. Furthermore, grasshopper species from China showed 36.78% and 65.30% similarity with those in South Korea and Russia, respectively. South Korea showed that 36% of the Russian-distributed species were similar (Table 3). In summary, Russia and China are closely tied to the number of Mongolian grasshopper species, whereas South Korea is distantly related.

Table 3.

Download as

CSV

XLSX

Similarity matrix of grasshopper distribution in neighbouring countries (China, Russia and South Korea).

Name of Country	Mongolia (%)	South Korea (%)	Russia (%)	China (%)
Mongolia	*	26.85	76.56	68.37
South Korea	*	*	36.00	36.78
Russia	*	*	*	65.31
China	*	*	*	*

In addition, 17 species of grasshoppers are endemic to Mongolia (13.17%), including Mongolotmethis gobiensis B.-Bien, Mongolotmethis kozlovi B.-Bien, Rhinotmethis hummeli Sjost, Podismopsis altaica Zub, Eclipophleps bogdanovi Tarb, Eclipophleps carinata Mistsh, Eclipophleps confinis Mistsh, Eclipophleps glacialis B.-Bien, Eclipophleps kerzhneri Mistsh, Eclipophleps lucida Mistsh, Eclipophleps similis Mistsh, Eclipophleps tarbinskii Oristsh, Stenobothrus newskii Zub, Bryodema gebleri (F.-W.), Bryodema (M.) orientalis B.-Bien and Compsorhipis bryodemoides B.-Bien. Also, Chorthippus (G.) mollis (Charp.), Chorthippus (G.) vagans (Ev.), Chorthippus (M.) chinensis Tarb., Aiolopus thalassinus (Fabr.), Bryodema heptapotanicum B.-Bien., Bryodema miramae B.-Bien, Bryodema (M.) semenovi Ikonn., Sphingonotus gobicus Chogs. species are new identified species from Mongolia Altanchimeg and Nonnaizab (2013) (Suppl. material 1). Furthermore, 34 grasshopper species are registered on the European Red List and two of them are listed as Endangered, four are listed as Near Threatened and 28 are classified as Least Concern (Suppl. material 2).

Discussion

The grasshopper fauna of Mongolia comprises 128 species, including three families, eight subfamilies, 19 tribes and 52 genera, of which 34 species are registered on the European Red List (Table 2). The 17 grasshopper species that are considered endemic to Mongolia are distributed in desert and desert steppe. Notably, some species are widely distributed in neighbouring countries, such as Russia, China and South Korea. The taxonomic keys of the superfamily Acridoidea in Mongolia were recently updated and it was reported that the superfamily includes three families, 49 genera and 127 species (Batkhuyag and Batnaran 2021). A difference of six species was observed between our checklist and the list updated by Batkhuyag and Batnaran (2021). These species included Gomphocerus licenti Chang, Chorthippus (Ch.) karelini Uv., Stenobothrus kirgizorum Ikonn., Sphingonotus halocnemi Uv., Sphingonotus turcmenus B.-Bien. and Eclipophleps confinis levis Mistsh. In addition, the records of specimens of four species were uncertain; these included Chorthippus (G.) buyanticus Batkhuyag et al., Chorthippus (G.) tseelicus Batlhuyag et al., Sphingonotus halocnemi Uv. and Sphingonotus turcmenus B.-Bien. In the present study, four species with uncertain distribution sources, namely Gomphocerus licenti Chang, Chorthippus (Ch.) karelini Uv., Stenobothrus kirgizorum Ikonn. and Eclipophleps levis Mistsh, were excluded from our checklist. In the future, the species list should be updated after previously unreported species have been identified and reported by undertaking additional research.

We compared the findings of our checklist with those of Batkhuyag and Batnaran (2021) and we found discrepancies related to three genera, namely Aiolopus Fieber, 1853, Pseudochorthippus Defaut, 2012 and Megaulacobothrus Caudell, 1921, as well as three species, which includes Aeropedellus baliolus Mistsh., Bryodema kozlovi B.-Bien., and Aiolopus thalassinus (Fabr.). These genus species are widely distributed in forests and steppe in Mongolia. When creating an annotated checklist, the species that were identified from literature were divided into natural zones, based on their distribution. However, Mongolia has the world’s largest intact grassland with respect to its biodiversity (Batsaikhan et al. 2014, Herbert et al. 2019), which has great importance for the preservation of native vascular plants (Baasanmunkh et al. 2022). Thus, it is important to study grasshopper's habitat and development, which negatively impact grassland. At the same time, there is an enormous shortage of taxonomists who can identify and describe species (Wheeler 2004). The loss of diversity coupled with the taxonomic impediment is one of the most challenging issues we biologists face today (Song 2010). A poor understanding of grasshopper fauna has impaired our understanding of grasshopper speciation and evolution. Therefore, further in-depth surveys of grasshoppers in Mongolia should be conducted and it is expected that the taxonomic uncertainty checked in this study can be solved through future studies.

Notably, some species are widely distributed in neighbouring countries, such as Russia, China and South Korea. The taxonomic keys of the superfamily Acridoidea in Mongolia were recently updated and it was reported that the superfamily includes three families, 49 genera and 127 species (Batkhuyag and Batnaran 2021). A difference of six species was observed between our checklist and the list updated by Batkhuyag and Batnaran (2021). These species included Gomphocerus licenti Chang, Chorthippus (Ch.) karelini Uv., Stenobothrus kirgizorum Ikonn., Sphingonotus halocnemi Uv., Sphingonotus turcmenus B.-Bien. and Eclipophleps levis Mistsh. In addition, the records of specimens of four species were uncertain; these included Chorthippus (G.) buyanticus Batkhuyag et al., Chorthippus (G.) tseelicus Batlhuyag et al., Sphingonotus halocnemi Uv. and Sphingonotus turcmenus B.-Bien. In the present study, four species had uncertain distribution sources, namely Gomphocerus licenti Chang, Chorthippus (Ch.) karelini Uv., Stenobothrus kirgizorum Ikonn. and Eclipophleps levis Mistsh and were excluded from our checklist. We are aware that this list is only a basis for further research and we hope that it will be further modified by the work of future scientists who devote their time and passion to researching new and interesting facts about Mongolian grasshoppers.

