Biodiversity Data Journal :
Taxonomic Paper
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Corresponding author:
Academic editor: Pavel Stoev
Received: 05 Feb 2016 | Accepted: 30 Mar 2016 | Published: 07 Apr 2016
© 2016 George Else, Barry Bolton, Gavin Broad
This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
Citation:
Else G, Bolton B, Broad G (2016) Checklist of British and Irish Hymenoptera - aculeates (Apoidea, Chrysidoidea and Vespoidea). Biodiversity Data Journal 4: e8050. https://doi.org/10.3897/BDJ.4.e8050
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The checklist of British and Irish aculeate Hymenoptera (Apoidea, Chrysidoidea and Vespoidea) is revised. Species distribution is summarised for all species at the level of country (England, Scotland, Wales, Ireland and Isle of Man).
The 601 native species represent an increase of 25 on the 1978 checklist, comprising mostly new discoveries. This increase is nearly balanced by the 23 species now presumed to be extinct in Britain and Ireland.
Britain, Ireland, bees, ants, wasps, fauna
The checklist of British and Irish aculeates is essentially that of
Unlike most groups of Hymenoptera, the aculeate ants, bees and wasps are well-served by identification literature and are actively collected and recorded by a relatively large number of amateur as well as professional entomologists. We thus have a better idea of the actual size of the fauna than we do for sawflies and parasitoid wasps. There is thus a much better idea of change in the aculeate fauna and numerous extinctions and colonisations have been documented.
The aculeate Hymenoptera encompass three superfamilies and 18 families in Britain and Ireland (as discussed in
Representative British crabronid and sphecid wasps
Representative British bees
Representative British Vespoidea
Representative British Vespoidea
The rationale and background to the updated Hymenoptera checklist are laid out in
Sources of classification are listed under each superfamily. The checklist of bees is essentially similar to that of Else's (in prep.) handbook to the British fauna, except that we employ six families rather than including all bees in Apidae and, as discussed in the introduction to this checklist series (
Country-level distribution data mostly derive from the distribution atlases produced by the Bees, Wasps and Ants Recording Society (
The following abbreviations and conventions are employed:
[species] taxon deleted from the British and Irish list
NHM Natural History Museum, London
# known or suspected introductions with at least temporarily self-sustaining populations
? status (including uncertain synonymy) or identification in the British Isles uncertain
misident. has been misidentified as this name
nom. dub. nomen dubium, a name of unknown or doubtful application
nom. ob. nomen oblitum, ‘forgotten name’, does not have priority over a younger name
nom. nov. nomen novum, a replacement name
nom. nud. nomen nudum, an unavailable name, with no type specimen
preocc. name preoccupied (junior homonym)
stat. rev. status revocatus, revived status (e.g., raised from synonymy)
unavailable not meeting the requirements of the International Code of Zoological Nomenclature
var. variety, only available as a valid name under the provisions of article 45.6 of the ICZN
Text and spreadsheet versions of the checklists are available as supplementary files: Suppl. materials
The higher-level classification of the Apoidea follows
DIMORPHA Panzer, 1806
Sphex boops Schrank, 1781
abdominalis (Panzer, 1798, Tiphia)
pompiliformis (Panzer, 1804, Larra)
oculata (Jurine, 1807, Dimorpha)
victor Curtis, 1829
vanderlindeni Robert, 1833
agilis Smith, 1875
Larra pinguis Dahlbom, 1832
stigma misident.
pinguis (Zetterstedt, 1838, Larra) preocc.
jaculator (Smith, 1845, Astata)
Pompilus lunicornis Fabricius, 1798
kenedii (Curtis, 1836, Alyson)
Gorytes fargeii Shuckard, 1837
campestris misident.
mongolensis Tsuneki, 1971
przewalskyi Kazenas, 1971
Sphex mystaceus Linnaeus, 1761
campestris (Linnaeus, 1761, Vespa)
inimica (Harris, 1776, Vespa)
longicornis (Rossi, 1790, Sphex)
bicinctus (Fabricius, 1793, Crabro)
arpactus (Fabricius, 1804, Mellinus)
flavicincta (Donovan, 1808, Vespa)
croceipes (Eversmann, 1849, Gorytes)
tonsus (Bondroit, 1933, Gorytes)
ARPACTUS Panzer, 1805
EUZONIA Stephens, 1829
EUSPONGUS Lepeletier, 1832
HOPLISUS Lepeletier, 1832
LAEVIGORYTES Zavadil, 1948
Euspongus laticinctus Lepeletier, 1832
Mellinus quadrifasciatus Fabricius, 1804
vicinus (Lepeletier, 1832, Euspongus)
montivagus (Mocsáry, 1878, Hoplisus)
ARPACTUS Jurine, 1807
HARPACTES Dahlbom, 1843
DIENOPLUS Fox, 1893
Pompilus tumidus Panzer, 1801
japonensis (Tsuneki, 1963, Dienoplus)
transiliensis Kazenas, 1989
HYPOMELLINUS Asmead, 1899
MELLINOGASTRA Asmead, 1899
Crabro bicinctus Rossi, 1794
SYNNEVRUS Costa, 1859
wesmaeli Lepeletier, 1845
distinguendus Chevrier, 1867
Mellinus interruptus Fabricius, 1798
spinosus (Fabricius, 1804, Ceropales)
panzeri Lepeletier, 1845
shukardi Wesmael, 1852
Sphex spinosus Forster, 1771
?bidens (Linnaeus, 1767, Vespa)
spinosus (Fabricius, 1775, Crabro) preocc.
tricinctus (Fabricius, 1793, Mellinus)
trilineata (Turton, 1802, Vespa)
geniculatus Lepeletier, 1845
malasei Gussakovskij, 1932
Crabro trimaculatus Rossi, 1790
THYREOPUS Lepeletier & Brullé, 1835
ANOTHYREUS Dahlbom, 1845
THYREOCNEMUS Costa, 1871
PARANOTHYREUS Ashmead, 1899
SYNOTHYREOPUS Ashmead, 1899
AGNOSICRABRO Pate, 1944
DYSCOLOCRABRO Pate, 1944
HEMITHYREOPUS Pate, 1944
NORUMBEGA Pate, 1947
PARANOTHYREUS Pate, 1944
PARATHYREOPUS Pate, 1944
PEMPHILIS Pate, 1944
OTHYREUS Marshakov, 1977
Vespa cribraria Linnaeus, 1758
patellarius (Schreber, 1784, Sphex)
argus (Christ, 1791, Sphex)
longa (Christ, 1791, Sphex)
lunatus (Christ, 1791, Sphex)
palmatus Panzer, 1797
inornatus Mocsáry, 1901
hypotheticus Kokujev, 1927
Sphex peltarius Schreber, 1784
patellatus Panzer, 1797
dentipes Panzer, 1797
mediatus Fabricius, 1798
Sphex scutellatus von Scheven, 1781
scutularius (Schreber, 1784, Sphex)
pterotus Panzer, 1801
reticulatus (Lepeletier & Brullé, 1835, Ceratocolus)
Crabro congener Dahlbom, 1844
added by
Crabro podagricus Vander Linden, 1829
vicinus Dahlbom, 1842
punctata Šnoflak, 1948
snoflaki Zavadil, 1948
CORENOCRABRO Tsuneki, 1974
Crabro vagabundus Panzer, 1798
varus (Panzer, 1799, Crabro) preocc.
bojus (Schrank, 1802, Crabro)
quinquemaculatus (Lepeletier & Brullé, 1835, Blepharipus)
lefebvrei Lepeletier & Brullé, 1835
fasciatus (Costa, 1871, Crabro)
esakii (Yasumatsu, 1942, Crabro)
ectemiformis (Tsuneki, 1974, Corenocrabro)
COELOCRABRO Thomson, 1874
Blepharipus annulipes Lepeletier & Brullé, 1835
gonager (Lepeletier & Brullé, 1835, Blepharipus)
nigritus (Gimmerthal, 1836, Crabro)
ambiguus (Dahlbom, 1842, Crabro)
capito (Dahlbom, 1845, Crabro)
parkeri (Banks, 1921, Blepharipus)
davidsoni (Sandhouse, 1938, Crabro)
Crabro capitosus Shuckard, 1837
annulus (Dahlbom, 1838, Crabro)
yezo Tsuneki, 1960
Crabro cetratus Shuckard, 1837
vanderlindeni (Dahlbom, 1838, Crabro)
dilatatus (Herrich-Schäffer, 1841, Crabro)
inornatus (Matsumura, 1912, Crabro)
dentsukanus Tsuneki, 1976
Sphex leucostoma Linnaeus, 1758
carbonarius (Dahlbom, 1838, Crabro)
rugosus (Herrich-Schäffer, 1841, Crabro)
melanarius (Wesmael, 1852, Crabro)
cinctipes (Provancher, 1882, Blepharipus)
niger (Provancher, 1888, Crabro)
nigror (Fox, 1895, Crabro)
servus (Dalla Torre, 1897, Crabro)
cinctitarsis (Ashmead, 1901, Stenocrabro)
columbiae (Bradley, 1906, Blepharipus)
stygius (Mickel, 1916, Thyreopus)
utensis (Mickel, 1916, Thyreopus)
Crabro megacephalus Rossi, 1790
leucostoma misident.
bidens (Haliday, 1833, Crabro)
niger Lepeletier & Brullé, 1835
rufipes Lepeletier & Brullé, 1835
laeviceps (Smith, 1856, Crabro)
bison (Costa, 1884, Crabro)
?zaidamensis (Radoszkowski, 1887, Crabro)
leucostomoides (Richards, 1935, Coelocrabro)
Blepharipus nigritus Lepeletier & Brullé, 1835
pubescens (Shuckard, 1837, Crabro)
diversipes Herrich-Schäffer, 1841
inermis (Thomson, 1870, Crabro)
melanogaster Kohl, 1880
nigricornis (Provancher, 1888, Blepharipus)
sambucicola (Verhoeff, 1891, Crabro)
verhoeffi Tsuneki, 1967
sudai Tsuneki, 1976
babai Tsuneki, 1979
Crabro styrius Kohl, 1892
pauxillus (Gussakovskij, 1932, Crabro)
sugiharai (Iwata, 1938, Crabro)
pilicornis Tsuneki, 1977
Crabro walkeri Shuckard, 1837
aphidium misident.
geniculatus (Shuckard, 1837, Crabro)
clypearis (Schenck, 1857, Crabro)
cloevorax (Nielson, 1901, Crabro)
STENOCRABRO Ashmead, 1899
ISCHNOLYNTHUS Holmberg, 1903
Crabro distinguendus Morawitz, 1866
mucronatus (Thomson, 1870, Crabro)
added by
Crabro elongatulus Vander Linden, Crabro
annulatus Lepeletier & Brullé, 1835
varipes Lepeletier & Brullé, 1835
affinis Lepeletier & Brullé, 1835
luteipalpis Lepeletier & Brullé, 1835
morio Lepeletier & Brullé, 1835
pallidipalpis Lepeletier & Brullé, 1835
proximus (Shuckard, 1837, Crabro)
hyalinus (Shuckard, 1837, Crabro)
transversalis (Shuckard, 1837, Crabro)
obliquus (Shuckard, 1837, Crabro)
propinquus (Shuckard, 1837, Crabro)
brevis (Eversmann, 1849, Crabro)
scutellaris (Smith, 1851, Crabro)
sulcus (Fox, 1895, Crabro)
plesius (Rohwer, 1912, Stenocrabro)
berlandi (Richards, 1928, Crabro)
The British population is considered to belong to the subspecies annulatus Lepeletier & Brullé (synonyms: proximus, hyalinus, transversalis, obliquus, propinquus, berlandi).
Crabro exiguus Vander Linden, Crabro
aphidum Lepeletier & Brullé, 1835
punctum (Zetterstedt, 1838, Crabro)
anxius (Wesmael, 1852, Crabro)
shuckardi (Smith, 1856, Crabro)
ovatus (Schulz, 1906, Crabro)
Sphex palmipes Linnaeus, 1767
palmarius (Schreber, 1784, Sphex)
scutatus (Fabricius, 1787, Crabro)
ornatus Lepeletier & Brullé, 1835
scutellaris (Gimmerthal, 1836, Crabro)
gracilis (Eversmann, 1849, Crabro)
decoratus (Smith, 1856, Crabro)
Crabro tarsatus Shuckard, 1837
palmipes misident.
palmatus De Stefani Perez, 1884 preocc,
varius misspelling
pusillus Lepeletier & Brullé, 1835
striatulus Lepeletier & Brullé, 1835
spinipectus (Shuckard, 1837, Crabro)
striatus Lepeletier, 1845
intricatus (Smith, 1856, Crabro)
lepeletieri (Smith, 1856, Crabro)
Crabro wesmaeli Vander Linden, Crabro
maurus (Lepeletier & Brullé, 1835, Ceratocolus)
ziegleri (Lepeletier & Brullé, 1835, Ceratocolus)
signatus (Panzer, 1798, Crabro) preocc.
monstrosus (Dahlbom, 1845, Crabro)
confusus (Schulz, 1906, Crabro)
Crabro dimidiatus Fabricius, 1781
subpunctatus (Rossi, 1790, Crabro)
sexmaculatus (Olivier, 1792, Crabro)
signatus (Olivier, 1792, Crabro)
serripes (Panzer, 1797, Crabro)
notatus (Illiger, 1807, Crabro)
pauperatus (Lepeletier & Brullé, 1835, Blepharipus)
armipes (von Sebold, 1844, Crabro)
Crabro quadrimaculatus Fabricius, 1793
quadripunctatus (Fabricius, 1793, Crabro)
murorum (Latreille, 1805, Crabro)
levipes Vander Linden, 1829, Crabro)
bimaculatus Lepeletier & Brullé, 1835
quniquemaculatus (Dahlbom, 1838, Crabro)
rotundarius (Dahlbom, 1838, Crabro)
Crabro cavifrons Thomson, 1870
cephalotes misident.