Acknowledgements

The authors appreciate all laboratory members of the Institute for Phylogenomics and Evolution at Kyungpook National University in South Korea, for helping with all their support and for helping us to draw and upgrade the figures on the manuscripts.

Author contributions

U.W.H conceived the study and Altanchimeg, D is the first author who analysed the data and helped to upgrade the tables on the manuscripts. All authors reviewed the manuscripts.

References

Adelung N (1906)

Materials on the fauna and flora of the Russian empire.

Zoology

(

)

‑

. [In

Russian

Adelung N (1910)

Horae Societatis Entomologicae Rossicae, variis sermonibus in Rossia usitatis editae

Typis V. Besobrasovi & Company

328-358

pp. URL: https://www.biodiversitylibrary.org/item/84670

Altanchimeg D, Nonnaizab N (2005)

Study on the karyotype of Bryodema holdereri (Acridoidea)

Proceedings of Institute of Biology

234

‑

236

. [In

Mongolian

Altanchimeg D (2011)

Acridoidea of Mongolia

Issues and current status of insect conservation in Mongolia

Mongolia

2011

Ulaanbaatar

15-16

pp. [In

Mongolian

Altanchimeg D, Nonnaizab N (2013)

Grasshoppers (Acridoidea) of Mongolian Plateau

In: Zhang L et al. (Eds) Orthoptera in scientific progress and human culture

11th International congress of Orthopterology

Kunming (China)

11-15th, August 2013

Metaleptea - The newsletter of the Orthopterists' Society (special issue)

81-82

pp.

Altanchimeg D, Lin C, Nonnaizab N (2013a)

Comparative study on the karyotype of two species of Megaulacobothrus Caud., 1921 (Acridoidea)

Mongolian Journal of Biological Sciences

(

1-2

‑

. URL: https://www.biotaxa.org/mjbs/article/view/26585

Altanchimeg D, Uranbileg G, Unurzaya K, Zulbayr M, Dorjderem S (2013b)

Grasshoppers (Acridoidea) of Khan khentii protected area

A multidisciplinary study of the eco-environment

Mongolia

64-69

pp. [In

Mongolian

Altanchimeg D, Chen L, Nonnaizab N (2014)

A new species of the genus Aeropedellus from the Hovsgol province of Mongolia (Orthoptera: Acrididae: Gomphocerinae)

Transactions of the American Entomological Society

140

(

133

‑

136

. https://doi.org/10.3157/061.140.0108

Altanchimeg D, Enkhnasan D, Arigunsudar P (2015)

Insect biodiversity of Gurvan tes district, Omnogovi province

Proceedings of Institute of Biology

‑

. [In

Mongolian

Baasanmunkh S, Urgamal M, Oyuntsetseg B, Sukhorukov AP, Tsegmed Z, Son DC, Erst A, Oyundelger K, Kechaykin AA, Norris J, Kosachev P, Ma JS, Chang KS, Choi HJ (2022)

Flora of Mongolia: annotated checklist of native vascular plants

PhytoKeys

192

‑

169

. https://doi.org/10.3897/phytokeys.192.79702

Baccetti B (1959)

Notulae Orthopterologicae. X. Ricerche sugli Ortotteridel Gran Sasso d’Italia per il Centro di Entomologia Alpina

Redia

(

351

‑

450

Batkhuyag B (1995)

Studying the biology and ecology of the main pest grasshoppers and pest management in Mongolia. Thesis of Ph.D

Mongolian Agricultural University

Ulaanbaatar, Mongolia

150

pp. [In

Mongolian

Batkhuyag B, Batnaran K, Dorjderem S (2014)

A new species of Mongolotmethis from the Gobi Region of Mongolia (Orthoptera: Pamphagidae)

Journal of Orthoptera Research

(

‑

. https://doi.org/10.1665/034.023.0202

Batkhuyag B, Bakey A, Batchuluun Y, Chimeddorj B, Dagvasuren M, Davaadorj G, Gombobaatar S, Munkhchuluun B (2019)

Sixth national report to the convention on biological diversity (2015-2018)

Ministry of Environment and Tourism of Mongolia

106-109p

Batkhuyag B, Batnaran B (2021)

Key of the short-horned orthopteroid insects in Mongolia.

ADMON Printing Company

Ulaanbaatar

134

pp. [In

Mongolian

]. URL: https://mongolia.panda.org/publications/?uNewsID=372940

Batnaran B, Mendjargal B, Davaasuren J (1999)

The history of pest control in Mongolia.

In: Chogsomzhav L (Ed.)

Conference of "Memories of the year" dedicated L. Chogsomjav

Mongolian Agricultural University

Ulaanbaatar

60-63

pp. [In

Mongolian

Batnaran K (2008)

Studying the biology and ecology of some pest grasshoppers and pest control in central Mongolia.Thesis of Ph.D.

Mongolian Agricultural University

Ulaanbaatar

126

pp. [In

Mongolian

Batnaran K, Batkhuyag B, Otgonchimeg T, Dorjderem S, Turbat T, Gandulam R (2016)

A study on the karyotype of some pest grasshoppers in Mongolia.