Crabro lapidarius Panzer, 1804
?cinctus (Spinola, 1806, Crabro)
chrysostomus (Lepeletier & Brullé, 1835, Crabro)
comptus (Lepeletier & Brullé, 1835, Crabro)
xylurgus (Shuckard, 1837, Crabro)
interstinctus (Smith, 1856, Crabro)
obscurus (Smith, 1856, Crabro)
gracilissimus (Packard, 1866, Crabro)
denticulatus (Packard, 1866, Crabro)
effosus (Packard, 1866, Crabro)
papagorum (Viereck, 1908, Crabro)
Crabro ruficornis Zetterstedt, 1838
aurilabris (Herrich-Schäffer, 1841, Crabro)
nigrifrons (Cresson, 1865, Crabro)
contiguus (Cresson, 1865, Crabro)
septentrionalis (Packard, 1866, Crabro)
planifrons (Thomson, 1870, Crabro)
longipalpis (Verhöff, 1892, Crabro)
lineatotarsis (Matsumura, 1911, Crabro)
chipsani (Matsumura, 1912, Crabro)
Crabro sexcinctus Fabricius, 1775
planifrons misident.
quadricinctus (Fabricius, 1787, Crabro)
interruptefasciatus (Retzius, 1783, Sphex)
tibialis (Olivier, 1792, Crabro)
maculata (Preyssler, 1793, Crabro)
zonatus (Panzer, 1797, Crabro)
vespiformis (Panzer, 1798, Crabro)
octomaculatus (Schrank, 1802, Crabro)
cinctus (Spinola, 1806, Crabro)
flavipes (Lepeletier & Brullé, 1835, Crabro)
tetraedrus (Blanchard, 1940, Crabro)
saundersi (Perkins, 1899, Crabro)
yosemite Pate, 1946
Crabro borealis Zetterstedt, 1838
bipunctatus (Zetterstedt, 1838, Crabro)
nigrinus (Herrich-Schäffer, 1841, Crabro)
parvulus (Packard, 1866, Crabro)
gredleri (Kohl, 1878, Lindenius)
proletarius (Mickel, 1916, Crabro)
Solenius dives Lepeletier & Brullé, 1835
octonotatus (Lepeletier & Brullé, 1835, Solenius)
octavonotatus (Lepeletier & Brullé, 1835, Solenius)
alatulus (Dahlbom, 1838, Crabro)
pictipes (Herrich-Schäffer, 1841, Crabro)
auratus (Smith, 1856, Crabro)
montanus (Cresson, 1865, Crabro) preocc.
cristatus (Packard, 1866, Crabro)
cubiceps (Packard, 1866, Crabro)
heraclei (Rohwer, 1908, Crabro)
montivagans (Strand, 1916, Crabro)
XESTOCRABRO Ashmead, 1899
Crabro continuus Fabricius, 1804
vagus misident.
sexmaculatus (Say, 1824, Crabro) preocc.
fuscitarsis (Herrich-Schäffer, 1841, Crabro)
vagatus (Smith, 1869, Crabro)
pumilus (Costa, 1871, Crabro)
granulatus (Walker, 1871, Crabro)
rugopunctatus (Taschenberg, 1875, Crabro)
validus (De Stefani Perez, 1884, Crabro)
bizexmaculatus (Viereck, 1910, Crabro)
sayi (Cockerell, 1910, Crabro)
giffardi (Rohwer, 1917, Solenius)
Solenius rubicola Dufour & Perris, 1840
microstictus (Herrich-Schäffer, 1841, Crabro)
larvatus (Wesmael, 1852, Crabro)
pumilus (Costa, 1871, Crabro)
Crabro cephalotes Olivier, 1792
quadricinctus misident.
floralis (Olivier, 1792, Crabro)
geniculatus (Olivier, 1792, Crabro)
tibialis (Olivier, 1792, Crabro)
?cephalotes (Panzer, 1799, Crabro)
striatus (Lepeletier & Brullé, 1835, Crabro)
ornatus (Lepeletier & Brullé, 1835, Crabro)
striatulus (Lepeletier & Brullé, 1835, Blepharipus)
lindenius (Shuckard, 1837, Crabro)
shuckardi (Dahlbom, 1838, Crabro)
interruptus (Dahlbom, 1845, Crabro) preocc.
fargeii (Smith, 1856, Crabro)
aciculatus (Provancher, 1882, Crabro)
ruthenicus (Morawitz, 1892, Crabro)
Crabro lituratus Panzer, 1805
petiolatus (Lepeletier & Brullé, 1835, Solenius)
fasciatus (Lepeletier & Brullé, 1835, Ceratocolus)
reticulatus (Lepeletier & Brullé, 1835, Ceratoculus [lapsus])
kollari (Dahlbom, 1845, Crabro)
argenteus (Schenck, 1857, Crabro)
vestitus (Smith, 1858, Crabro)
intermedius (Morawitz, 1866, Crabro)
luxuriosus (Costa, 1871, Crabro)
KOXINGA Pate, 1944
MASHONA Pate, 1944
TONCAHUA Pate, 1944
FLORKINUS Leclercq, 1956
BIHARGNATHUS Leclercq, 1977
Crabro brevis Vander Linden, 1829
apicalis (Lepeletier & Brullé, 1835, Lindenius)
nasutus (Gribodo, 1884, Lindenius)
THYREUS Lepeletier & Brullé, 1835 preocc.
Apis clypeata Schreber, 1759
ovata (Christ, 1791, Sphex)
vexillata (Panzer, 1797, Crabro)
lapidaria (Fabricius, 1804, Crabro) preocc.
clypeata (Thunberg, 1815, Philanthus)
nigridens (Herrich-Schäffer, 1841, Crabro)
quadrifer (Dufour, 1841, Crabro)
CHALCOLAMPRUS Wesmael, 1852
TRACHELOSIMUS Morawitz, 1866
Crabro albilabris Fabricius, 1793
aenescens (Dahlbom, 1838, Crabro)
Crabro panzeri Vander Linden, 1829
venustus Lepeletier & Brullé, 1835
latebrosus (Kohl, 1905, Crabro)
harbinensis Tsuneki, 1967
mongolicus Tsuneki, 1972
Crabro pygmaeus Rossi, 1794
curtus Lepeletier & Brullé, 1835
kratochvili (Šnoflak, 1948, Crabro)
Represented by the subspecies armatus (Vander Linden, 1829, Crabro).
EUPLILIS Risso, 1826 nom. ob.
PHYSOSCELUS Lepeletier & Brullé, 1835
Sphex clavipes Linnaeus, 1758
rufiventris (Panzer, 1799, Crabro)
DRYPHUS Herrich-Schäffer, 1840
ALLIOGNATHUS Ashmead, 1899
Sphex coarctata Scopoli, 1763
crassipes (Fabricius, 1798, Crabro)
tibiale (Fabricius, 1798, Crabro) preocc.
modestum Rohwer, 1908
nigrinum Kiesenwetter, 1849 preocc.
kiesenwetteri (Morawitz, 1866, Crabro)
simplicipes (Morawitz, 1888, Corynopus)
SCHISTOSPHEX Arnold, 1922
ATELOSPHEX Arnold, 1923
Astata nitida Spinola, 1805
unicolor misident.
ibericus (de Saussure, 1867, Tachytes)
Added by
Tachytes obscuripennis Schenck, 1857
lativalvis (Thomson, 1870, Tachytes)
Only recorded as a vagrant in Deal, Kent; resident on the Channel Isles, which lie outside of the geographical coverage of this checklist.
Larra pompiliformis Panzer, 1805
pectinipes misident.
nigripennis (Spinola, 1808, Tachytes)
dimidiata (Panzer, 1809, Larra)
jokischiana (Panzer, 1809, Larra)
parvula (Cresson, 1865, Larrada)
quebecensis (Provancher, 1882, Larra)
decorus Fox, 1894
tenuipunctus Fox, 1894
consimilis Fox, 1894
rufoniger Bingham, 1897
projectus Nurse, 1903
argyrotrichus Rohwer, 1911
granulosus Mickel, 1916
erythraeus Mickel, 1916
angularis Mickel, 1916
Larra unicolor Panzer, 1809
nitidus misident.
jurinii (Drapiez, 1819, Larra)
NITELOPTERUS Ashmead, 1897
HYPOMISCOPHUS Cockerell, 1898
MISCOPHINUS Ashmead, 1898
maritimus Smith, 1858
dubius (Panzer, 1809, Larra)
metallicus Verhöff, 1890
tsunekii de Andrade, 1960
added by
bicolor misident.
moravicus Balthazar, 1957
TENILA Brèthes, 1913
RHINONITELA Williams, 1928
added by
spinolae misident.
Added by
NOTOGLOSSA Dahlbom, 1845
ALEPIDASPIS Costa, 1882
ANOXYBELUS Kohl, 1924
GONIOXYBELUS Pate, 1937
ORTHOXYBELUS Pate, 1937
LATROXYBELUS Noskiewicz & Chudoba, 1950
mucronatus misident.
?decimmaculata (Donovan, 1806, Vespa)
ferox Shuckard, 1837
nigricornis Shuckard, 1837
sericatus Gerstäcker, 1867
Vespa uniglumis Linnaeus, 1758
punctata (Fabricius, 1793, Nomada)
tridens (Fabricius, 1798, Crabro)
?decimmaculata (Donovan, 1806, Vespa)
pygmaeus Olivier, 1812
quadrinotatus Say, 1824
impatiens Smith, 1856
interruptus Cresson, 1865
fallax Gerstäcker, 1867
brodiei Provancher, 1883
hispanicus Giner Marí, 1943
APIUS Panzer, 1806
APIUS Jurine, 1807
TRYPARGILUM Richards, 1934
ASACONOTON Arnold, 1959
tibiale Zetterstedt, 1840
batumicum Antropov, 1985
Sphex figulus Linnaeus, 1758
fuliginosa (Scopoli, 1763, Sphex)
apicale Fox, 1891
yezo Tsuneki, 1956
barbarum de Beaumont, 1957
fieuzeti Giner Marí, 1959
added by
koma Tsuneki, 1956
added by
DINETUS Jurine, 1807 preocc.
Crabro pictus Fabricius, 1793
guttatus (Fabricius, 1793, Sphex) preocc.
Vespa arvensis Linnaeus, 1758
vagus (Linnaeus, 1758, Sphex)
superbus (Harris, 1776, Vespa)
tricinctus (Schrank, 1781, Vespa)
clavatus (Retzius, 1783, Sphex)
?infundibuliformis (Geoffroy, 1785, Vespa)
?petiolatus (Geoffroy, 1785, Vespa)
bipunctatus (Fabricius, 1787, Crabro)
?gibbus (Villers, 1789, Sphex) preocc.
melanostictus (Gmelin, 1790, Vespa)
?arthriticus (Rossi, 1790, Crabro)
?rachiticus (Rossi, 1790, Crabro)
annularis (Christ, 1791, Sphex)
succinctus (Olivier, 1792, Vespa)
diversus (Olivier, 1792, Vespa)
labiatus (Olivier, 1792, Crabro)
quinquemaculatus (Fabricius, 1793, Philanthus)
u-flavum (Panzer, 1794, Crabro)
capistratus (Schrank, 1796, Crabro)
pratensis Jurine, 1807
compactus Handlirsch, 1888
Sphex crabronea Thunberg, 1791
sabulosus (Fabricius, 1787, Crabro) preocc.
ruficornis (Villers, 1789, Sphex) preocc.
sabulosus (Olivier, 1792, Crabro) preocc.
ruficornis Fabricius, 1793 preocc.
frontalis (Panzer, 1797, Crabro)
petiolatus (Panzer, 1797, Crabro)
fulvicornis Fabricius, 1804
XYLOCELIA Rohwer, 1915
NEODIODONTUS Tsuneki, 1972
CORENIUS Tsuneki, 1974
friesei misident.
Crabro minutus Fabricius, 1793
franclemonti Krombein, 1939
Pemphredon tristis Vander Linden, 1829
XYLOECUS Shuckard, 1837
COELOECUS Verhöff, 1890
HEROECUS Verhöff, 1890
angustus Gussakovskij, 1952
added by
Diodontus gracilis Curtis, 1834
insignis misident.
turionum misident.
brevicornis Morawitz, 1864
Pemphredon insignis Vander Linden, 1829
roettgeni Verhöff, 1890
shuckardi Yasumatsu, 1934
gracilis misident.
tenuis Morawitz, 1864
gertrudis Krombein, 1938
added by
CEMONUS Panzer, 1806
DINEURUS Westwood, 1837
CERATOPHORUS Shuckard, 1837
DIPHLEBUS Westwood, 1840
CHEVRIERIA Kohl, 1883
SUSANOWO Tsuneki, 1972
Taxonomy follows
Diphlebus austriacus Kohl, 1888
enslini misident.
coracina Valkeila, 1972
tener Valkeila, 1972
nescia Merisuo, 1972
England
shuckardi (Morawitz, 1864, Cemonus)
dentata (Puton, 1871, Cemonus)
tenax Fox, 1829
Cemonus lethifer Shuckard, 1837
lethifera misspelling
austriaca misident.
strigatus (Chevrier, 1870, Cemonus)
fabricii (Müller, 1911, Cemonus)
fuscatus (Wagner, 1918, Diphlebus)
littoralis (Wagner, 1918, Diphlebus)
neglectus (Wagner, 1918, Diphlebus)
minutus (Wagner, 1918, Diphlebus)
enslini Wagner, 1931
brevipetiolata Wagner, 1932
platyura Gussakovskij, 1952
minor Gussakovskij, 1952
levinota Merisuo, 1972
nannophyes Merisuo, 1972
dispar Valkeila, 1972
gemina Valkeila, 1972
trichogastor Valkeila, 1972
?sudaorum Tsuneki, 1977
Crabro lugubris Fabricius, 1793
concolor Say, 1824
ocellaris Gimmerthal, 1836
luctuosa Shuckard, 1837
morio Cresson, 1865 preocc.
cressoni Dalla Torre, 1897
provancheri Dalla Torre, 1897
tinctipennis Cameron, 1908
shawii Rohwer, 1917
pacifica Gussakovkij, 1932
anthracinus (Smith, 1851, Ceratophorus)
carinata Thomson, 1870
clypealis Thomson, 1870
intermedia Tsuneki, 1951
Cemonus rugifer Dahlbom, 1844
rugifera misspelling
unicolor (Panzer, 1798, Sphex) preocc.
pilosa (Gimmerthal, 1836, Cemonus)
wesmaeli (Morawitz, 1864, Cemonus)
scotica Perkins, 1929
solivaga (Bondroit, 1931, Cemonus)
bucharicus Gussakovskij, 1952
mortifer Valkeila, 1972
punctifer Valkeila, 1972
scytica Valkeila, 1972
CELIA Shuckard, 1837 preocc.