Report of Science and Technology Foundation

‑

. [In

Mongolian

Batsaikhan N, Buuveibaatar B, Chimed B, Enkhtuya O, Galbrakh D, Ganbaatar O, Lkhagvasuren B, Nandintsetseg D, Berger J, Calabrese JM (2014)

Conserving the world's finest grassland amidst ambitious national development

Conservation Biology

(

1736

‑

1739

. https://doi.org/10.1111/cobi.12297

Bazelet C, Samways M (2012)

Grasshopper and butterfly local congruency in grassland remnants

Journal of Insect Conservation

(

‑

. https://doi.org/10.1007/s10841-011-9394-7

Benediktov AA (1999)

To little-known taxa of Chorthippus biguttulus group (Orthoptera, Acrididae, and Gomphocerinae)

Vestnik Moskovskogo Universiteta Seriya

XVI Biologiya 1

‑

Benediktov AA (2016)

Variability of wings pattern of the locusts from the tribe Bryodemini Bey-Bienko (Orthoptera: Acrididae)

320

Proceedings of the Zoological Institute of the Russian Academy of Sciences

467-472

pp. https://doi.org/10.31610/trudyzin/2016.320.4.467

Bey-Bienko GY (1926)

Notes on some Orthoptera from Palaearctic Asia

Transactions Siberian Academic Agriculture Forest

(

199

‑

211 [1-13, 3 figs.]

Bey-Bienko GY (1930)

A monograph of the genus Bryodema Fieb. (Orthoptera, Acrididae) and its nearest allies.

Annuaire du Musée Zoologique de l'Académie Impériale des Sciences de Sant-Pétersbourg

(

‑

127. Plates XVIII–XX

Bey-Bienko GY (1932)

Notes on the genus Compsorhipis Sauss. (Orthoptera: Acrididae).

A Journal of Taxonomic Entomology

‑

Bey-Bienko GY (1933)

Orthoptera collected by Prof. V. Baranov in north western Mongolia.

Boletín de la Real Sociedad Española de Historia Natural

105

‑

119

Bey-Bienko GY (1941)

New and little known Orthoptera found in the USSR.

Zapiski Leningradskogo Selskokhozjastvennogo

147

‑

159

. [In

Russian

Bey-Bienko GY (1948)

Grasshoppers of the tribe Thrinchini (Orthoptera, Acrididae) collected by Russian investigators in Mongolia and limitrophic China

Entomologicheskoe Obozrenie

‑

316

. [In

Russian

Bey-Bienko GY, Mistshenko LL (1951)

Keys to the Fauna of the U.S.S.R. [1964 English translation no. 40]

. In: Staff TaEbI, Technical Editorial Consultant: Robert Lathan Randell MRAAoNSoPPP,USA (Eds)

Locusts and Grasshoppers of the U.S.S.R. and Adjacent Countries.

Zoological Institute of the U.S.S.R. Academy of Sciences

Moskva, Leningrad

385-667[1-291]

pp. [In

Russian

]. URL: https://www.biodiversitylibrary.org/item/224097#page/9/mode/1up

Bi D (1986)

Description of five new grasshoppers from China (Orthoptera: Acridoidea)

Contributions from the Shanghai Institute of Entomology

195

‑

206

Bolívar I (1898)

Contributions à l'étude des Acridiens espèces de la Faune indo et austro-malaisienne du Museo Civico di Storia Naturale di Genova

Annali del Museo Civico di Storia Naturale di Genova

219

(

‑

101

. https://doi.org/10.5962/bhl.part.9541

Bolívar I (1901)

In Zichy

Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy

223

‑

243

Cejchan A, Maran J (1966)

Orthoptera aus der Mongolischen Volksrepublik. Zugleich ergebnisse der Mongolisch-Deutschen biologischen expedition seit 1962. Nr. 11

Mitteilungen aus dem Zoologischen Museum in Berlin

(

177

‑

195

. [In

German

Childebaev MK, Storozhenko SY (2001)

An annotated list of brachycerous orthopterous insects (Orthoptera: Caelifera) occurring in Kazakhstan

Tethys Entomological Research

‑

Chogsomzhav L, Shurovenkov (1963)

Fauna of grasshoppers (Orthoptera, Acrididae) of the Mongolian People's Republic. - 5th meeting

Fauna of grasshoppers (Orthoptera, Acrididae) of the Mongolian People's Republic

USSR Academy of Sciences

Proceedings of the All-Union Entomological

61-63

pp.

Chogsomzhav L (1968)

The distribution of Mongolian grasshopper (Acrididae, Orthoptera).

Journal of University of Agriculture, Mongolia

‑

. [In

Mongolian

Chogsomzhav L (1969a)

New record of orthopteroid insect

News of Mongolian Academy of Science 3:76-80

Chogsomzhav L (1969b)

Study of Orthoptera

Proceedings of Institute of Biology

123

‑

130

. [In

Mongolia

Chogsomzhav L (1970)

Orthoptera in the basin of the great lakes of the Mongolian People's Republic (Orthoptera).

Proceedings of Institute Biology

169

‑

177

. [In

Mongolian

Chogsomzhav L (1971)

Acridoidea and Tettigonioidea of Mongolian People's Republic.

Insects of Mongolia

‑

108

. [In

Russian

Chogsomzhav L (1972)

Acridoidea and Tettigonioidea of the Mongolian People’s Republic.

Insects of Mongolia

151

‑

198

. [In

Russian

Chogsomzhav L (1974a)

A new species of the genus Mongolotettix Rehn (Orthoptera, Acrididae) from Mongolia.

Entomological Review

‑

. [In

Russian

Chogsomzhav L (1974b)

Orthopteroid insects (Orthopteroidea) of western and southern Mongolia.

[

Insects of Mongolia

Nauka

Leningrad

23-33

pp. [In

Russian

Chogsomzhav L (1975)

Orthopteroidea collected by the entomological group of the Soviet-Mongolian complex biological expedition in the year 1971

Insects of Mongolia

(

‑

. [In

Russian

Chogsomzhav L (1977)

Orthopteroidea of the Gobi Desert.