MICROGLOSSA Rayment, 1930 preocc.
MICROGLOSSELLA Rayment, 1935
TAIALIA Tsuneki, 1971
exspectata Valkeila, 1957
curruca misident.
England
Has been called curruca (Dahlbom, 1844, Celia) in several publications, following the synonymy by
Celia curruca Dahlbom, 1844
pulawskii Dolfuss, 1983
nikkoensis Tsuneki, 1971
Stigmus troglodytes Vander Linden, 1829
minutissimus (Radoszkowski, 1877, Stigmus)
vagans Blüthgen, 1953
ANTRONIUS Zetterstedt, 1838
GONOSTIGMUS Rohwer, 1911
ATOPOSTIGMUS Krombein, 1973
ater Jurine, 1807
added by
europaeus Tsuneki, 1954
verhoeffi Tsuneki, 1954
APORIA Wesmael, 1852 preocc.
APORINA Gussakovskij, 1937 preocc.
Psen bicolor Jurine, 1807
equestris misident.
?rufa (Panzer, 1805, Psen)
rossicus (Gussakovskij, 1937, Psen)
Trypoxylon equestre Fabricius, 1804
bicolor misident.
Sphex lutarius Fabricius, 1787
shuckardi Wesmael, 1852
basirufa Packard, 1867
nebrascensis Smith, 1908
dispar (Gussakovskij, 1937, Psen)
mallochi Finnamore, 1980
Mimesa atratina Morawitz, 1891
carbonaria (Tournier, 1899, Mimesa)
belgica (Bondroit, 1932, Mimesa)
longula (Gussakovskij, 1932, Mimesa)
sameshimai (Yasumatsu, 1937, Psen)
Mimesa dahlbomi Wesmael, 1852
Mimesa littoralis Bondroit, 1934
unicolor misident.
fulvitarsis (Gussakovskij, 1934, Psen)
fulvitarsis (Gussakovskij, 1937, Psen) preocc.
celtica (Spooner, 1948, Mimesa)
Mimesa spooneri Richards, 1948
Psen unicolor Vander Linden, 1829
borealis (Dahlbom, 1842, Mimesa)
palliditarsis (Saunders, 1904, Mimesa)
oresterus (van Lith, 1976, Psen)
added by
PSENUS Rafinesque, 1815
PSENIA Stephens, 1829
DAHLBOMIA Wissmann, 1849
MESOPORA Wesmael, 1852
CAENOPSEN Cameron, 1899
Crabro ater Olivier, 1792
ater (Fabricius, 1794, Sphex) preocc.
compressicornis (Fabricius, 1804, Pelopoeus)
atratus (Jurine, 1807, Psen)
serraticornis (Jurine, 1807, Psen)
DIODONTUS misident.
NEOFOXIA Viereck, 1901
STENOMELLINUS Schulz, 1911
EOPSENULUS Gussakovskij, 1934
NIPPONOPSEN Yasumatsu, 1938
Diodontus is a misidentification by American authors in the older literature
Psen concolor Dahlbom, 1843
intermedius (Schenck, 1857, Psen)
ambiguus (Schenck, 1857, Psen)
Sphex pallipes Panzer, 1798
atratus (Fabricius, 1804, Trypoxylon)
montanus (Costa, 1861, Psen)
haemorrhoidalis (Costa, 1871, Psen)
minutus (Tournier, 1899, Psen)
chevrieri (Tournier, 1899, Psen)
nigricornis (Tournier, 1899, Psen)
pygmaeus (Tournier, 1899, Psen)
rubicola Harttig, 1931
brevitarsis Merisuo, 1937
Psen schencki Tournier, 1889
simplex (Tournier, 1899, Psen)
longulus (Tournier, 1899, Psen)
NECTANEBUS Spinola, 1839
DIAMMA Dahlbom, 1844
DIDESMUS Dahlbom, 1845
APIRAPTRIX Shestakov, 1923
PARACERCERIS Brèthes, 1913
BUCERCERIS Minkiewicz, 1934
STERCOBATA Gussakovskij, 1935
APICERCERIS Pate, 1937
Sphex arenarius Linnaeus, 1758
xanthocephala (Forster, 1771, Sphex)
exulta (Harris, 1776, Vespa)
petulans (Harris, 1776, Vespa)
serripes (Fabricius, 1781, Vespa)
arenosa (Gmelin, 1790, Vespa)
aurita (Fabricius, 1794, Philanthus)
striolata Schletterer, 1887
Philanthus quadricinctus Panzer, 1799
fasciata Spinola, 1806
Crabro quinquefasciatus Rossi, 1792
interrupta misident.
nasuta Dahlbom, 1844 preocc.
subdepressa Lepeletier, 1845
Philanthus ruficornis Fabricius, 1793
labiata misident.
bidens (Schrank, 1802, Crabro)
cunicularia (Schrank, 1802, Crabro)
quadricincta (Fabricius, 1804, Mellinus)
trifidus (Fabricius, 1804, Philanthus)
?nasuta Latreille, 1809
laminifera Costa, 1867
Sphex rybyensis Linnaeus, 1771
ornata (Fabricius, 1790, Philanthus)
apifalco (Christ, 1791, Sphex)
semicincta (Panzer, 1797, Philanthus)
hortorum (Panzer, 1799, Philanthus)
variabilis (Schrank, 1802, Crabro)
biguttata (Thunberg, 1815, Philanthus)
colon (Thunberg, 1815, Philanthus)
kashmirensis Nurse, 1903
dittrichi Schulz, 1904
reginae Eck, 1979
Philanthus sabulosus Panzer, 1799
emarginata (Panzer, 1799, Philanthus)
tricincta (Spinola, 1805, Philanthus)
?pygmaea (Thunberg, 1815, Philanthus)
minuta Lepeletier, 1845
superba Shestakov, 1923
SYMBLEPHILUS Panzer, 1806
SIMBLEPHILUS Jurine, 1807
CHEILOPOGONUS Westwood, 1835
ANTHOPHILUS Dahlbom, 1844
EPHIPHILANTHUS Ashmead, 1899
PSEUDOPHILANTHUS Ashmead, 1899
OCLOCLETES Banks, 1913
Vespa triangulum Fabricius, 1775
ruspatrix (Linnaeus, 1767, Vespa) nom. ob.
fasciatus (Geoffroy, 1785, Vespa)
maculatus (Christ, 1791, Sphex)
limbatus (Olivier, 1792, Vespa)
androgynus (Rossi, 1792, Crabro)
pictus Panzer, 1797
discolor Panzer, 1799
apivorus Latreille, 1799
allionii Dahlbom, 1845
SPHEX misident.
MISCUS Jurine, 1807
COLOPTERA Latreille, 1845
ARGYRAMMOPHILA Gussakovskij, 1928
APYCNEMIA Leclercq, 1961
campestris misident.
arvensis (Dahlbom, 1843, Miscus)
susterai Šnoflak, 1943
adriaansei Wilcke, 1945
Sphex sabulosus Linnaeus, 1758
hortensis (Poda, 1761, Sphex)
frischii (Geoffroy, 1785, Ichneumon)
dimidiata (Christ, 1791, Sphex) preocc.
vulgaris Kirby, 1798
pulvillata Sowerby, 1805
mucronata (Jurine, 1807, Sphex)
cyanescens Dahlbom, 1845
?vischu Cameron, 1889
kamtschatica Gussakovskij, 1932
PSAMMOPHILA Dahlbom, 1842 preocc.
PODALONIA Spinola, 1853 suppressed
Ammophila affinis Kirby, 1798
lutaria misident.
ariasi (Mercet, 1906, Ammophila)
Sphex hirsutus Scopoli, 1763
viatica misident.
arenaria (Fabricius, 1787, Sphex) preocc.
arenosa (Gmelin, 1790, Sphex)
argentea (Kirby, 1798, Ammophila)
Taxonomy mostly follows
ANTHRENA Illiger, 1801
ANTHOCHARESSA Gistel, 1850
Melitta clarkella Kirby, 1802
Apis fulva Müller, 1766
armata (Gmelin, 1790, Apis)
Apis helvola Linnaeus, 1758
subdentata (Kirby, 1802, Melitta)
Apis praecox Scopoli, 1763
smithella (Kirby, 1802, Melitta)
clypeata Smith, 1855 preocc.
ambigua Perkins, 1895, preocc.
Melitta varians Kirby, 1802
varians (Rossius, 1792, Apis): misident.
angulosa (Kirby, 1802, Melitta)
Nomada hattorfiana Fabricius, 1775
lathamana (Kirby, 1802, Melitta)
fulvescens Smith, 1847
Apis fulvago Christ, 1791
constricta Smith, 1849
Melitta denticulata Kirby, 1802
listerella (Kirby, 1802, Melitta)
Melitta fuscipes Kirby, 1802
Melitta nigriceps Kirby, 1802
lanifrons (Kirby, 1802, Melitta)
Melitta tridentata Kirby, 1802
rufitarsis (Kirby, 1802, Melitta)
XANTHANDRENA Lanham, 1949
GEANDRENA LaBerge, 1964
picicornis (Kirby, 1802, Melitta)
pilosula (Kirby, 1802, Melitta)
gwynana (Kirby, 1802, Melitta)
proxima Smith, 1847 preocc.
aestiva Smith, 1849
consimilis Smith, 1849 preocc.
Melitta labialis Kirby, 1802
separata Smith, 1847
eximia Smith, 1847
longipes Smith, 1847
distincta Smith, 1847
eximia misident.
zonalis (Kirby, 1802, Melitta)
stragulata Illiger, 1806
strangulata misspelling
stragulata is the summer brood of rosae
carantonica misident.
jacobi Perkins, 1921
johnsoni Perkins, 1921
England, Scotland, Wales, Ireland, Isle of Man
Although usually referred to as scotica in Britain (e.g.
Melitta trimmerana Kirby, 1802
spinigera (Kirby, 1802, Melitta)
carantonica Peréz, 1916
Melitta barbilabris Kirby, 1802
sericea (Christ, 1791, Apis) preocc.
albicrus (Kirby, 1802, Melitta)
cetii (Schrank, 1781, Apis)
schrankella (Kirby, 1802, Melitta)
frontalis Smith, 1849
GYMNANDRENA Hedicke, 1933
CRYPTANDRENA Lanham, 1949 preocc.
BYTHANDRENA Lanham, 1950
Apis cineraria Linnaeus, 1758
Melitta nigroaenea Kirby, 1802
aprilina Smith, 1848
The population on the Isles of Scilly and Channel Islands has been described as the subspecies sarnia Richards, 1979.
Apis nitida Müller, 1776
pubescens Olivier, 1789
consimilis Smith, 1847
Apis thoracica Fabricius, 1775
melanocephala (Kirby, 1802, Melitta)
ANDRENELLA Hedicke, 1933
moricella Perkins, 1914
Melitta minutula Kirby, 1802
parvula (Kirby, 1802, Melitta)
nigrifrons Smith, 1855 preocc.
parvuloides Perkins, 1914
Melitta nana Kirby, 1802
spreta misident.
nana misident.
saundersella Perkins, 1914
Melitta chrysosceles Kirby, 1802
connectens (Kirby, 1802, Melitta)
lucens Imhoff, 1868
Melitta coitana Kirby, 1802
shawella (Kirby, 1802, Melitta)
SCHIZANDRENA Hedicke, 1933
GLYPHANDRENA Hedicke, 1933
MITSUKURIELLA Hirashima & LaBerge, 1965
MITSUKURIAPIS Hirashima, LaBerge & Ikudomem 1884
Melitta bimaculata Kirby, 1802
articulata Smith, 1847
conjuncta Smith, 1847
decorata Smith, 1847
vitrea Smith, 1847
Although treated as a synonym of pilipes by
carbonaria misident.
spectabilis Smith, 1853
praetexta Smith, 1872
Melitta tibialis Kirby, 1802
mouffetella (Kirby, 1802, Melitta)
atriceps (Kirby, 1802, Melitta)
cingulata misident.
rubricata Smith, 1847
Melitta proxima Kirby, 1802
digitalis (Kirby, 1802, Melitta)
EREMANDRENA LaBerge, 1964
Melitta angustior Kirby, 1802
lacinia Smith, 1847
PLATANDRENA Viereck, 1924
STENANDRENA Timberlake, 1949
confinis Stöckhert, 1930
Melitta dorsata Kirby, 1802
collinsonana (Kirby, 1802, Melitta)
lewinella (Kirby, 1802, Melitta)
nudiuscula (Kirby, 1802, Melitta)
subincana (Kirby, 1802, Melitta)
Melitta ovatula Kirby, 1802
afzeliella (Kirby, 1802, Melitta)
barbata (Kirby, 1802, Melitta)
fuscata (Kirby, 1802, Melitta)
picipes (Kirby, 1802, Melitta)
albofasciata Thomson, 1870
?ocreata Christ, 1791 nom. dub.
Melitta wilkella Kirby, 1802
barbatula (Kirby, 1802, Melitta)
convexiuscula (Kirby, 1802, Melitta)
xanthura (Kirby, 1802, Melitta)
analis misident.
Apis haemorrhoa Fabricius, 1781
albicans misident.
contigua (Kirby, 1802, Melitta)
fulvicrus (Kirby, 1802, Melitta)
extricata Smith, 1849
fasciata Nylander, 1852 preocc.
picicrus Schenck, 1853
Apis banksiana Kirby, 1802
ursinus misident.
Apis calcarata Scopoli, 1763
linnaeella (Kirby, 1802, Apis)
Apis plumipes Pallas, 1772
acervorum misident.
pilipes (Fabricius, 1775, Apis)
Apis furcata Panzer, 1798
Apis quadrimaculata Panzer, 1798
subglobosa (Kirby, 1802, Apis)
SAROPODA Latreille, 1809
Apis bimaculata Panzer, 1798
Apis retusa Linnaeus, 1758
haworthana (Kirby, 1802, Apis)
pennipes (Kirby, 1802, Apis)
mellifica Linnaeus, 1761
There have been various subspecies names proposed for honey bees, with the supposedly native north west European population being A. mellifera mellifera. See
Various subspecific names have been used for British Bombus but these are generally poorly justified and we have treated these as synonymous here. Subgeneric classification follows
TERRESTRIBOMBUS Vogt, 1911
Full synonymy for this subgenus is given by
Apis cryptarum Fabricius, 1775
added by
Apis lucorum Linnaeus, 1761
Apis terrestris Linnaeus, 1758
audax (Harris, 1776, Apis)
Apis cullumana Kirby, 1802
Apis soroeensis Fabricius, 1777
HORTOBOMBUS Vogt, 1911
Apis hortorum Linnaeus, 1761
?flavonigrescens Smith, 1846
ivernicus Sladen, 1912
splendida Stelfox, 1938
Populations in Ireland have been referred to the subspecies B. hortorum ivernicus.
Apis ruderata Fabricius, 1775
perniger (Harris, 1776, Apis)
harrisellus (Kirby, 1802, Apis)
tunstallanus (Kirby, 1802, Apis)
English populations have been referred to the subspecies B. ruderatus perniger.
LAPIDARIOBOMBUS Vogt, 1911
Apis lapidaria Linnaeus, 1758
England, Scotland, Wales, Ireland, Isle of Man
ALLOPSITHYRUS Popov, 1931
ASHTONIPSITHYRUS Frison, 1927
FERNALDAEPSITHYRUS Frison, 1927
METAPSITHYRUS Popov, 1931
Apis barbutella Kirby, 1802
Apis bohemicus Seidl, 1837
distinctus (Pérez, 1884, Psithyrus)
Apis campestris Panzer, 1801
rossiellus (Kirby, 1802, Apis)
leeanus (Kirby, 1802, Apis)
francisanus (Kirby, 1802, Apis)
swynnertoni (Richards, 1936, Psithyrus)
Populations in Western Scotland have been referred to the subspecies B. campestris swynnertoni.
Apis rupestris Fabricius, 1793
albinellus (Kirby, 1802, Apis)
Psithyrus sylvestris Lepeletier, 1832
quadricolor misident.
Apis vestalis Geoffroy, 1785
PRATOBOMBUS Vogt, 1911
Apis hypnorum Linnaeus, 1758
England, Scotland, Wales
added by
Apis jonella Kirby, 1802
nivalis misident.
scrimshiranus (Kirby, 1802, Apis)
atrocorbiculosus Vogt, 1911
hebridensis Wild, 1931
vogtii Richards, 1933 preocc.
monapiae Kruseman, 1953
vogtianus Rasmont, 1983
The following populations have been given subspecies names: B. jonellus hebridensis on the Hebrides; B. jonellus monapiae on the Isle of Man; B. jonellus vogtianus on the Shetlands (vogtianus is a replacement name for the preccupied vogtii).
lapponicus misident.
scoticus Pittioni, 1942
Apis pratorum Linnaeus, 1761
subinterruptus (Kirby, 1802, Apis)
donovanella (Kirby, 1802, Apis)
burrellana (Kirby, 1802, Apis)
Apis subterranea Linnaeus, 1758
collinus Smith, 1844
latreillellus (Kirby, 1802, Apis)
Extinct in Britain since the 1980s but an attempt is being made to reintroduce the species to southern England. British populations have been referred to the subspecies B. subterraneus latreillellus whereas the introduced individuals from Sweden have been ascribed to the nominate subspecies.
POMOBOMBUS Krüger, 1917
solstitialis Panzer, 1806
helferanus Seidl, 1837
venustus Smith, 1876 preocc.
anglicus Yarrow, 1978
Apis muscorum Linnaeus, 1758
arcticus misident.
smithianus misident.
pallidus Evans, 1901 preocc.
laevis Vogt, 1909
sladeni Vogt, 1911
allenellus Stelfox, 1933
orcadensis Richards, 1935
scyllonius Richards, 1935
celticus Yarrow, 1978
agricolae Baker, 1996
The following populations have been given subspecies names: B. muscorum sladeni on the Irish mainland and southern England; B. muscorum allenellus on the Aran islands; B. muscorum orcadensis on Orkney; B. muscorum scyllonius on the Isles of Scilly; B. muscorum celticus in mainland Scotland and Northern England; B. muscorum agricolae on the Shetland islands.
Apis pascuorum Scopoli, 1763
vulgo (Harris, 1776, Apis)
agrorum (Fabricius, 1787, Apis) preocc.
floralis (Gmelin, 1790, Apis)
francillonellus (Kirby, 1802, Apis)
sowerbianus (Kirby, 1802, Apis)
beckwithella (Kirby, 1802, Apis)
curtisellus (Kirby, 1802, Apis)
forsterellus (Kirby, 1802, Apis)
cognatus Stevens, 1846
smithianus White, 1851
septentrionalis Vogt, 1909
The following populations have been given subspecies names: B. pascuorum vulgo in southern England and Wales; B. pascuorum floralis in Ireland; B. pascuorum septentrionalis in Scotland and Northern England.
Bremus pomorum Panzer, 1805
There is confusion surrounding the dates of capture of the supposedly British specimens of pomorum, collected near Deal in Kent (summarised by
Apis ruderaria Müller, 1776
derhamellus (Kirby, 1802, Apis)
raiellus (Kirby, 1802, Apis)
Apis sylvarum Linnaeus, 1761
nigrescens Pérez, 1879
distinctus Vogt, 1909
Apis longicornis Linnaeus, 1758
linguaria (Fabricius, 1775, Apis)
tuberculata misident. [?]
Apis albifrons Forster, 1771
punctata (Fabricius, 1775, Apis)
armata (Panzer, 1799, Andrena) preocc.
Apis luctuosa Scopoli, 1770
Nomada crucigera Panzer, 1799
rufipes Thomson, 1870
Apis variegata Linnaeus, 1758
notatus misident.
productus Thomson, 1870
atrata Smith, 1846
kirbyella Stephens, 1846
alboguttata misident.
Apis fabriciana Linnaeus, 1767
fabriciella (Kirby, 1802, Apis)
quadrinotata (Kirby, 1802, Apis)
Apis ferruginata Linnaeus, 1767
lateralis misident.
xanthosticta (Kirby, 1802, Apis)
bridgmaniana Smith, 1876
Apis flavoguttata Kirby, 1802
rufocincta (Kirby, 1802, Apis)
Apis flavopicta Kirby, 1802
jacobaeae misident. [?]
lineola Panzer, 1798
sexcincta (Kirby, 1802, Apis)
caprea (Kirby, 1802, Apis)
cornigera (Kirby, 1802, Apis)
subcornuta (Kirby, 1802, Apis)
Apis goodeniana Kirby, 1802
succincta misident.
alternata (Kirby, 1802, Apis)
bucephalae Perkins, 1917
ferruginata misident.
germanica misident.
pleurosticta misident.
stigma misident.
cinctiventris Friese, 1921
Apis lathburiana Kirby, 1802
rufiventris (Kirby, 1802, Apis)
Apis leucophthalma Kirby, 1802
borealis Zetterstedt, 1838
inquilina Smith, 1844
Apis marshamella Kirby, 1802
alternata misident.
mistura Smith, 1851
ruficornis misident.
tormentillae Alfken, 1901
Apis ruficornis Linnaeus, 1758
bifida Thomson, 1872
solidaginis misident.
picta (Kirby, 1802, Apis)
rufopicta (Kirby, 1802, Apis)
connexa (Kirby, 1802, Apis)
schaefferella (Kirby, 1802, Apis)
Apis sheppardana Kirby, 1802
furva misident.
dalii Curtis, 1832
hillana (Kirby, 1802, Apis)
ochrostoma (Kirby, 1802, Apis)
vidua Smith, 1844
Apis cyanea Kirby, 1802
Apis violacea Linnaeus, 1758
Apis cunicularia Linnaeus, 1761
The British population has been described as a separate subspecies, C. cunicularius celticus O’Toole, 1974, and Else, Field & O’Toole (in
montanus Morawitz, 1876
Apis fodiens Geoffroy, 1785
England
added by
picistigma Thomson, 1872
Apis succincta Linnaeus, 1758
plantarius Smith, 1842
rupestris (Smith, 1872, Prosopis)
Mellita annularis Kirby, 1802
euryscapus misident.
spilotus Forster, 1871
masoni (Saunders, 1894, Prosopis)
Mellita dilatata Kirby, 1802
annularis misident.
varipes (Smith, 1853, Prosopis)
NESOPROSOPIS Perkins, R.C.L., 1899
rubicola (Smith, 1869, Prosopis) preocc.
The subspecies perkinsi Blüthgen, 1926, is recognised from Britain.
gibbus misident.
genalis Thomson, 1872
kriechbaumeri Förster, 1871
palustris (Perkins, 1900, Prosopis)
Sphex signata Panzer, 1798
quadricinctus misident.
tetrazonius misident.
Apis rubicunda Christ, 1791
quadrifasciatus Smith, 1870
nesiotis Perkins, 1922
alpinus Alfken, 1907
confusus perkinsi Blüthgen, 1926
flavipes misident.
Apis subaurata Rossi, 1792
gramineus Smith, 1849
Apis tumulorum Linnaeus, 1758
Nomenclature for some species ('Evylaeus' s. l.) updated from
Halictus cupromicans Pérez, 1903
The Irish population has been described as the subspecies hibernicum Ebmer, 1970, with other populations referred to the subspecies scoticum Ebmer, 1970.
Melitta leucopus Kirby, 1802
aeratum (Kirby, 1802, Melitta)
semiaeneum (Brullé, 1832, Halictus)
viridaeneum (Blüthgen, 1918, Halictus)
Hylaeus morio Fabricius, 1793
Melitta smeathmanella Kirby, 1802
Halictus angusticeps Perkins, 1895
Halictus brevicornis Schenck, 1869
Melitta minutissima Kirby, 1802
arnoldi (Saunders, 1910, Halictus)
Melitta nitidiuscula Kirby, 1802
Hylaeus parvulus Schenck, 1853
minutum misident.
Halictus pauperatus Brullé, 1832
breviceps (Saunders, 1879, Halictus)
Hylaeus punctatissimus Schenck, 1853
longiceps (Saunders, 1879, Halictus)
Halictus puncticollis Morawitz, 1872
Halictus rufitarsis Zetterstedt, 1838
atricorne (Smith, 1870, Halictus)
Halictus semilucens Alfken, 1914
Halictus sexstrigatus Schenck, 1870
sabulosum (Warncke, 1986, Halictus)
added by
Melitta villosula Kirby, 1802
punctulatum (Kirby, 1802, Melitta)
Melitta laevigata Kirby, 1802
lugubris (Kirby, 1802, Melitta)
Hylaeus lativentris Schenck, 1853
decipiens (Perkins, 1913, Halictus)
Halictus prasinus Smith, 1848
Melitta quadrinotata Kirby, 1802
Melitta sexnotata Kirby, 1802
nitidum misident.
Melitta xanthopus Kirby, 1802
tricingulum Curtis, 1833
Apis leucozonia Schrank, 1781
similis (Smith, 1853, Halictus)
Halictus zonulus Smith, 1848
Apis albipes Fabricius, 1781
Apis calceata Scopoli, 1763
cylindricum (Fabricius, 1793, Hylaeus)
fulvocinctum (Kirby, 1802, Melitta)
obovatum (Kirby, 1802, Melitta)
Halictus fratellus Pérez, 1903
nigrum misident.
subfasciatum (Nylander, 1848, Halictus)
freygessneri (Alfken, 1905, Halictus)
Melitta fulvicornis Kirby, 1802
subfasciatum misident.
Melitta laevis Kirby, 1802
Halictus laticeps Schenck, 1869
semipunctulatum misident.
Melitta malachura Kirby, 1802
longulum (Smith, 1848, Halictus)
Hylaeus pauxillus Schenck, 1853
immarginatum (Schenck, 1853, Hylaeus)
variegatus von Hagens, 1874
Sphex ephippia Linnaeus, 1767
divisus (Kirby, 1802, Melitta)
similis Wesmael, 1835
Melitta geofrella Kirby, 1802
affinis von Hagens, 1882
fasciatus von Hagens, 1882
Sphex gibba Linnaeus, 1758
picea (Kirby, 1802, Melitta)
sphecoides (Kirby, 1802, Melitta)
dimidiatus von Hagens, 1882
Melitta monilicornis Kirby, 1802
subquadratus Smith, 1845
pilifrons Thomson, 1870
ruficrus misident.
rufiventris misident.
Rhophites halictulus Nylander, 1852
vulgaris Schenck, 1861
Apis manicata Linnaeus, 1758
Represented by the subspecies nigrithorax Dalla Torre, 1877
added by
Gyrodroma ornatula Klug, 1807
octomaculata Smith, 1843
Apis phaeoptera Kirby, 1802
Apis punctulatissima Kirby, 1802
aterrima (Panzer, 1798, Apis) preocc.
Apis inermis Kirby, 1802
acuminata Nylander, 1852 preocc.
England, Wales
Anthophora conoidea Illiger, 1806
vectis Curtis, 1831
sponsa Smith, 1855
Apis quadridentata Linnaeus, 1758
umbrina Smith, 1843
dorsalis Pérez, 1879 synonymy by
argentata misident.
Apis centuncularis Linnaeus, 1758
Apis ligniseca Kirby, 1802
Includes the subspecies hiberniae Perkins, 1925.
Megachile ericetorum Lepeletier, 1841
fasciata (Smith, 1844, Megachile)
rufitarsis (Smith, 1844, Megachile)
Apis circumcincta Kirby, 1802
Apis maritima Kirby, 1802
Apis willughbiella Kirby, 1802
Includes the nominate subspecies and hibernica Perkins, 1925
Apis campanularum Kirby, 1802
Apis florisomnis Linnaeus, 1758
maxillosum (Linnaeus, 1767, Apis)
Apis truncorum Linnaeus, 1758
Osmia claviventris Thomson, 1872
leucomelana misident.
Apis leucomelana Kirby, 1802
parvula (Dufour & Perris, 1840, Osmia)
Apis spinulosa Kirby, 1802
Apis caerulescens Linnaeus, 1758
aenea (Linnaeus, 1761, Apis)
Apis niveata Fabricius, 1804
fulviventris (Panzer, 1798, Apis)
Apis leaiana Kirby, 1802
fulviventris misident.