Insects of Mongolia

‑

. [In

Russian

Chogsomzhav L (1989)

Composition and distribution of fauna of the Orthopteroidea in the Mongolian People's Republic

Insects of Mongolia

‑

. [In

Russian

Chopard L (1949)

Note sur les Orthoptéroïdes du Sahara marocain

Bulletin of the Society of Natural Sciences of Morocco

25-27 [1945-1947]

191

‑

199

Cigliano M, Braun H, Eades D, Otte D (2022)

Orthoptera species file. Version 5.0/5.0.

URL: http://Orthoptera.SpeciesFile.org

Creutzer C (1799)

Entomologische Versuche

Wien: Bey Karl Schaumburg und Comp., 1799

142

pp. URL: http://www.biodiversitylibrary.org/item/41692#3

De Geer (1773)

Mémoires pour servir à l'histoire des insectes, Pierre Hesselberg, Stockholm

L.L. Grefing, 1773

696

pp.

Dey L, Seidel M, Lchagvasuren D, Husemann M (2021)

From the steppe to the desert: Survey of band-winged grasshoppers from Mongolia (Orthoptera: Acrididae: Oedipodinae) based on material from 50 Years of expeditions

Erforschung Biologischer Ressourcen der Mongolei/ Exploration into the Biological Resources of Mongolia

(

)

329

‑

360

. URL: https://digitalcommons.unl.edu/biolmongol/262

Fang J, Bai Y, Wu J (2015)

Towards a better understanding of landscape patterns and ecosystem processes of the Mongolian Plateau

Landscape Ecology

(

1573

‑

1578

. https://doi.org/https://doi.org/10.1007/s10980-015-0277-2

Fartmann T, Krämer B, Stelzner F, Poniatowski D (2012)

Orthoptera as ecological indicators for succession in steppe grassland

Ecological Indicators

337

‑

344

. https://doi.org/10.1016/j.ecolind.2012.03.002

Fieber FX (1853)

Synopsis der Europaischen Orthopteren mit besonderer Rücksicht der Böhmischen Arten.

Lotus (Prag)

(

separate issued 1854, Prag (Gerzabek), pp.1-79 + (index) I-IV

90-104, 115-131, 138 154, 168-176, 184-188, 201-207, 232-238, 252-261

Fischer LH (1849)

Beitraege zur Insekten-fauna Freiburgs, Orthoptera

Jahresbericht, Mannheimer Verein für Naturkunde

‑

Fischer von Waldheim G (1836)

Orthoptera duo e montibus Catunicis descripta et icone illustrata.

Bulletin de la Société Impériale des Naturalistes de Moscou

346

‑

349

Fischer von Waldheim G (1846)

Entomographia Imperii Rossici. IV. Orthoptera Imperii Rossici.

Nouveaux Mémoires de la Société Impériale des Naturalistes de Moscou

(

i-iv

1–443

‑

pls 1–37

Furukawa H (1929)

Some alpine orthopterans from Mt. Ogahana, with description of a new subspecies.

Kontyu

165

‑

178

Furukawa H, Shiraki T, Omachi F (1950)

In iconographia insectorum Japonicorum.

In: Shiraki T (Ed.)

Orthoptera

Hokuryukan, Tokyo

22-52

pp. [In

Japanese

Ganbold Y (2009)

Study of medicinal properties from raw materials of wide distributed Mongolian Acridoidea grasshopper and to produced preparation technology “Acritract”

Mongolian Agricultural University

Ulaanbaatar, Mongolia

. URL: http://data.stf.gov.mn/Publication/Thesis/ThesisViewPublic.aspx?id=129150

Gandulam R (2016)

Species composition of grasshopper (Acrididae) in the Khentii province

Mongolian Agricultural University

Ulaanbaatar

pp. [In

Mongolian

Garai A (2001)

Orthopteroid insects of the Adrienne Garai and Péter Gyulai expedition to Mongolia in 1997

Esperiana

747

‑

754

Garai A (2010)

Contribution to the knowledge of the Iranian Orthopteroid insects I

Esperiana

393

‑

418

Gmelin JF (1789)

Caroli a Linné, Systema naturae per regna tria naturae, secundum classes, ordines, genera, species; cum characteribus, differentiis, synonymis, locis

Editio decimo tertia, aucta, reformata. G.E. Beer, Lipsiae [= Leipzig]

(

1517

‑

2224

. URL: http://www.biodiversitylibrary.org/item/10288#page/1/mode/1up

Gombobaatar S, Myagmarsuren S, Conaboy N, Мunkhjargal M (2014)

Convention on biological diversity: The 5th national report of Mongolia

. URL: https://www.cbd.int/doc/world/mn/mn-nr-05-en.pdf

Gorochov AV, Mistshenko LL, Podgornaya LI (1989)

Materials on the fauna and ecology of Orthoptera of the Transaltai Gobi

Nasekomye Mongolii [Insects of Mongolia]

‑

117

Günther KK (1971)

Blattoidea-Orthopteroidea-Ausbeute 1964, Teil II (Tetrigidae und Acrididae). Ergebnisse der Mongolisch-Deutschen Biologischen Expeditionen seit 1962, Nr. 55.

Mitteilungen aus dem Zoologischen Museum in Berlin

(

109

‑

130

. https://doi.org/10.1002/mmnz.19710470113

Gupta VK (1983)

The locust and grasshopper agricultural manual 1982

Oriental Insects

(

‑

126

. https://doi.org/10.1080/00305316.1983.10433700

Herbert H, Nikolai F, Karl-Georg B, Barbara NS, Smirnov Alexander S, Frank B (2019)

The Eurasian steppe belt: Status quo, origin and evolutionary history

Turczaninowia

(

‑

. https://doi.org/10.14258/turczaninowia.22.3.1

Hewitt G (1977)

Review of forage losses caused by rangeland grasshoppers

Agricultural Research Service, United States.