Anthophora inermis Zetterstedt, 1838
parietina misident.
added by Else in
Apis xanthomelana Kirby, 1802
atricapilla Curtis, 1828
Apis bicolor Schrank, 1781
Apis bicornis Linnaeus, 1758
rufa (Linnaeus, 1758, Apis)
hedera Smith, 1844
According to
Andrena hirtipes Fabricius, 1793
?altercator (Harris, 1776, Apis) nom. dub.
plumipes (Panzer, 1797, Andrena)
swammerdamella (Kirby, 1802, Melitta)
labiata misident.
PSEUDOCILISSA Radoszkowski, 1891
Andrena haemorrhoidalis Fabricius, 1775
chrysura Kirby, 1802
Apis leporina Panzer, 1799
melanura (Nylander, 1852, Kirbya)
Some distribution records taken from
Taxonomy mostly follows
PERISEMUS Förster, 1856
ANOXUS Thomson, 1862 synonymy by
EPISEMUS Thomson, 1862
DIGONIOZUS Kieffer, 1905
Anoxus boops Thomson, 1862
added by
Perisemus cephalotes Förster, 1860
fuscicornis var. tibialis Kieffer, 1905
Omalus fuscicornis Jurine, 1807
sygenesiae Haliday, 1834
fulvicornis Curtis, 1838
triareolatus Förster, 1851
variabilis (Thomson, 1862, Episemus)
hyalinus (Marshall, 1874, Perisemus) synonymy by
fuscicornis var. maurus Kieffer, 1905
brevipennis Hellén, 1920
berlandi Arle, 1929
PARASIEROLA Cameron, 1883
PROGONIOZUS Kieffer, 1905
PERISIEROLA Kieffer, 1914
Bethylus claripennis Förster, 1851
fuscipennis (Förster, 1851, Bethylus)
distigmus Thomson, 1862
audouinii Westwood, 1874
claripennis var. fuscipennis Kieffer, 1905 preocc.
claripennis var. tibialis Kieffer, 1905
HOLOPEDINA Förster, 1850
CEPHALOMIA Kirchner, 1867
polypori (Förster, 1850, Holopedina)
brevipennis Kieffer, 1906
formiciformis var. sulcata Kieffer, 1906
Cephalonomia brevipennis is listed as a separate species (described from English specimens, now missing) by
Sclerochroa gallicola Ashmead, 1887
nubilipennis (Ashmead, 1887, Holopedina)
xambeui Girad, 1898
quadridentata Duchaussoy, 1920
strandi Hoffer, 1936
England
Ateleopterus tarsalis Ashmead, 1893
carinata Kieffer, 1907
meridionalis Brethes, 1913
kiefferi Fouts, 1920
DOLUS Motshultsky, 1863
MUELLERELLA Saussure, 1892
HOMOGLENUS Kieffer, 1904 synonymy by
PAREPYRIS Kieffer, 1913
PSILEPYRIS Kieffer, 1913
ARTIEPYRIS Kieffer, 1913
fraternus Westwood, 1874
saeva Westwood, 1874
multidentatus Keiffer, 1906
multidentatus var. angustipennis Kieffer, 1906
England
Supposedly described from the New Forest, the type is lost and the species has not been satisfactorily interpreted since.
MISEPYRIS Kieffer, 1913
PAREPYRIS Brethes, 1913
Tiphia glabrata Fabricius, 1798
hawaiiensis (Ashmead, 1901, Holepyris)
Parepyris sylvanidis Brethes, 1913
zeae (Turner & Waterston, 1921, Rhabdepyris)
see
Bethylus femoralis Förster, 1860
microneurus (Kieffer, 1906, Allepyris)
nigricrus (Kieffer, 1906, Allepyris)
England
Bethylus pedatus Say, 1836
England
Added by
SNAPPANIA Hedqvist, 1975
Anoxus chittendenii Ashmead, 1893
Cephalonomia westwoodi Kieffer, 1914
kiefferi Gahan, 1931
ACREPYRIS Kieffer, 1905
MANGESIA Kieffer, 1911
TRICHELOBRACHIUM Kieffer, 1914
Bethylus depressus Fabricius, 1805
roubali (Menozzi, 1925, Pseudisobrachium)
MONEPYRIS Kieffer, 1905
XESTOBETHYLUS Cameron, 1909
PLUTOBETHYLUS Kieffer, 1910
LYSSEPYRIS Kieffer, 1913
XANTEPYRIS Kieffer, 1913
PARISOBRACHIUM Kieffer, 1914
AFRISOBRACHIUM Benoit, 1957
EDAPHOLIGON Oglobin, 1963
Epyris subcyaneus Haliday, 1838
halidaii (Westwood, 1874, Epyris)
carpentieri Kieffer, 1906
carpentieri var. septemfasciatum Kieffer, 1906
cantianum Chitty, 1906
concolor Kieffer, 1906
Sphex semiaurata Linnaeus, 1761
auratus (Panzer, 1798, Ichneumon)
pallipes Lepeletier, 1805
diana Mocsáry, 1889
Synonymy follows
Ichneumon nitidulus Fabricius, 1793
Some distribution data from
The number of species recognised in the Chrysis ignita group has varied from author to author.
brevidens Tournier, 1879
England, Scotland, Wales
added by
saussurei Chevrier, 1862
added by
Sphex ignita Linnaeus, 1758
chrysoprasina Hellén, 1919 preocc.
helleni Linsenmaier, 1959
succincta misident.
auripes Wesmael, 1839
schenckiana Linsenmaier, 1959 preocc.
added by
rutiliventris misident.
bidentata Linnaeus, 1767
Chrysis hirsuta Gerstäcker, 1869
osmiae (Thomson, 1870, Chrysis)
Chrysis radians Harris, 1776
pustulosa (Abeille de Perrin, 1878, Chrysis)
Chrysis neglecta Shuckard, 1836
Sphex cyanea Linnaeus, 1758
NOTOZUS Förster, 1853
Chrysis panzeri Fabricius, 1804
scutellaris (Panzer, 1798, Chrysis) preocc.
constrictus misident.
Chrysis ardens Latreille, 1801
Hedychrum coriaceum Dahlbom, 1854
Hedychrum cupreum Dahlbom, 1845
integrum (Dahlbom, 1854, Hedychrum) synonymy by
Chrysis rosea Rossi, 1790
nobile misident.
Sphex nobilis Scopoli, 1763
lucidula (Fabricius, 1775, Chrysis)
regia (Fabricius, 1793, Chrysis)
added by
intermedium misident.
Chrysis aenea Fabricius, 1787
Ellampus puncticollis Mocsáry, 1887
Treated as a subspecies of aeneus in Fauna Europaea.
Chrysis truncata Dahlbom, 1831
Sphex aurata Linnaeus, 1758
Sphex violacea Scopoli, 1763
Some distributional data and synonymy from
CHELOGYNUS Haliday, 1838
NEOCHELOGYNUS Perkins, 1905
imberbis Kieffer, 1905
bensoni Richards, 1939
jurineanum in part, Perkins, 1976
Dryinus brachycerus Dalman, 1823
lyde (Walker, 1837, Dryinus)
nigricornis Kieffer, 1905
triareolatus Kieffer, 1905
brevicollis Kieffer, 1905
flavitarsis Kieffer, 1905
indivisus Kieffer, 1905
nigroclavatus Kieffer, 1905
curvatus Kieffer, 1906
obscuricornis Kieffer, 1906
suffolciensis (Chitty, 1908, Antaeon)
curvus Kieffer, 1914
Gonatopus ephippiger Dalman, 1818
collaris (Dalman, 1818, Gonatopus)
facialis (Thomson, 1860, Dryinus)
albidicollis Kieffer, 1905
rubrifrons Kieffer, 1905
rufovariegatus (Berland, 1928, Chelogynus)
albidocolle Richards, 1939
pyonganensis Moczar, 1983
Chelogynus exiguus Haupt, 1941
subarcticus Hellén, 1935 nom. nud.
flaviscapus Jansson, 1950
subarcticus Hellén, 1953
added by
Dryinus facialis Thomson, 1860
pseudohilare Burn, 1990
Gonatopus flavicornis Dalman, 1818
sericeus Kieffer, 1905
subflavicornis Haupt, 1941
Dryinus fulviventris Haliday, 1828
fuscipes (Thomson, 1860, Dryinus)
similis Kieffer, 1905
gracilicollis Kieffer, 1905
flavinervis Kieffer, 1905
parvulus Kieffer, 1905
xanthostigma Kieffer, 1905
flaviscapus Kieffer, 1905
parvus Kieffer, 1906
alutaceus (Richards, 1935, Chelogynus)
cameroni Kieffer, 1905
trivialis Kieffer, 1905
rufulocollis (Chitty, 1908, Antaeon)
Dryinus infectus Haliday, 1837
luteicornis misident. sensu
inclytus (Haliday, 1837, Dryinus)
lateralis (Thomson, 1860, Dryinus)
fusiformis Kieffer, 1905
punctatus Kieffer, 1905
ellimani (Chitty, 1908, Antaeon)
brevicornis (Dalman, 1818, Gonatopus)
cursor (Haliday, 1828, Dryinus)
otiartes (Walker, 1837, Dryinus)
sisithrus (Walker, 1837, Dryinus)
nanus (Haliday, 1837, Dryinus)
crenulatus Kieffer, 1905
thomsoni Kieffer, 1905
vicinus Kieffer, 1905
marginatus Kieffer, 1905
rectus Kieffer, 1905
scoticus Kieffer, 1905
barbatus (Chitty, 1908, Antaeon)
brunneipes (Berland, 1928, Xenanteon)
nec jurineanum Richards, 1939
nec jurineanum Perkins, 1976
Gonatopus pubicornis Dalman, 1818
tenuicornis (Dalman, 1823, Dryinus)
cephalotes (Ljungh, 1824, Gonatopus)
lucidus (Haliday, 1828, Dryinus)
penidas (Walker, 1837, Dryinus)
alorus (Walker, 1837, Dryinus)
fuscoclavatus Kieffer, 1905
triangularis Kieffer, 1905
divisus Kieffer, 1905
vulgaris Kieffer, 1905
breviventralis (Chitty, 1908, Antaeon)
delicatulus (Chitty, 1908, Antaeon)
serratus (Maneval, 1935, Chelogynus)
exiguus (Haupt, 1941, Chelogynus)
mongolicum Moczar, 1983
Included as a synonym of brachycerum by
added by
Dryinus scapularis Haliday, 1837
longiforceps Kieffer, 1905
carinatus Kieffer, 1905
lanionis (Haupt, 1941, Chelogynus)
tricarinatus Kieffer, 1905
kiefferi (Chitty, 1908, Antaeon)
angusticollis (Berland, 1928, Chelogynus)
prehensor (Maneval, 1935, Chelogynus)
berlandi (Richards, 1936, Chelogynus)
silvaticus (Ponomarenko, 1970, Chelogynus)
PRENANTEON Kieffer, 1913
PSILANTEON Kieffer, 1913
Gonatopus ruficornis Dalman, 1818
basalis (Dalman, 1818, Gonatopus)
frontalis (Dalman, 1818, Gonatopus)
fuscicornis (Dalman, 1818, Gonatopus)
longicornis (Dalman, 1823, Dryinus)
crassimanus (Haliday, 1828, Dryinus)
daos (Walker, 1837, Dryinus)
ilus (Walker, 1837, Dryinus)
misor (Walker, 1837, Dryinus)
lapponicus (Thomson, 1860, Dryinus)
retusus (Thomson, 1860, Dryinus)
lepidus (Förster, 1861, Chelogynus)
subapterus (Kieffer, 1905, Anteon)
aequalis (Kieffer, 1905, Anteon)
melanocera (Kieffer, 1905, Anteon)
vitellinipes (Kieffer, 1905, Anteon)
procericornis (Kieffer, 1905, Anteon)
declivis (Kieffer, 1905, Anteon)
pallidinervis (Kieffer, 1905, Anteon)
integer (Kieffer, 1905, Anteon)
curvinervis (Kieffer, 1905, Anteon)
fractinervis (Kieffer, 1905, Anteon)
hyalinipennis (Kieffer, 1905, Anteon)
longifilis (Kieffer, 1906, Anteon)
halidayi (Kieffer, 1906, Anteon)
morleyi (Chitty, 1908, Antaeon)
luffnessensis (Chitty, 1908, Antaeon)
beaumonti (Chitty, 1908, Antaeon)
walkeri (Kieffer, 1914, Chelogynus)
parcepunctatus (Kieffer, 1914, Prenanteon)
palustris (Oglobin, 1924, Prenanteon)
filicornis (Oglobin, 1924, Prenanteon)
euscelisi (Haupt, 1941, Prenanteon)
semenovi (Ponomarenko, 1970, Prenanteon)
foveatus (Richards, 1971, Prenanteon)
pektusanense (Moczar, 1983, Prenanteon)
clavatum (Moczar, 1983, Prenanteon)
ANTAPHELOPUS Benoit, 1951
GYMNAPHELOPUS Benoit, 1951
Data on five species reared in Wales from
Dryinus atratus Dalman, 1823
holomelas Richards, 1939
heidelbergensis Richards, 1939
Gonatopus melaleucus Dalman, 1818
albipes (Ratzeburg, 1848, Ceraphron)
added by
CAMPYLONYX Westwood, 1835
PARADRYINUS Perkins, 1905
CHLORODRYINUS Perkins, 1905
PLASTODRYINUS Kieffer, 1906
MESODRYINUS Kieffer, 1906
HESPERODRYINUS Perkins, 1907
Sphex collaris Linnaeus, 1767
formicarius Latreille, 1805
ampuliciformis (Westwood, 1835, Campylonyx)
corsicae (Kieffer, 1914, Lestodryinus)
brittanicus (Richards, 1939, Mesodryinus)
DICONDYLUS Haliday, 1829-30
LABEO Haliday, 1833
PSEUDOGONATOPUS Perkins, 1905
NEOGONATOPUS Perkins, 1905
PACHYGONATOPUS Perkins, 1905
CHALCOGONATOPUS Perkins, 1905
EUGONATOPUS Perkins, 1905
PLATYGONATOPUS Kieffer, 1906
AGONATOPUS Perkins, 1907
AGONATOPOIDES Perkins, 1907
EUCAMPTONYX Perkins, 1907
CYRTOGONATOPUS Kieffer, 1907
DIGONATOPUS Kieffer, 1913
LABERIUS Kieffer, 1914
TRICHOGONATOPUS Hellén, 1930
METAGONATOPUS Oglobin, 1932
ALLOGONATOPUS Haupt, 1938
DONISTHORPINA Richards, 1939
PLECTROGONATOPUS Richards, 1939
TETRODONTOCHELYS Richards, 1939
EPIGONATOPOIDES Richards, 1939
RHYNCHOGONATOPUS Benoit, 1953
MADECAGONATOPUS Benoit, 1953
CYRTOGONATOPOIDES Ponomarenko, 1966
MEGAGONATOPUS Olmi & Currado, 1976
TETRADONTOCHELYS Perkins, 1976
separatus (Richards, 1939, Pseudogonatopus) synonymy by
England, Wales
Dryinus bicolor Haliday, 1828
vitripennis (Haliday, 1833, Labeo)
excisus (Westwood, 1833, Antaeon)
conjunctus Kieffer, 1905
bifarius Kieffer, 1906
decretorius Haupt, 1916
lindbergi (Heikinheimo, 1957, Dicondylus)
Gelis clavipes Thunberg, 1827
sepsoides Westwood, 1833
pilosus Thomson, 1860
nigerrimus (Förster, 1861, Labeo)
pusillus Szépligeti, 1901, Labeo)
hispanicus Kieffer, 1905
sociabilis Kieffer, 1907
borealis Sahlberg, 1910
wagneri Strand, 1919
barbatellus Richards, 1939
campestris Ponomarenko, 1965
rhaensis Ponomarenko, 1970
septemdentatus Sahlberg, 1910
robustus (Ceballos, 1927, Dicondylus)
flavicornis Thomson, 1860 preocc.
luteicornis Kieffer, 1905
excavatus Sahlberg, 1910
liechtensteini Picard, 1932
procerus (Haupt, 1938, Allogonatopus)
thomsoni Hellén, 1953
rossicus Ponomarenko, 1965
Donisthorpina formicicola Richards, 1939
England, Wales
dichromus Kieffer, 1906
rufescens Hellén, 1935
added by
bifasciatus Kieffer, 1904
gracilicornis Kieffer, 1904
filicornis Kieffer, 1905
gracilis Kieffer, 1905
marshalli Kieffer, 1905
myrmecophilus Kieffer, 1905
gracilipes Kieffer, 1906
raptoripes Strand, 1919
ljunghii Westwood, 1833
leucostomus Sahlberg, 1910
arnoldii (Ponomarenko, 1966, Pachygonatopus)
richardsi (Moczar, 1965, Plectrogonatopus)
tauricus (Ponomarenko, 1965, Agonatopoides)
MONOGONATOPUS Richards, 1939
Gonatopus oratorius Westwood, 1833
England
MYRMECOMORPHUS Westwood, 1833
POLYPLANUS Nees, 1834
EMBOLIMUS Agassiz, 1846
FORMILA De Romand, 1846
PEDINOMMA Förster, 1856
AMPULICOMORPHA Ashmead, 1893
AMPULICIMORPHA Brues, 1933
Myrmecomorphus ruddii Westwood, 1833
rufescens (Westwood, 1833, Myrmecomorphus)
sickershusanus (Nees, 1834, Polyplanus)
antennalis (Kieffer, 1906, Pedinomma) synonymy by
holochlora (Kieffer, 1906, Pedinomma)
hypochlora (Kieffer, 1906, Pedinomma)
rufus Kieffer, 1906
Some distribution data for Mutillidae, Sapygidae, Tiphiidae and Vespidae from
Nomenclature, synonymy and higher classification based on
[LINEPITHEMA Mayr, 1866 # humile (Mayr, 1868, Hypoclinea)] Although this highly invasive species has the potential to establish itself in hothouses, given the correct climate, we are unaware of any established colonies being found in Britain. Any introductions have been of isolated workers.
MICROMYRMA Dufour, 1857
SEMONIUS Forel, 1910
TAPINOPTERA Santschi, 1925
ZATAPINOMA Wheeler, 1928
NEOCLYSTOPSENELLA Kurian, 1955
Formica erratica Latreille, 1798
caerulescens (Losana, 1834, Formica)
glabrella (Nylander, 1849, Formica)
collina Förster, 1850
bononiensis Emery, 1925
breve Emery, 1925
tauridis Emery, 1925
transcaucasica Karavaiev, 1927
ambiguum misident.
madeirense misident.
Formica melanocephala Fabricius, 1793
nana (Jerdon, 1851, Formica) preocc.
pellucida (Smith, 1857, Myrmica)
familiaris (Smith, 1860, Formica) preocc.
australe Santschi, 1928
australis Santschi, 1928
FORMICINA Schuckard, 1840
NEOFORMICA Wheeler, 1913
RAPTIFORMICA Forel, 1913
SERVIFORMICA Forel, 1913
COPTOFORMICA Müller, 1923
ADFORMICA Lomnicki, 1925
schmidti Ruzsky, 1920 preocc.
fuscorufibarbis Forel, 1874
glauca Ruzsky, 1896
rubescens Forel, 1904
caucasica Wheeler, 1913
volgensis Ruzsky, 1914
katuniensis Ruzsky, 1915
montivaga Santschi, 1928
montaniformis Kuznetsov-Ugamsky, 1929
fuscoides Dlussky, 1967
exsectopressilabris Forel, 1874
rubens Forel, 1874
etrusca Emery, 1909
dalcqi Bondroit, 1918
sudetica Scholz, 1924
wheeleri Creighton, 1935
kontuniemii Betrem, 1954
nemoralis Dlussky, 1964
libera Scopoli, 1763
flavipes Geoffroy, 1785
barbata Razoumowski, 1789
tristis Christ, 1791
glebaria Nylander, 1846
marcida Wheeler, 1913
pallipes Kuznetsov-Ugamsky, 1926
rufipes Stitz, 1930
borealis Vashkevich, 1924
congerens Nylander, 1846
santschii Wheeler, 1913
nylanderi Bondroit, 1920
unicolor Ruzsky, 1926
montana Sadil, 1953
candida misident.
glabra White, 1884 preocc.
transkaucasica Nasonov, 1889
orientalis Ruzsky, 1915
piceoinplana Emery, 1925
inplana Emery, 1925
lochmatteri Stärcke, 1935
Not a homonym under article 23.9.5 of the Code (
nigricans Bondroit, 1912
cordieri Bondroit, 1917
grouvellei Bondroit, 1918
ciliata Ruzsky, 1926
thyssei Stärcke, 1942
pratensoides Gösswald, 1951
Probably extinct in Britain (
ferruginea Christ, 1791
dorsata Panzer, 1798
major Nylander, 1849
piniphila Schenck, 1852
apicalis Smith, 1858
rufopratensis Forel, 1874
meridionalis Nasonov, 1889
gaullei Bondroit, 1917
nicaeensis Leach, 1825
stenoptera Förster, 1850
cinereorufibarbis Forel, 1874
defensor Smith, 1878
fraterna Smith, 1878
piligera Lomnicki, 1925
dominula Nylander, 1846
fusciceps Emery, 1895
mollesonae Ruzsky, 1903
clarior Ruzsky, 1905
flavorubra Forel, 1909
borea Santschi, 1925
strennua Santschi, 1925
griseopubescens Kuznetsov-Ugamsky, 1926
monticola Kuznetsov-Ugamsky, 1926
rotundata Kuznetzsov-Ugamsky, 1926
arenicola Kuznetsov-Ugamsky, 1928
leninei Santschi, 1928
tristis Karavaiev, 1929
CHTHONOLASIUS Ruzsky, 1912
DENDROLASIUS Ruzsky, 1912
CAUTOLASIUS Wilson, 1955
AUSTROLASIUS Faber, 1967
Formica aliena Foerster, 1850
americanus Emery, 1893
pannonica Röszler, 1942
Formica brunnea Latreille, 1798
pallida (Latreille, 1798, Formica)
timida (Förster, 1850, Formica)
alienobrunneus Forel, 1874
nigrobrunneus (Donisthorpe, 1926, Acanthomyops)
Formica emarginata Olivier, 1792
brunneoemarginatus Forel, 1874
brunneoides Forel, 1874
nigroemarginatus Forel, 1874
illyricus Zimmermann, 1935
pontica Stärcke, 1944
added by
Formica flava Fabricius, 1781
ruficornis (Fabricius, 1804, Formica)
brevicornis Emery, 1893
fuscoides Ruzsky, 1902
odoratus Ruzsky, 1905
claripennis Wheeler, 1917
microps Wheeler, 1917
morbosa (Bondroit, 1918, Formicina)
ibericus Santschi, 1925
apennina Menozzi, 1925
olivacea Karavaiev, 1926
helvus Cook, 1953
Formica fuliginosa Latreille, 1798
Formicina meridionalis Bondroit, 1920
Formica mixta Nylander, 1846
added by
Formica nigra Linnaeus, 1758
nigerrima (Christ, 1791, Formica)
pallescens (Schenck, 1852, Formica)
alienoniger Forel, 1874
emeryi Ruzsky, 1905
nitidus (Kuznetzsov-Ugamsky, 1927, Acanthomyops)
minimus (Kuznetsov-Ugamsky, 1928, Acanthomyops)
transylvanica Röszler, 1943
added by
added by
Formicina sabularum Bondroit, 1918
Formica umbrata Nylander, 1846
aphidicola (Walsh, 1863, Formica)
exacutus Ruzsky, 1902
affinoumbratus Donisthorpe, 1914
belgarum (Bondroit, 1918, Formicina)
silvestrii Wheeler, 1928
hirtiscapus Stärcke, 1937
osakana Santschi, 1941
nyaradi (Röszler, 1943, Chthonolasius)
epinotalis Buren, 1944
Formica longicornis Latreille, 1802
vagans (Jerdon, 1851, Formica)
gracilescens (Nylander, 1856, Formica)
currens Motschoulsky, 1863
APOROMYRMEX Faber, 1969
PARAPLAGIOLEPIS Faber, 1969
barbara Santschi, 1911
crosi Santschi, 1920
madeirensis Emery, 1921
Two nests persisted on the Isle of Wight from 2007-2008 (BB, pers. obs., specimens in NHM, BB coll. and B. Seifert coll.) but appear to have died out since. A native of the Azores and Madeira.
The tribal classification follows
Formica megacephala Fabricius, 1793
edax (Forskål, 1775, Formica)
trinodis (Losana, 1834, Myrmica)
pusilla (Heer, 1852, Oecophthora)
laevigata (Smith, 1855, Myrmica)
agilis (Smith, 1857, Myrmica)
testacea (Smith, 1858, Atta)
janus Smith, 1858
suspiciosa (Smith, 1859, Myrmica)
laevigata Mayr, 1862
perniciosa (Gerstäcker, 1862, Oecophthora)
picata Forel, 1891
scabrior Forel, 1891
gietleni Forel, 1905
bernhardae Emery, 1915
A tramp species, in the past regarded as a proper introduction in Britain but no established colonies have been reported in the period 1970–present. On a world-wide scale its importance as a tramp species seems to be diminishing. The origin of megacephala is certainly Afrotropical as it is a member of a large and successful species group that is otherwise confined to that region.
Cardiocondyla britteni Crawley, 1920 was described from a worker casually introduced in Britain; included here only because the type-locality is Britain.
Crematogaster scutellaris (Olivier, 1792, Formica) is an occasional accidental import with cork from southern Europe. It has never established in Britain and, with the decrease in the cork trade, imports of this species will probably become even rarer.
SYMMYRMICA Wheeler, 1904
Myrmica nitidula Nylander, 1846
laeviuscula (Förster, 1850, Myrmica)
picea (Wasmann, 1906, Leptothorax)
DORONOMYRMEX Kutter, 1945
MYCHOTHORAX Ruzsky, 1904
Many species previously placed in Leptothorax have been transferred to Temnothorax by
Formica acervorum Fabricius, 1793
lacteipennis (Zetterstedt, 1838, Myrmica)
nigrescens Ruzsky, 1905
superus Ruzsky, 1905
kamtschaticum Ruzsky, 1920
orientalis (Kuznetsov-Ugamsky, 1928, Mychothorax)
ARCHAEOMYRMEX Mann, 1921
Formica graminicola Latreille, 1802
latreillei Curtis, 1829
striatula (Nylander, 1849, Myrmica)
bidens (Förster, 1850, Myrmica)
kutteri Forel, 1914
grouvellei Bondroit, 1918
gotlandica Karavaiev, 1930
oelandica Karavaiev, 1930
dentata Santschi, 1939
MYRMUS Schenck, 1853
Eciton testaceum Schenck, 1852
emarginatus (Schenck, 1852, Myrmus)
diveri Donisthorpe, 1936
MACROMISCHA Roger, 1863
DICHOTHORAX Emery, 1895
MYRMOXENUS Ruzsky, 1902
PROTOMOGNATHUS Wheeler, 1905
ANTILLAEMYRMEX Mann, 1920
CROESOMYRMEX Mann, 1920
CHALEPOXENUS Menozzi, 1923
MYRMAMMOPHILUS Menozzi, 1925
MYRAFANT Smith, 1950
ICOTHORAX Hamann & Klemm, 1967
Stenamma albipennis Curtis, 1854
tuberum misident.;
tuberointerruptus (Bondroit, 1918, Leptothorax)
Myrmica interrupta Schenck, 1852
simpliciuscula (Nylander, 1856, Myrmica)
tuberoaffinis (Bondroit, 1918, Leptothorax)
Myrmica nylanderi Foerster, 1850
cingulata (Schenck, 1852, Myrmica)
nylanderocorticalis (Forel, 1874, Leptothorax)
nylanderotuberum (Ruzsky, 1902, Leptothorax)
Formica unifasciata Latreille, 1798
anoplogynus (Emery, 1869, Leptothorax)
unifasciatointerruptus (Forel, 1874, Leptothorax)
kirillovi (Ruzsky, 1905, Leptothorax)
brauneri (Karavaiev, 1937, Leptothorax)
salina (Karavaiev, 1937, Leptothorax)
ucrainicus (Arnol’di, 1977, Leptothorax)
England
Recorded from England as an accidental introduction by
TETROGMUS Roger, 1857
ANERGATES Forel, 1874
XIPHOMYRMEX Forel, 1887
TRIGLYPHOTHRIX Forel, 1890
RHOPTROMYRMEX Mayr, 1901
ATOPULA Emery, 1912
MACROMISCHOIDES Wheeler, 1920
LOBOMYRMEX Kratochvil, 1941
TELEUTOMYRMEX Kutter, 1950
Myrmica atratula Schenck, 1852
friedlandi (Creighton, 1934, Anergates)
Myrmica bicarinata Nylander, 1846
guineense misident.