Department of Agriculture

Washington District of Columbia

Miscellaneous Publications No. 1348.24

pp. URL: https://doi.org/10.5962/bhl.title.65355

Hodjat SH, Tork M, Seiedy M, Defaut B (2018)

A taxonomic review of recorded species of Caelifera (Orthoptera) in Iran

Matériaux orthoptériques et entomocénotiques

‑

Huang C (1981)

Orthoptera: Acrididae, Catantopinae, Pyrgomorphinae, Oedipodinae. In: Insects of Xizang

Science Press, Beijing

‑

Ikonnikov N (1911)

Zur Kenntnis der Acridiodeen Sibiriens.

Annuaire du Musée Zoologique de l'Académie Impériale des Sciences de Saint-Pétersbourg

242

‑

270

. URL: http://www.biodiversitylibrary.org/item/34698#page/280/mode/1up

IUCN (2022)

The IUCN red list of threatened species. Version 2022-September.

URL: https://www.iucnredlist.org. Accessed on [Feb,2022].

Ivanova IV (1967)

On the fauna of Orthoptera of the southern part of the Krasnojarsk Region, Central Siberia.

Entomologicheskoe Obozrenie

(

127

‑

138

Jacobson GG, Bianchi. VL (1905)

Orthopteroid and pseudoneuropteroid insects of the Russian empire and adjacent countries

pls 1–25

Cambridge University Press

1–952

pp. URL: https://doi.org/10.4039/Ent34130-5

Jin X (1994)

Genus Haplotropis Saussure. In Xia, K.-L. & et al. Acridoidea: Pamphagidae, Chrotogonidae, Pyrgomorphidae.

Fauna Sinica, Insecta

249

‑

252, 319

Karny HH (1908)

Orthoptera. A Dictyoptera, Tettigonioidea, Acridoidea

Wissenschaftliche Ergebnisse der Expedition Filchner nach China und Tibet 1903-1905.

(

‑

. URL: http://books.google.com/books?id=GbKCAAAAIAAJ

Kietzka, Lecoq GJ, Samways M, J M (2021)

Ecological and human diet value of locusts in a changing world

Agronomy

(

1856

. https://doi.org/10.3390/agronomy11091856

Kittary M (1849)

Orthoptères observés dans les steppes des Kirguises par MM. le Professeur P. Wagner et le Docteur Kittary, en 1846, déterminés et décrits

Bulletin de la Société Impériale des Naturalistes de Moscou

437

‑

479

. URL: http://books.google.com/books?id=aioWAQAAIAAJ

Krauss HA (1901)

Orthopteren vom Kuku-nor-Gebiet in Centralasien, gesammelt von Dr. J. Holderer im Jahre 1898.

Zoologischer Anzeiger

235

‑

239

. URL: http://www.biodiversitylibrary.org/item/37592#7

Latchininsky A, Sword G, Sergeev M, Cigliano MM, Lecoq M (2011)

Locusts and Grasshoppers: Behavior, Ecology, and Biogeography

Psyche-A journal of Entomology

2011

4pp

. https://doi.org/10.1155/2011/578327

Lecoq M, Zhang L (2019)

Encyclopedia of pest Orthoptera of the world

China Agricultural University Press

311

pp. [ISBN

978-7-5655-2232-1

]

Lecoq M, Cease A (2022)

What Have We Learned after Millennia of Locust Invasions?

Agronomy

(

472

. https://doi.org/10.3390/agronomy12020472

Lee HS, Lee C (1985)

Taxonomic revision of the Catantopinae from Korea (Orthoptera: Acrididae) IV. Calliptamini and Eyprepocnemidini.

The Korean Journal of Entomology

(

‑

Lee HS, Park. WH (1992)

A new genus of the subfamily Oedipodinae (Orthoptera: Acrididae)

Basic Science Research Institute of Hyosung Women's University

‑

Lian Z, Zheng Z (1984)

New genera and new species of grasshoppers from Gansu, China.

Entomotaxonomia

(

299

‑

311

Li H (1981)

Studies on the fauna of the genus Angaracris B.-Bienko (Orthoptera: Acrididae)

Acta Zootaxonomica Sinica

(

167

‑

173

Li H, Hao. S, Kang. L (2007)

Regional differentiation of the Acridoidea ecofaunas in different vegetational zones (subzones) of Inner Mongolia region

Acta Entomologica Sinica

(

361

‑

375

. URL: http://www.insect.org.cn/EN/abstract/abstract9838.shtml

Lockwood DR, Lockwood JA (2008)

Grasshopper population ecology: catastrophe, criticality, and critique

Ecology and Society

(

. https://doi.org/10.5751/ES-02496-130134

Marini L, Fontana P, Scotton M, Klimek S (2008)

Vascular plant and Orthoptera diversity in relation to grassland management and landscape composition in the European Alps

Journal of Applied Ecology

(

361

‑

370

. https://doi.org/10.1111/j.1365-2664.2007.01402.x

Matsumura S (1911)

Erster Beitrag zur Insekten-Fauna von Sachalin.

The Journal of the College of Agriculture, Tohoku Imperial University, Sapporo, Japan

(

‑

145

. URL: https://eprints.lib.hokudai.ac.jp/dspace/bitstream/2115/12498/1/4(1)_p1-145.pdf

Miksic S (1981)

Mitteleuropäische und mediterrane Orthopteren in der Fauna des herzegowinischen Karstes.

Acta Entomological Jugoslavica

‑

Mistshenko LL (1937)

Revision of palaearctic species of the genus Sphingonotus Fieber (Orthoptera: Acrididae)

Eos, Revista Española de Entomología

(

3-4

‑

282

. URL: https://archive.org/details/eos-0013-9440-147638

Mistshenko LL, Bey-Bienko GY (1951)

Keys to the fauna of the U.S.S.R. [1964 English translation no. 40]

Locusts and grasshoppers of the U.S.S.R. and adjacent countries

385

‑

667[1-291]

. URL: http://www.biodiversitylibrary.org/item/224097#page/7/mode/1up

Mistshenko LL (1952)

Locusts and grasshoppers, Catantopinae.

Fauna of the U.S.S.R.

(

79-80,

‑

96-97,610, 520 figs

Mistshenko LL (1968)

Orthopteroid insects (Orthopteroidea) collected by the entomological expedition of the zoological institute, USSR academy of sciences in the Mongolian People’s Republic in 1967.

Entomological Review

482

‑

498

Mistshenko LL (1973)

Grasshoppers of the genus Eclipophleps S. Tarb. (Orthoptera: Acrididae)

Entomological Review

(

‑

107

Mistshenko LL, Gorochov AV (1989)

In Gorochov, Mistshenko & Podgornaya. Materials on the fauna and ecology of Orthoptera of the Transaltai Gobi

Nasekomye Mongolii [Insects of Mongolia]

‑

117

. URL: http://gryllus.smart21.ru/gryllus/Gorochov%27s%20papers%20in%20PDF%20and%20list%20of%20his%20publications/1989%20%2846-49%29/46%20Orthoptera%20Transaltai%20Gobi%20%5BInsects%20of%20Mongolia.%20Leningrad.%20ZIN%20AN%20SSSR%5D%201989.pdf

Mohamed, A, Kimura R (2014)

Applying the moisture availability index (NTDI) over vegetated land in Central Asia: Mongolian steppe.

Journal of Water Resource and Protection

1335

‑

1343

. https://doi.org/10.4236/jwarp.2014.614123

Munkhbat J (2010)

Study of Orthoptera insect communities from Hustai National Park

ESPA is an electronic archive of research works and papers of biological science of Mongolia

. URL: https://biology.mn/?p=product&viewby=single&id=112#description

Myagmar G, Dorzhiev TZ, Gantigmaa C (2019)

The fauna of orthopteran insects of the Galba desert in the south eastern Mongolia

Regional problems of ecology and wildlife protection

Ulan-Ude

1^st-2^nd Feb, 2019

Buryat State University Publishing Department

246

pp. [In

Russian

]. URL: https://my.bsu.ru/content/pbl/publications/publication_64.pdf

Nonnaizab N, Qi B, Li Y (1999)

Insects of Inner Mongolia China

Inner Mongolia People’s Publishing House

Hohhot

506

pp. [In

Chinese

Olivier GA (1791)

Criquet, Acrydium. In: Olivier M, Encyclopédie méthodique. Histoire naturelle. Insectes

Chez Panckoucke

Panckoucke, Paris

204-236

pp. URL: https://books.google.co.kr/books?id=8IIPAAAAQAAJ

Orishchenko. (1960)

Eclipophleps tarbinskii is een rechtvleugelig insect uit de familie veldsprinkhanen (Acrididae).

De wetenschappelijke naam van deze soort is voor het eerst geldig gepubliceerd in 1960 door Orishchenko

Otgonchimeg T (2017)

Study of grasshopper (Acrididae) species composition and distribution patterns of forest-steppe zone of Mongolia

Mongolian Agricultural University

Ulaanbaatar, Mongolia

pp. URL: http://data.stf.gov.mn/Publication/Thesis/ThesisViewPublic.aspx?id=128628

Podgornaya LI, Gorochov AV (1989)

In Gorochov, Mistshenko & Podgornaya. Materials on the fauna and ecology of Orthoptera of the Transaltai Gobi.

Nasekomye Mongolii [Insects of Mongolia]

‑

117

Popova K, Molodtsov V, Sergeev M (2020)

Rare grasshoppers (Orthoptera, Acridoidea) of the Baraba and Kulunda steppes (South Siberia)

Acta Biologica Sibirica

595

‑

609

. https://doi.org/10.3897/abs.6.e59519

Predtechenskii (1928)

Notes of the Astrakhan plant protection station against pests

Russian State Library

Astrakhan

116

pp. [In

Russian and German

Pylnov E (1916)

Contributions à la faune des Acridoidea et des Locustodea de la Mongolie boréale.

Russian Entomological Review

275

‑

284

Qian H, Altanchimeg D, Naizab N, Wang S, Wen S, Lin C (2021)

The complete mitochondrial genome of Eclipophleps carinata (Orthoptera: Acridoidea: Gomphoceridae)

Mitochondrial DNA Part B

(

1310

‑

1312

. https://doi.org/10.1080/23802359.2021.1907803

Ramme W (1939)

Beiträge zur Kenntnis der palaearktischen Orthopteren fauna (Tettig. u. Acrid.) III

Mitteilungen aus dem Zoologischen Museum in Berlin

‑

150

Ramme W (1952)

Orthopteren der Sven Hedin-Expedition nach China 1927-30. Ergänzungen und Berichtigungen zur Bearbeitung durch Y. Sjöstedt 1933.

Archive for Zoology

(

)

‑

Redtenbacher J (1889)

Beitrag zur Orthopteren-Fauna von Turkmenien.

Wiener Entomologische Zeitschrift

‑

Rentsendorj G, Khodroi B (2020)

Study review of the composition grasshoppers in Mongolia

Mongolian Journal of Agricultural Sciences

(

‑

. https://doi.org/10.5564/mjas.v29i1.1375

Saussure H (1884)

Prodromus oedipodiorum insectorum ex ordine orthopterorum

28 (6)

Smithsonian Libraries

Georg H, Geneva

254

pp. https://doi.org/10.5962/bhl.title.9514

Saussure H (1888)

Additamenta ad prodromum Oedipodiorum

pl. 2

30(1)

Imprimerie Charles Schuchardt

182

pp. URL: http://www.biodiversitylibrary.org/item/50100#page/71/mode/1up

Sergeev MG (1995)

The general distribution of Orthoptera in the eastern parts of the Saharan-Gobian and Scythian subregions.