cariniceps (Guérin-Méneville, 1852, Myrmica)
kollari (Mayr, 1853, Myrmica)
modesta (Smith, 1860, Myrmica)
reticulata (Smith, 1862, Myrmica)
Formica caespitum Linnaeus, 1758
fusca (Leach, 1825, Formica)
fuscula (Nylander, 1846, Myrmica)
modesta (Foerster, 1850, Myrmica)
himalayanum Viehmeyer, 1914
hammi Donisthorpe, 1915
immigrans Santschi, 1927
indocile Santschi, 1927
transbaicalense Ruzsky, 1936
transversinodis (Enzmann, 1946, Myrmica)
fusciclavum Consani & Zngheri, 1952
jiangxiense Wang & Xiao, 1988
Tetramorium caespitum s.l. is now recognised as comprising several species but the species present in Britain is apparently the true caespitum, at least on the basis of specimens from southern England (
Myrmica simillima Smith, 1851
parallela (Smith, 1859, Myrmica)
pygmaeum Emery, 1877
denticulatum Forel, 1902
bantouana Santschi, 1910
opacior Forel, 1913
exoleta Santschi, 1914
brevispinosa (Borgmeier, 1928, Wasmannia)
insulare Santschi, 1928
SIFOLINIA Emery, 1907
SOMMIMYRMA Menozzi, 1925
SYMBIOMYRMA Arnol’di, 1930
PARAMYRMICA Cole, 1957
DODECAMYRMICA Arnol’di, 1968
Symbiomyrma karavajevi Arnol’di, 1930
laurae misident.
pechei (Samsinak, 1957, Sifolinia)
faniensis van Boven, 1970
winterae (Kutter, 1973, Sifolinia)
denticornis Curtis, 1854
arduennae Bondroit, 1911
angustifrons Stärcke, 1927
brunescens Karavaiev, 1929
burtshakabramovitshi Karavaiev, 1929
starki Karavaiev, 1929
foreli Santschi, 1931
alpestris Arnol’di, 1934
kievensis Karavaiev, 1934
lissahorensis Stitz, 1939
added by
Formica rubra Linnaeus, 1758
laevinodis Nylander, 1846
longiscapus Curtis, 1854
champlaini Forel, 1901
europaea Finzi, 1926
bruesi Weber, 1947
microrubra Seifert, 1993
dimidiata Say, 1836
diluta Nylander, 1849
ruginodolaevinodis Forel, 1874
silvestrii Wheeler, 1928
sontica Santschi, 1937
yoshiokai Weber, 1947
macrogyna Brian & Brian, 1949
microgyna Brian & Brian, 1949
mutata Sadil, 1952
scabrinodolobicornis Forel, 1874
rugulosoides Forel, 1915
pilosiscapus Bondroit, 1920
ahngeri Karavaiev, 1926
scabrinodosabuleti Sadil, 1952
kutteri Finzi, 1926
subopaca Arnol'di, 1934
betuliana Ruzsky, 1946
schenckioides Boer & Noordijk, 2005
silvestrianum Emery, 1924
striata Finzi, 1926
sancta Karavaiev, 1926
nevodovskii (Karavaiev, 1926, Leptothorax)
turcica Santschi, 1931
puerilis Stärcke, 1942
dolens Stärcke, 1942
balcanica Sadil, 1952
scabrinodoides Sadil, 1952
tschuliensis Arnol’di, 1976
kozakorum Radchenko & Elmes, 2010
perelegans Curtis, 1854
nigripes Ruzsky, 1895
myrmecophila Wasmann, 1910
sulcinodoruginodis Donisthorpe, 1915
sulcinodoscabrinodis Forel, 1915
derzhavini Ruzsky, 1920
vicaria Kuznetsov-Ugamsky, 1928
eximia Kupyanskaya, 1990
England, Wales
added by
Atta floricola Jerdon, 1851
cinnabari Roger, 1863
poecilum Roger, 1863
specularis Mayr, 1866
impressum Smith, 1876
philippinensis Forel, 1910
furina Forel, 1911
floreanum Stitz, 1932
angusticlava Donisthorpe, 1947
Formica pharaonis Linnaeus, 1758
antiguensis (Fabricius, 1793, Formica)
domestica (Shuckard, 1838, Myrmica)
minuta (Jerdon, 1851, Atta)
vastator (Smith, 1857, Myrmica)
contigua (Smith, 1858, Myrmica)
fragilis (Smith, 1858, Myrmica)
DIPLORHOPTRUM Mayr, 1855
OCTELLA Forel, 1915
SYNSOLENOPSIS Forel, 1918
DIAGYNE Santschi, 1923
EUOPHTHALMA Creighton, 1930
LABAUCHENA Santschi, 1930
OEDALEOCERUS Creighton, 1930
BISOLENOPSIS Kusnezov, 1953
PARANAMYRMA Kusnezov, 1953
GRANISOLENOPSIS Kusnezov, 1957
LILIDRIS Kusnezov, 1957
Formica fugax Latreille, 1798
flavidula (Nylander, 1849, Myrmica)
latroides Ruzsky, 1905
orientalis Ruzsky, 1905
kasalinensis Emery, 1909
pontica Santschi, 1934
scythica Santschi, 1934
furtiva Santschi, 1934
balachowskyi Bernard, 1950
banyulensis Bernard, 1950
duboscqui Bernard, 1950
laevithorax Bernard, 1950
monticola Bernard, 1950
nicaeensis Bernard, 1950
provincialis Bernard, 1950
pygmaea Bernard, 1950
richardi Bernard, 1950
robusta Bernard, 1950
rugosa Bernard, 1950
tertialis Ettershank, 1966
avium (Bernard, 1978, Diplorhoptrum)
delta (Bernard, 1978, Diplorhoptrum)
insulare (Bernard, 1978, Diplorhoptrum)
pilosum (Bernard, 1978, Diplorhoptrum)
ASEMORHOPTRUM Mayr, 1861
THERYELLA Santschi, 1921
Myrmica debilis Förster, 1850
minkii (Förster, 1850, Myrmica)
golosejevi Karavaiev, 1926
ucrainicum Arnol’di, 1928
polonicum Begdon, 1932
orousseti Casevitz-Weulerrse, 1990
added by
Records from Ireland may relate to debile (Fox in
Hypoponera gibbinota (Forel, 1912, Ponera) was described from a worker casually introduced to Britain.
Ponera punctatissima Roger, 1859
androgyna (Roger, 1859, Ponera)
tarda (Charsley, 1877, Ponera)
jugata (Forel, 1892, Ponera)
brevis (Santschi, 1911, Ponera)
cognata (Santschi, 1912, Ponera)
durbanensis (Forel, 1914, Ponera)
incisa (Santschi, 1914, Ponera)
sordida (Santschi, 1914, Ponera)
petri (Forel, 1916, Ponera)
exacta (Santschi, 1923, Ponera)
mina (Wheeler, 1927, Ponera)
mumfordi (Wheeler, 1931, Ponera)
argonautorum (Arnol’di, 1932, Ponera)
mesoepinotalis (Weber, 1942, Ponera)
breviceps (Bernard, 1953, Ponera)
ursoidea (Bernard, 1953, Ponera)
sulcitana (Stefani, 1970, Ponera)
Ponera ergatandria Forel, 1893
schauinslandi (Emery, 1899, Ponera) synonymy by
kalakauae (Forel, 1899, Ponera)
aemula (Santschi, 1911, Ponera)
bondroiti (Forel, 1911, Ponera)
PSEUDOCRYPTOPONE Wheeler, 1933
SELENOPONE Wheeler, 1933
PTEROPONERA Bernard, 1950
Formica coarctata Latreille, 1802
contracta (Latreille, 1802, Formica)
lucida Emery, 1898
atlantis Santschi, 1921
crassisquama Emery, 1916
added by
Nomenclature follows
Mutilla rufipes Fabricius, 1787
melanocephala (Fabricius, 1793, Mutilla) preocc.
The nominate subspecies occurs in England and Wales, atra erythrocephala Yarrow, 1954 in Ireland and the Isle of Man.
PSAMMOCHARIDAE Banks, 1910
Classification follows
PILPOMUS Costa, 1859
PSEUDAGENIA Kohl, 1844
Sphex carbonaria Scopoli, 1763
punctum (Fabricius, 1804, Ceropales)
canaliculatus (Schenck, 1857, Agenia)
submarginatus Lepeletier, 1845
albigena Lepeletier, 1845
collinus Haupt, 1962
silvalis Haupt, 1962
CALICURGUS Lepeletier, 1845
Pompilus fasciatellus Spinola, 1808
calcaratus (Dahlbom, 1829, Pompilus)
maculipennis (Dahlbom, 1829, Pompilus)
albispinus (Herrich-Schäffer, 1830, Pompilus)
curtus (Zetterstedt, 1838, Pompilus)
gyllenhali (Dahlbom, 1843, Priocnemis)
labiatus (Lepeletier, 1845, Anoplius)
unimacula (Lepeletier, 1845, Anoplius)
odontellus (Lepeletier, 1845, Calicurgus)
bivirgulatus (Costa, 1881, Pompilus)
fuscopennis (Verhoeff, 1892, Priocnemis)
CALICURGUS Brullé, 1833
CHLOROCHEILUS Wolf, 1965
SALIUS Fabricius, 1804
HOMONOTUS Dahlbom, 1845
Sphex notata Rossi, 1792
?guttus (Spinola, 1808, Pompilus)
affinis Vander Linden, 1827, Pompilus)
iracundus (Dufour, 1841, Pompilus)
apricus (Lepeletier, 1845, Calicurgus)
melanius (Lepeletier, 1845, Calicurgus)
?binotatus (Marquet, 1879, Priocnemis) preocc.
marquetii (Dalla Torre, 1897, Salius)
orientalis Haupt, 1927
Ichneumon bifasciatus Geoffroy, 1785
hircanus (Fabricius, 1798, Pompilus)
intermedius (Dahlbom, 1843, Agenia)
Pogonius subintermedius Magretti, 1886
nitidus (Haupt, 1927, Deuteragenia)
Sphex variegata Linnaeus, 1758
erythropus (Kohl, 1888, Agenia)
structor (Ferton, 1897, Agenia)
faggiolii (Haupt, 1927, Deuteragenia)
MACULIPENNIS Junco, 1946
Pompilus agilis Shuckard, 1837
obtusiventris Schiødte, 1837
fraterculus Junco, 1946
gracilis Haupt, 1927 preocc.
gussakowskiji Wolf, 2004 preocc.
Sphex exaltata Fabricius, 1775
gibba (Scopoli, 1763, Sphex)
revo (Harris, 1780, Sphex)
nudipes Dahlbom, 1845
longicornis Haupt, 1927
valkeilai Wolf, 1959
Sphex hyalinata Fabricius, 1793
femoralis (Dahlbom, 1829, Pompilus)
discrepans (Costa, 1887, Pseudagenia)
trifurcus Radoszkowski, 1888
vitripennis Verhoeff, 1892
notatulus (Saunders, 1896, Salius)
pseudofemoralis Sustera, 1938
taigaica Wolf, 1967
minor (Zetterstedt, 1879, Pompilus)
mocsaryi Gussakovskij, 1930
klosei Haupt, 1937
haupti Sustera, 1938
vinetorum Blüthgen, 1944
minutalis Wahis, 1979 nom. auct.
Calicurgus propinquus Lepeletier, 1845
agenoides Dubois, 1920
Pompilus pusillus Schiødte, 1837
PRIOCNEMISSUS Haupt, 1949
capciosus Junco, 1946
Sphex perturbator Harris, 1780
fusca misident.
ambustor (Panzer, 1804, Ichneumon)
serripes (Dahlbom, 1829, Pompilus)
ambulator (Lepeletier, 1845, Calicurgus)
sepicola (Smith, 1851, Pompilus)
ater Wolf, 1960
clementi Haupt, 1927
gasconia Wolf, 1975
APOROIDEUS Ashmead, 1902
GYMNOCHARES Banks, 1917
Pompilus cinctellus Spinola, 1808
clypeatus (Dahlbom, 1829, Pompilus)
punctipes (Dahlbom, 1832, Pompilus)
tibialis (Lepeletier, 1845, Anoplius)
Pompilus sericeus Vander Linden, 1827
vicinus (Lepeletier, 1845, Pompilus)
subserricornis (Kohl, 1879, Pompilus)
declivus (Tournier, 1889, Pompilus)
gaullei (Tournier, 1889, Pompilus)
hungaricus (Moczar, 1944, Anospilus)
added by
Pompilus caviventris Aurivillius, 1907
cardui (Perkins, 1917, Pompilus)
carbonarius Haupt, 1937
atricolor Moczar, 1944
Pompilus concinnus Dahlbom, 1845
vacillans (Wesmael, 1851, Pompilus)
approximatus (Smith, 1877, Pompilus)
bifidus (Morawitz, 1891, Pompilus)
distinguendus (Morawitz, 1891, Pompilus)
haereticus (Tournier, 1889, Pompilus)
Sphex nigerrima Scopoli, 1763
nigrus (Fabricius, 1775, Sphex)
incisus (Tischbein, 1850, Pompilus)
melanarius (Schenck, 1857, Pompilus) preocc.