Acta Zoologica Cracoviensia

(

213

‑

256

. URL: http://www.isez.pan.krakow.pl/journals/azc/pdf/azc_i/38(2)/38(2)_06.pdf

Sergeev MG, Jirong MM, N.E. H (2009)

Diversity and distribution patterns of Orthoptera in the Altai Mountains.

Amurian Zoological Journal

(

106

‑

112

. [In

Russian

]. https://doi.org/10.33910/1999-4079-2009-1-2-106-112

Sergeev MG, Storozhenko SY, Benediktov AA (2019)

An annotated check-list of Orthoptera of Tuva and adjacent regions. Part 2. Suboder Caelifera. Tridactylidae, Tetrigidae, Acrididae: Melanoplinae, Calliptaminae, and Gomphocerinae (except Gomphocerini).

Far Eastern Entomologist

389

‑

. https://doi.org/10.25221/fee.389.2

Sergeev MG, Storozhenko SY, Benediktov AA (2020)

An annotated check-list of Orthoptera of Tuva and adjacent regions. Part 3. Suboder Caelifera (Acrididae: Gomphocerinae: Gomphocerini, Locustinae).

Far Eastern Entomologist

402

‑

. https://doi.org/10.25221/fee.402.1

Sergeev MG (2021)

Distribution patterns of grasshoppers and their kin over the Eurasian steppes

Insects

(

. https://doi.org/10.3390/insects12010077

Song H (2010)

Grasshopper Systematics: Past, Present and Future

Journal of Orthoptera Research

(

‑

68, 12

. https://doi.org/10.1665/034.019.0112

Stål C (1861)

Orthoptera species novas descripsit.

Kongliga Svenska Fregatten Eugenies Resa Omkring Jorden under befäl af C.A. Virgin åren 1851-1853 (Zoologi)

(

299

‑

350

. URL: http://www.biodiversitylibrary.org/item/20004#page/311/mode/1up

Steinmann H (1964)

Ergebnisse der zoologischen Forschungen von Dr. Z. Kaszab in der Mongolei 20. Tetrigidae und Acrididae (Orthoptera).

Folia Entomologica Hungarica

381

‑

384

. URL: http://publication.nhmus.hu/folent/cikkreszletes.php?idhoz=5529

Steinmann H (1967)

Tetrigidae and Acridiidae. Ergebnisse der zoologischen Forschungen von Dr. Z. Kaszab in der Mongolei 99. (Orthoptera).

Reichenbachia

107

‑

120

Steinmann H (1968)

Tetrigidae und Acridiidae. Ergebnisse der zoologischen Forschungen von Dr. Z. Kaszab in der Mongolei (Orthoptera).

Reichenbachia

239

‑

248

Steinmann H (1971)

Tetrigidae und Acrididae. Ergebnisse der zoologischen Forschungen von Dr. Z. Kaszab in der Mongolei (Orthoptera).

Faunistische Abhandlungen. Staatliches Museum für Tierkunde in Dresden

145

‑

157

Stoll C (1813)

Représentation exactement colorée d'après nature des spectres ou phasmes, des mantes, des sauterelles, des grillons, des criquets et des blattes, qui se trouvent dans les quatre parties du monde, l'Europe, l'Asie, l'Afrique et l'Amerique

JC Sepp et Fils

Amsterdam

. URL: http://books.google.com/books?id=mTlnAAAAcAAJ

Storozhenko SY (1985)

New representatives of the genus Stenobothrus (Orthoptera, Acrididae) from the Sovjet Far East.

Zoologicheskij Zhurnal

144

‑

148

. [In

Russian, English abstract

Storozhenko SY (1986)

Revision of the genus Zubovskya Dov.-Zap. (Orthoptera, Acrididae).

Zoological Institute (Leningrad), Academy of Sciences SSSR

143

‑

Storozhenko SY, Paik J-C (2007)

Orthoptera of Korea

Dalnauka, Vladivostok

232

pp.

Storozhenko SY, Kim TW, Jeon M (2015)

Monograph of Korean Orthoptera

National Institute of Biological Resources

Incheon

. [ISBN

978-89-6811-229-4 93490

]

Sulzer JH (1776)

Abgekuerzte Geschichte der Insecten nach dem Linnei’schen System

Steiner, Winterthur

Switzerland

71 pp + 32 plates [plate 8-9 orthoptera, discussed in Fuessly C., 1778]

pp.

Tarbinsky SP (1925)

Zur Kenntnis der Gattung Chorthippus Fieb. (Orthoptera: Acridinae)

Konowia

135

‑

140

. URL: http://www.biologiezentrum.at/pdf_frei_remote/KON_4_0135-0140.pdf

Tarbinsky SP (1927)

On some new and little-known Orthoptera from Palaearctic Asia.

Annals and Magazine of Natural History

(

119

489

‑

502

. https://doi.org/10.1080/00222932708655478

Tarbinsky SP (1931)

A revision of the Palaearctic species of the genera Gomphocerus Thunb. and Dasyhippus Uvar. (Acrididae)

Bulletin of the Leningrad Institute for Controlling Farm and Forest Pests.

127

‑

157

. [In

Russian

Tishechkin DY, Bukhvalova MA (2009)

New data on and calling signal of Gomphocerinae grasshopper (Orthoptera: Acrididae) from South Siberia and Russian Far East

Russian Entomological Journal

‑

Tsyplenkov E (1956)

A new genus of the tribe Thrinchini (Orthoptera, Acrididae) from western China

Entomologicheskoe Obozrenie, Moscow

883

‑

885

Uchida K, Ushimaru A (2014)

Biodiversity declines due to abandonment and intensification of agricultural lands: patterns and mechanisms

Ecological Monographs

(

637

‑

658

. https://doi.org/10.1890/13-2170.1

Ünal (2016)

Pamphagidae (Orthoptera: Acridoidea) from the Palaearctic Region: taxonomy, classification, keys to genera and a review of the tribe Nocarodeini I. Bolívar.

Zootaxa

4206

(

). https://doi.org/10.11646/zootaxa.4206.1.1

Uvarov B (1910)

Contributions sur la faune des Orthoptères de la province de l’Oural

Horae Societatis entomologicae Rossicae

367

‑

390

Uvarov BP (1914)

Contributions to the fauna of orthoptera in the province of Transbaikalia.

Directory of the Zoological Museum of the Imperial Academy of Sciences in Saint Petersburg.

167

‑

172

. URL: http://www.biodiversitylibrary.org/item/105349#page/191/mode/1up

Uvarov BP (1934)

Studies in the Orthoptera of Turkey, Iraq and Syria.

Revista Española de Entomología

‑

119

Werner F (1905)

Ergebnisse einer zoologischen Forschungsreise nach Ägypten und dem ägyptischen Sudan. I. Die Orthopterenfauna Ägyptens mit besonderer Burücksichtigung der Eremiaphilen

Sitzungsberichte der kaiserlichen Akademie der Wissenschaften, Mathematisch -naturwissenschaftliche Klasse, Wien

114

(

357

‑

436

. URL: http://www.biodiversitylibrary.org/item/31082#page/403/mode/1up

Wheeler QD (2004)

Taxonomic triage and the poverty of phylogeny

Philosophical Transactions of the Royal Society of London. Series B: Biological Sciences

359

(

1444

571

‑

583

. https://doi.org/10.1098/rstb.2003.1452

Willemse LM (2009)

In Çiplak, K.-G. Heller & F.M.H. Willemse. Review of the genus Eupholidoptera (Orthoptera, Tettigoniidae): different genitalia, uniform song.

Zootaxa.

2156

‑

. https://doi.org/10.11646/zootaxa.2156.1.1

Wnukowskij W (1926)

Zur Fauna der Orthopteren und Dermapteren des Bezirks Kamenj (südwestliches Sibirien, früheres Gouvernement Tomsk)

Mitteilungen der Münchner Entomologischen Gesellschaft

‑

Worden R, Savada AM (1991)

Mongolia: A country study

2d ed

Federal Research Division Library of Congress

320

pp. URL: https://apps.dtic.mil/sti/pdfs/ADA233772.pdf

Yembuu Be (2021)

The physical geography of Mongolia

Springer

218

pp. [ISBN

9783030614331 (hardcover)

] https://doi.org/10.1007/978-3-030-61434-8

Yin X-C, Chou. (1979)

Two new genera and three new species of Acrididae from Shaanxi province.

Entomotaxonomia

(

125

‑

130

Yin X-C, Wang W (2005)

A new species of Compsorhipis Saussure (Orthoptera, Acrididae, Oedipodinae), with a key to the known species from China and adjacent areas

. URL: http://www.biodiversitylibrary.org/item/113865#page/31/mode/1up

Zhang D, Wang W, Yin X (2006)

A new species of Bryodema (Orthoptera: Acridoidea) from China, with a key to the described species

Entomological News

117

‑

. https://doi.org/10.3157/0013-872X(2006)117[11:ANSOBO]2.0.CO;2

Zhang L, Lecoq L, Latchininsky M, Hunter A, David (2019)

Locust and grasshopper management

Annual Review of Entomology

(

‑

. https://doi.org/10.1146/annurev-ento-011118-112500

Zheng Z, Ren. G (1993)

Four new species of grasshoppers from northern west of China (Orthoptera: Acridoidea).

Journal of Hubei University (Natural Science).

(

424

‑

429

Zheng Z, He. D (1994)

Two new species of grasshoppers from Ningxia (Orthoptera: Acridoidea).

Journal of Hubei University (Natural Science).

(

308

‑

311

Zheng Z, Han. Y (1998)

Two new species of grasshoppers (Orthoptera: Acridoidea) from Nei Mongol.

Entomotaxonomia

(

‑

Zheng Z, Zeng H, Zhang H, Tao S, Su. S (2012)

A survey of grasshoppers from Helan mountain national reserve in Inner Mongolia (Orthoptera).

Journal of Shaanxi Normal University (Natural Science Edition)

(

‑

Zubovski N (1900)

Beitrag zur Kenntniss der sibirischen Acridiodeen

Trudy Russkago Entomologicheskago Obshchestva [Horae Societatis Entomologicae Rossicae].

‑

. URL: http://www.biodiversitylibrary.org/item/46209#page/105/mode/1up

Supplementary materials

Suppl. material 1: Species list of grasshoppers’ geographical natural distribution in six types of zones

Authors: Altanchimeg

Data type: table

Brief description:

Captions: (*) = Endemic of Mongolia, (+) = geographical distribution natural zone, (★) = Indicator species of geographical natural zones, (-) = poor species. Abbreviation: 1-High mountain, 2-Taiga, 3-Forest steppe, 4-Steppe, 5-Desert steppe, 6-Desert natural zone.

Download file (49.45 kb)

Suppl. material 2: Species registered in the European Red List of grasshopper

Authors: Enkhtsetseg

Data type: table

Download file (16.54 kb)

Suppl. material 3: Shannon and Berger Parker’s index of natural zone

Authors: Enkhtsetseg

Data type: table

Download file (14.12 kb)

Suppl. material 4: Shannon and Berger-Parker index of neighbouring boundary countries grasshopper distribution

Authors: Enkhtsetseg

Data type: table

Download file (13.85 kb)

Endnotes