excerptus (Tournier, 1889, Pompilus)
difficilis (Tournier, 1889, Pompilus)
wheeleri Banks, 1939
banksi Dreisbach, 1950
POMPILINUS Ashmead, 1902
Pompilus infuscatus Vander Linden, 1827
minor (Herrich-Schäffer, 1830, Pompilus)
sericatus (Shuckard, 1835, Pompilus)
chalybeatus (Schiødte, 1837, Pompilus)
difformis (Schiødte, 1837, Pompilus)
dispar (Dahlbom, 1843, Pompilus)
sabulicola (Thomson, 1874, Pompilus)
meticulosa (Costa, 1882, Pompilus)
aeruginosus (Tournier, 1890, Pompilus)
aerarius (Tournier, 1890, Pompilus)
argentatus (Tournier, 1890, Pompilus)
calcatus (Tournier, 1890, Pompilus)
onus (Tournier, 1890, Pompilus)
stellatus (Tournier, 1890, Pompilus)
utendus (Tournier, 1890, Pompilus)
vivus (Tournier, 1890, Pompilus)
xysticus (Tournier, 1890, Pompilus)
petulans Haupt, 1962
cinctellus (Haupt, 1962, Paracyphonyx)
lusitanicus Wolf & Diniz, 1970
fortunatus Wolf, 1975
simii Wolf, 1978
England, Wales
Sphex viatica Linnaeus, 1758
fuscus (Linnaeus, 1761, Sphex)
paganus (Dahlbom, 1843, Pompilus)
propinquus (Smith, 1879, Pompilus)
tibialis (Tournier, 1890, Pompilus) preocc.
delatorius (Tournier, 1890, Pompilus)
immixtus (Tournier, 1890, Pompilus)
pleropicus (Tournier, 1890, Pompilus)
valesicus (Tournier, 1890, Pompilus)
macrurus (Dalla Torre, 1897, Pompilus)
holomelas (Mantero, 1905, Pompilus) preocc.
femoralis Vander Linden, 1827
castor (Kohl, 1838, Pompilus)
ANOPOMPILINUS Dreisbach, 1949
ARIDOPOMPILUS Wolf, 1965
BOREOPOMPILUS Wolf, 1965
HOLARCTOPOMPILUS Wolf, 1965
SAXATILIPOMPILUS Wolf, 1965
Pompilus anceps Wesmael, 1851
?vaga (Harris, 1870, Sphex) preocc.
unguicularis (Thomson, 1870, Pompilus)
crobaci (Tournier, 1890, Pompilus)
expleta (Tournier, 1890, Pompilus)
lustrica (Tournier, 1890, Pompilus)
nava (Tournier, 1890, Pompilus)
radiosa (Tournier, 1890, Pompilus)
saxea (Tournier, 1890, Pompilus)
peninsulana (Wolf, 1966, Pompilus)
serica Wolf & Moczar, 1972
Pompilus consobrinus Dahlbom, 1843
ater (Brullé, 1840, Pompilus) preocc.
excisa (Pérez, 1895, Pompilus) preocc.
nivariae (Dalla Torre, 1897, Pompilus)
guimarensis (Saunders, 1904, Pompilus)
lanuginosa (Haupt, 1927, Psammochares)
heringi (Haupt, 1928, Psammochares)
emissa (Haupt, 1930, Psammochares)
laufferi (Junco, 1960, Pompilus)
alpina (Wolf, 1965, Pompilus)
pyrenaica (Wolf, 1965, Pompilus)
continentalis (Wolf, 1966, Pompilus)
sicula (Wolf, 1966, Pompilus)
Pompilus trivialis Dahlbom, 1843
gibba misident.
aerumnata (Tournier, 1889, Pompilus)
corruptor (Haupt, 1927, Psammochares)
michalki (Blüthgen, 1961, Ammosphex)
insubrica (Wolf, 1965, Pompilus)
Pompilus wesmaeli Thomson, 1870
Pompilus minutulus Dahlbom, 1842
cellularis (Dahlbom, 1843, Pompilus)
neglecta (Dahlbom, 1843, Pompilus)
inermis (Lepeletier, 1845, Anoplius)
anoplius (Dalla Torre, 1897, Pompilus)
simplicicra (Priesner, 1960, Pompilus)
apenninusurata Wolf, 1970
Pompilus spissus Schiødte, 1837
apennina Wolf, 1970
PYCNOPOMPILUS Ashmead, 1902
Psammochares rufus Haupt, 1927
adulterina (Haupt, 1937, Psammochares)
melanota Wolf, 1975
SPILOPOMPILUS Ashmead, 1902
Pompilus gallicus Tournier, 1889
intermedius Haupt, 1930
tertius Blüthgen, 1944
added by
Sphex rufipes Linnaeus, 1758
laevigata (Rossius, 1794, Sphex)
gracilis (Lepeletier, 1845, Pompilus) preocc.
septemmaculatus (Wesmael, 1851, Pompilus)
argyrolepis (Costa, 1887, Pompilus)
compressus (Tournier, 1889, Pompilus) preocc.
aequatus (Tournier, 1889, Pompilus)
pygidialis (Tournier, 1889, Pompilus)
deuterus (Dalla Torre, 1897, Pompilus)
ephialtes (Dalla Torre, 1897, Pompilus)
sardous Wolf, 1961
SOPHROPOMPILUS Ashmead, 1902
ASTHENOCTENUS Arnold, 1937
ASTHENOCTENIDIA Pate, 1946
LEUCHIMON Haupt, 1930
TRICHOSYRON Haupt, 1930
PSAMMOCHAROIDES Moczar, 1946
STREPTOSELLA Dreisbach, 1950
CARINEVAGETES Wolf, 1970
CONTEMPTEVAGETES Wolf, 1970
Pompilus crassicornis Shuckard, 1835
dahlbomi (Thomson, 1870, Pompilus)
subarcticus Wolf, 1964
Aporus dubius Vander Linden, 1827
bicolor Lepeletier, 1845
servillei Costa, 1882
rattus (Dalla Torre, 1897, Pompilus)
obscurodubius Wolf, 1970
theodori Wolf, 1970
Sphex pectinipes Linnaeus, 1758
quadrispinosus (Kohl, 1886, Pompilus)
aculeatus (Thomson, 1870, Pompilus)
minotaurus Wolf, 1970
ISONOTUS Dahlbom, 1842
WESMAELINIUS Costa, 1886
Sphex sanguinolenta Fabricius, 1793
dispar (Latreille, 1809, Pompilus)
bidens (Lepeletier, 1845, Anoplius)
affinis (Eversmann, 1849, Pompilus) preocc.
nigrus (Marquet, 1879, Ferrola)
nasutus Morawitz, 1888
doctor (Dalla Torre, 1897, Pompilus)
CHIONOPOMPILUS Priesner, 1955
Sphex cinerea Fabricius, 1775
plumbeus (Fabricius, 1787, Sphex)
pulcher Fabricius, 1798
pruinosus Smith, 1879 preocc.
chevrieri Tournier, 1889
leprosus Dalla Torre, 1897
lusitanicus Wolf & Diniz, 1970
gotlandicus Wolf, 1972
CERATOPALES Schulz, 1906
HYPSICERAEUS Morice & Durrant, 1915
Evania maculata Fabricius, 1775
rustica (Müller, 1776, Sphex)
multicolor (Fourcroy, 1785, Ichneumon)
frontalis (Panzer, 1799, Pompilus)
perligerus (Costa, 1882, Priocnemis)
balearica Costa, 1893
Evania variegata Fabricius, 1798
destefanii Costa, 1887
Apis clavicornis Linnaeus, 1758
Scolia quinquepunctata Fabricius, 1781
METHOCA misspelling
Taxonomy follows
?formicaria (Latreille, 1792, Mutilla) nom. dub.
ichneumonides Latreille, 1805
mutillaria Latreille, 1806
sanvitali (Latreille, 1809, Tengyra)
italica (Costa, 1858, Spinolia)
Taxonomy mostly follows
Vespa antilope Panzer, 1798
pictus (Curtis, 1826, Odynerus)
?quadratus (Panzer, 1799, Vespa) nom. dub.
Although this species has been referred to as Ancistrocerus quadratus in British faunistic works (e.g.
Vespa gazella Panzer, 1798
emarginata (Fabricius, 1793, Vespa) preocc.
Odynerus nigricornis Curtis, 1826
callosus (Thomson, 1870, Odynerus)
excisus (Thomson, 1870, Odynerus)
sexpunctata (Christ, 1791, Vespa)
Odynerus oviventris Wesmael, 1836
constans (Herrich-Schäffer, 1839, Odynerus)
viduus (Herrich-Schäffer, 1839, Odynerus)
The subspecies hibernicus Blüthgen, 1937 occurs in Ireland and the Outer Hebrides.
Vespa parietina Linnaeus, 1758
domesticus (Christ, 1791, Sphex)
affinis (Herrich-Schäffer, 1839, Odynerus)
Vespa parietum Linnaeus, 1758
Odynerus scoticus Curtis, 1826
trimarginatus misident.
albotricinctus (Zetterstedt, 1838, Odynerus)
Vespa trifasciata Müller, 1776
trimarginatus (Zetterstedt, 1838, Odynerus)
tricinctus (Herrich-Schäffer, 1839, Odynerus)
Vespa coarctata Linnaeus, 1758
?papillarius (Christ, 1791, Sphex)
Vespa quadrifasciata Fabricius, 1793
tomentosus (Thomson, 1870, Odynerus)
Odynerus laevipes Shuckard, 1837
Odynerus exilis Herrich-Schäffer, 1839
bivittatus (Lepeletier, 1841, Odynerus)
Vespa melanocephala Gmelin, 1790
Vespa spinipes Linnaeus, 1758
quinquefasciata (Fabricius, 1793, Vespa) preocc.
muticus (Zetterstedt, 1838, Odynerus)
Vespa reniformis Gmelin, 1790
Odynerus herrichii Saussure, 1855
variegata misident.
basalis (Smith, 1857, Odynerus)
Vespa bifasciata Linnaeus, 1761
sinuatus (Fabricius, 1793, Vespa) preocc.
mutinensis (Baldeni, 1894, Odynerus)
sinuatissimus Richards, 1935
Odynerus connexus Curtis, 1826
bifasciatus misident.
Vespa crassicornis Panzer, 1798
Odynerus gracilis Brullé, 1832
Taxonomy follows
Vespa dominula Christ, 1791
gallicus misident.
italica Herrich-Schäffer, 1840 nom. nud.
pectoralis Herrich-Schäffer, 1841
lefebvrei Guérin, 1844
bucharensis Erichson, 1849
maculatus Rudow, 1889
merceti Dusmet, 1903
rufescens Buysson, 1912
ornata Weyrauch, 1938
pacfica Weyrauch, 1939 preocc.
pseudopacificus Giordani Soika, 1970
muchei Gusenleitner, 1976
Vespa gallica Linnaeus, 1767
pictior Radoszkowski, 1872
foederata Kohl, 1898
omissa (Weyrauch, 1938, Polistula)
Added by
Nomenclature follows
PSEUDOVESPULA Bischoff, 1931
BOREOVESPULA Blüthgen, 1943
METAVESPULA Blüthgen, 1943
Vespa medius Retzius, 1783
geerii (Lepeletier, 1836, Vespa)
crassa (Herrich-Schäffer, 1841, Vespa)
similis (Schenck, 1853, Vespa)
rufoscutellata (Schenck, 1853, Vespa)
flavicincta (Schenck, 1853, Vespa)
lineolata (Pérez, 1910, Vespa)
conjugens Paul, 1943
sugare Ishikawa, 1969
borealis Lee, 1986 preocc.
added by
Vespa norwegica Fabricius, 1781
britannica (Leach, 1814, Vespa)
marginata (Kirby, 1837, Vespa) preocc.
borealis (Zetterstedt, 1838, Vespa) preocc.
albida (Sladen, 1918, Vespa)
arctica (Friese, 1919, Vespa) preocc.
zetterstedti Blüthgen, 1937
Vespa saxonica Fabricius, 1793
bavarica (von Schrank, 1802, Vespa)
tridens (Schenck, 1853, Vespa)
nipponica Yamane, 1975
kamtschatkensis Eck, 1983
nigrescens Eck, 1983
added by
Vespa sylvestris Scopoli, 1763
parietum (Harris, 1776, Vespa)
holsatica (Fabricius, 1793, Vespa)
frontalis (Latreille, 1802, Vespa)
campanaria (Fowler, 1833, Vespa)
anglica (Smith, 1843, Vespa)
pilosella (Costa, 1858, Vespa)
sumptuosa (De Buysson, 1905, Vespa)
xinjiangensis Lee, 1986
MACROVESPA Dalla Torre, 1904
NYCTOVESPA van der Vecht, 1959
vexator Harris, 1776
major Retzius, 1783
?pratensis Geoffroy, 1785
germana Christ, 1791
crabroniformis Smith, 1852
borealis Radoszkowski, 1863
anglica Gribodo, 1892 preocc.
oberthuri du Buysson, 1902
flavo-fasciata Cameron, 1903
tartarea du Buysson, 1905
altaica Pérez, 1910
caspica Pérez, 1910
vulgata Birula, 1925
meridionalis Birula, 1925
chinensis Birula, 1925 preocc.
birulai Bequaert, 1931
gribodoi Bequaert, 1931
British populations are considered to belong to the subspecies gribodoi.
PSEUDOVESPA Schmiedeknecht, 1881
PARAVESPULA Blüthgen, 1938
ALLOVESPULA Blüthgen, 1943
RUGOVESPULA Archer, 1982
Vespa austriaca Panzer, 1799
borealis (Smith, 1843, Vespa) preocc.
arborea (Smith, 1849, Vespa)
biloba (Schilling, 1850, Vespa)
Vespa germanica Fabricius, 1793
?maculata (Scopoli, 1763, Vespa) preocc.
?macularis (Olivier, 1792, Vespa)
Vespa rufa Linnaeus, 1758
schrenckii (Radoszkowski, 1861, Vespa)
sibiria (André, 1884, Vespa)
grahami Archer, 1981
obscura Lee, 1986
?yichunensis Lee, 1986
Vespa vulgaris Linnaeus, 1758
sexcincta (Panzer, 1799, Vespa)
We are especially grateful to Jeremy Early for enlivening the checklist with so many lovely photographs of aculeates. Thanks also to Phil Adams for some additional, excellent photos. Paul Williams, Mike Edwards and Michael Kuhlmann have given helpful advice on apoid classification and nomenclature. Natalie Dale-Skey very kindly created the import spreadsheets from Word documents.
GE compiled an earlier version of this checklist and revised the current list of bees. BB compiled the checklist of ants, Formicidae. GRB updated all sections, added distribution data and edited the paper.
Word document version of the checklist
Excel spreadsheet version of the checklist